PMC:329117 JSON TXT 8 Projects

Annnotations TAB TSV DIC JSON TextAE Lectin_function IAV-Glycan

Id	Subject	Object	Predicate	Lexical cue
T246	0-7	NN	denotes	Odorant
T248	0-151	sentence	denotes	Odorant receptor expressed sequence tags demonstrate olfactory expression of over 400 genes, extensive alternate splicing and unequal expression levels
T247	8-16	NN	denotes	receptor
T249	17-26	VBN	denotes	expressed
T251	27-35	NN	denotes	sequence
T250	36-40	NNS	denotes	tags
T252	41-52	VBP	denotes	demonstrate
T253	53-62	JJ	denotes	olfactory
T254	63-73	NN	denotes	expression
T255	74-76	IN	denotes	of
T256	77-81	IN	denotes	over
T257	82-85	CD	denotes	400
T258	86-91	NNS	denotes	genes
T259	91-93	,	denotes	,
T260	93-102	JJ	denotes	extensive
T262	103-112	JJ	denotes	alternate
T261	113-121	NN	denotes	splicing
T263	122-125	CC	denotes	and
T264	126-133	JJ	denotes	unequal
T266	134-144	NN	denotes	expression
T265	145-151	NNS	denotes	levels
T267	151-152	sentence	denotes
T268	611-621	sentence	denotes	Background
T269	611-621	NN	denotes	Background
T270	621-703	sentence	denotes	The olfactory receptor gene family is one of the largest in the mammalian genome.
T271	622-625	DT	denotes	The
T273	626-635	JJ	denotes	olfactory
T274	636-644	NN	denotes	receptor
T275	645-649	NN	denotes	gene
T272	650-656	NN	denotes	family
T276	657-659	VBZ	denotes	is
T277	660-663	CD	denotes	one
T278	664-666	IN	denotes	of
T279	667-670	DT	denotes	the
T280	671-678	JJS	denotes	largest
T281	679-681	IN	denotes	in
T282	682-685	DT	denotes	the
T284	686-695	JJ	denotes	mammalian
T283	696-702	NN	denotes	genome
T285	702-703	.	denotes	.
T286	703-901	sentence	denotes	Previous computational analyses have identified approximately 1,500 mouse olfactory receptors, but experimental evidence confirming olfactory function is available for very few olfactory receptors.
T287	704-712	JJ	denotes	Previous
T289	713-726	JJ	denotes	computational
T288	727-735	NNS	denotes	analyses
T291	736-740	VBP	denotes	have
T290	741-751	VBN	denotes	identified
T292	752-765	RB	denotes	approximately
T293	766-771	CD	denotes	1,500
T295	772-777	NN	denotes	mouse
T296	778-787	JJ	denotes	olfactory
T294	788-797	NNS	denotes	receptors
T297	797-799	,	denotes	,
T298	799-802	CC	denotes	but
T299	803-815	JJ	denotes	experimental
T300	816-824	NN	denotes	evidence
T302	825-835	VBG	denotes	confirming
T303	836-845	JJ	denotes	olfactory
T304	846-854	NN	denotes	function
T301	855-857	VBZ	denotes	is
T305	858-867	JJ	denotes	available
T306	868-871	IN	denotes	for
T307	872-876	RB	denotes	very
T308	877-880	JJ	denotes	few
T310	881-890	JJ	denotes	olfactory
T309	891-900	NNS	denotes	receptors
T311	900-901	.	denotes	.
T312	901-1170	sentence	denotes	We therefore screened a mouse olfactory epithelium cDNA library to obtain olfactory receptor expressed sequence tags, providing evidence of olfactory function for many additional olfactory receptors, as well as identifying gene structure and putative promoter regions.
T313	902-904	PRP	denotes	We
T315	905-914	RB	denotes	therefore
T314	915-923	VBD	denotes	screened
T316	924-925	DT	denotes	a
T318	926-931	NN	denotes	mouse
T320	932-941	JJ	denotes	olfactory
T319	942-952	NN	denotes	epithelium
T321	953-957	NN	denotes	cDNA
T317	958-965	NN	denotes	library
T322	966-968	TO	denotes	to
T323	969-975	VB	denotes	obtain
T324	976-985	JJ	denotes	olfactory
T325	986-994	NN	denotes	receptor
T326	995-1004	VBN	denotes	expressed
T328	1005-1013	NN	denotes	sequence
T327	1014-1018	NNS	denotes	tags
T329	1018-1020	,	denotes	,
T330	1020-1029	VBG	denotes	providing
T331	1030-1038	NN	denotes	evidence
T332	1039-1041	IN	denotes	of
T333	1042-1051	JJ	denotes	olfactory
T334	1052-1060	NN	denotes	function
T335	1061-1064	IN	denotes	for
T336	1065-1069	JJ	denotes	many
T338	1070-1080	JJ	denotes	additional
T339	1081-1090	JJ	denotes	olfactory
T337	1091-1100	NNS	denotes	receptors
T340	1100-1102	,	denotes	,
T341	1102-1104	RB	denotes	as
T343	1105-1109	RB	denotes	well
T342	1110-1112	IN	denotes	as
T344	1113-1124	VBG	denotes	identifying
T345	1125-1129	NN	denotes	gene
T346	1130-1139	NN	denotes	structure
T347	1140-1143	CC	denotes	and
T348	1144-1152	JJ	denotes	putative
T350	1153-1161	NN	denotes	promoter
T349	1162-1169	NNS	denotes	regions
T351	1169-1170	.	denotes	.
T352	1170-1179	sentence	denotes	Results
T353	1172-1179	NNS	denotes	Results
T354	1179-1266	sentence	denotes	We identified more than 1,200 odorant receptor cDNAs representing more than 400 genes.
T355	1180-1182	PRP	denotes	We
T356	1183-1193	VBD	denotes	identified
T357	1194-1198	JJR	denotes	more
T359	1199-1203	IN	denotes	than
T358	1204-1209	CD	denotes	1,200
T361	1210-1217	JJ	denotes	odorant
T362	1218-1226	NN	denotes	receptor
T360	1227-1232	NNS	denotes	cDNAs
T363	1233-1245	VBG	denotes	representing
T364	1246-1250	JJR	denotes	more
T366	1251-1255	IN	denotes	than
T365	1256-1259	CD	denotes	400
T367	1260-1265	NNS	denotes	genes
T368	1265-1266	.	denotes	.
T369	1266-1468	sentence	denotes	Using real-time PCR to confirm expression level differences suggested by our screen, we find that transcript levels in the olfactory epithelium can differ between olfactory receptors by up to 300-fold.
T370	1267-1272	VBG	denotes	Using
T372	1273-1277	JJ	denotes	real
T374	1277-1278	HYPH	denotes	-
T373	1278-1282	NN	denotes	time
T375	1283-1286	NN	denotes	PCR
T376	1287-1289	TO	denotes	to
T377	1290-1297	VB	denotes	confirm
T378	1298-1308	NN	denotes	expression
T379	1309-1314	NN	denotes	level
T380	1315-1326	NNS	denotes	differences
T381	1327-1336	VBN	denotes	suggested
T382	1337-1339	IN	denotes	by
T383	1340-1343	PRP$	denotes	our
T384	1344-1350	NN	denotes	screen
T385	1350-1352	,	denotes	,
T386	1352-1354	PRP	denotes	we
T371	1355-1359	VBP	denotes	find
T387	1360-1364	IN	denotes	that
T389	1365-1375	NN	denotes	transcript
T390	1376-1382	NNS	denotes	levels
T391	1383-1385	IN	denotes	in
T392	1386-1389	DT	denotes	the
T394	1390-1399	JJ	denotes	olfactory
T393	1400-1410	NN	denotes	epithelium
T395	1411-1414	MD	denotes	can
T388	1415-1421	VB	denotes	differ
T396	1422-1429	IN	denotes	between
T397	1430-1439	JJ	denotes	olfactory
T398	1440-1449	NNS	denotes	receptors
T399	1450-1452	IN	denotes	by
T400	1453-1455	RB	denotes	up
T402	1456-1458	IN	denotes	to
T401	1459-1467	RB	denotes	300-fold
T403	1467-1468	.	denotes	.
T404	1468-1612	sentence	denotes	Differences for one gene pair are apparently due to both unequal numbers of expressing cells and unequal transcript levels per expressing cell.
T405	1469-1480	NNS	denotes	Differences
T407	1481-1484	IN	denotes	for
T408	1485-1488	CD	denotes	one
T410	1489-1493	NN	denotes	gene
T409	1494-1498	NN	denotes	pair
T406	1499-1502	VBP	denotes	are
T411	1503-1513	RB	denotes	apparently
T412	1514-1517	IN	denotes	due
T413	1518-1520	IN	denotes	to
T414	1521-1525	CC	denotes	both
T416	1526-1533	JJ	denotes	unequal
T415	1534-1541	NNS	denotes	numbers
T417	1542-1544	IN	denotes	of
T418	1545-1555	VBG	denotes	expressing
T419	1556-1561	NNS	denotes	cells
T420	1562-1565	CC	denotes	and
T421	1566-1573	JJ	denotes	unequal
T423	1574-1584	NN	denotes	transcript
T422	1585-1591	NNS	denotes	levels
T424	1592-1595	IN	denotes	per
T425	1596-1606	VBG	denotes	expressing
T426	1607-1611	NN	denotes	cell
T427	1611-1612	.	denotes	.
T428	1612-1764	sentence	denotes	At least two-thirds of olfactory receptors exhibit multiple transcriptional variants, with alternative isoforms of both 5' and 3' untranslated regions.
T429	1613-1615	RB	denotes	At
T431	1616-1621	RBS	denotes	least
T430	1622-1625	CD	denotes	two
T433	1625-1626	HYPH	denotes	-
T434	1626-1632	NNS	denotes	thirds
T435	1633-1635	IN	denotes	of
T436	1636-1645	JJ	denotes	olfactory
T437	1646-1655	NNS	denotes	receptors
T432	1656-1663	VBP	denotes	exhibit
T438	1664-1672	JJ	denotes	multiple
T440	1673-1688	JJ	denotes	transcriptional
T439	1689-1697	NNS	denotes	variants
T441	1697-1699	,	denotes	,
T442	1699-1703	IN	denotes	with
T443	1704-1715	JJ	denotes	alternative
T444	1716-1724	NNS	denotes	isoforms
T445	1725-1727	IN	denotes	of
T446	1728-1732	CC	denotes	both
T447	1733-1734	CD	denotes	5
T449	1734-1735	SYM	denotes	'
T450	1736-1739	CC	denotes	and
T451	1740-1741	CD	denotes	3
T452	1741-1742	SYM	denotes	'
T453	1743-1755	JJ	denotes	untranslated
T448	1756-1763	NNS	denotes	regions
T454	1763-1764	.	denotes	.
T455	1764-1896	sentence	denotes	Some transcripts (5%) utilize splice sites within the coding region, contrary to the stereotyped olfactory receptor gene structure.
T456	1765-1769	DT	denotes	Some
T457	1770-1781	NNS	denotes	transcripts
T459	1782-1783	-LRB-	denotes	(
T461	1783-1784	CD	denotes	5
T460	1784-1785	NN	denotes	%
T462	1785-1786	-RRB-	denotes	)
T458	1787-1794	VBP	denotes	utilize
T463	1795-1801	NN	denotes	splice
T464	1802-1807	NNS	denotes	sites
T465	1808-1814	IN	denotes	within
T466	1815-1818	DT	denotes	the
T468	1819-1825	VBG	denotes	coding
T467	1826-1832	NN	denotes	region
T469	1832-1834	,	denotes	,
T470	1834-1842	JJ	denotes	contrary
T471	1843-1845	IN	denotes	to
T472	1846-1849	DT	denotes	the
T474	1850-1861	VBN	denotes	stereotyped
T475	1862-1871	JJ	denotes	olfactory
T476	1872-1880	NN	denotes	receptor
T477	1881-1885	NN	denotes	gene
T473	1886-1895	NN	denotes	structure
T478	1895-1896	.	denotes	.
T479	1896-2014	sentence	denotes	Most atypical transcripts encode nonfunctional olfactory receptors, but can occasionally increase receptor diversity.
T480	1897-1901	JJS	denotes	Most
T482	1902-1910	JJ	denotes	atypical
T481	1911-1922	NNS	denotes	transcripts
T483	1923-1929	VBP	denotes	encode
T484	1930-1943	JJ	denotes	nonfunctional
T485	1944-1953	JJ	denotes	olfactory
T486	1954-1963	NNS	denotes	receptors
T487	1963-1965	,	denotes	,
T488	1965-1968	CC	denotes	but
T489	1969-1972	MD	denotes	can
T491	1973-1985	RB	denotes	occasionally
T490	1986-1994	VB	denotes	increase
T492	1995-2003	NN	denotes	receptor
T493	2004-2013	NN	denotes	diversity
T494	2013-2014	.	denotes	.
T495	2014-2027	sentence	denotes	Conclusions
T496	2016-2027	NNS	denotes	Conclusions
T497	2027-2134	sentence	denotes	Our cDNA collection confirms olfactory function of over one-third of the intact mouse olfactory receptors.
T498	2028-2031	PRP$	denotes	Our
T500	2032-2036	NN	denotes	cDNA
T499	2037-2047	NN	denotes	collection
T501	2048-2056	VBZ	denotes	confirms
T502	2057-2066	JJ	denotes	olfactory
T503	2067-2075	NN	denotes	function
T504	2076-2078	IN	denotes	of
T505	2079-2083	IN	denotes	over
T506	2084-2087	CD	denotes	one
T507	2087-2088	HYPH	denotes	-
T508	2088-2093	NN	denotes	third
T509	2094-2096	IN	denotes	of
T510	2097-2100	DT	denotes	the
T512	2101-2107	JJ	denotes	intact
T513	2108-2113	NN	denotes	mouse
T514	2114-2123	JJ	denotes	olfactory
T511	2124-2133	NNS	denotes	receptors
T515	2133-2134	.	denotes	.
T516	2134-2240	sentence	denotes	Most of these genes were previously annotated as olfactory receptors based solely on sequence similarity.
T517	2135-2139	JJS	denotes	Most
T519	2140-2142	IN	denotes	of
T520	2143-2148	DT	denotes	these
T521	2149-2154	NNS	denotes	genes
T522	2155-2159	VBD	denotes	were
T523	2160-2170	RB	denotes	previously
T518	2171-2180	VBN	denotes	annotated
T524	2181-2183	IN	denotes	as
T525	2184-2193	JJ	denotes	olfactory
T526	2194-2203	NNS	denotes	receptors
T527	2204-2209	VBN	denotes	based
T528	2210-2216	RB	denotes	solely
T529	2217-2219	IN	denotes	on
T530	2220-2228	NN	denotes	sequence
T531	2229-2239	NN	denotes	similarity
T532	2239-2240	.	denotes	.
T533	2240-2407	sentence	denotes	Our finding that different olfactory receptors have different expression levels is intriguing given the one-neuron, one-gene expression regime of olfactory receptors.
T534	2241-2244	PRP$	denotes	Our
T535	2245-2252	NN	denotes	finding
T537	2253-2257	IN	denotes	that
T539	2258-2267	JJ	denotes	different
T541	2268-2277	JJ	denotes	olfactory
T540	2278-2287	NNS	denotes	receptors
T538	2288-2292	VBP	denotes	have
T542	2293-2302	JJ	denotes	different
T544	2303-2313	NN	denotes	expression
T543	2314-2320	NNS	denotes	levels
T536	2321-2323	VBZ	denotes	is
T545	2324-2334	JJ	denotes	intriguing
T546	2335-2340	VBN	denotes	given
T547	2341-2344	DT	denotes	the
T549	2345-2348	CD	denotes	one
T551	2348-2349	HYPH	denotes	-
T550	2349-2355	NN	denotes	neuron
T552	2355-2357	,	denotes	,
T553	2357-2360	CD	denotes	one
T555	2360-2361	HYPH	denotes	-
T554	2361-2365	NN	denotes	gene
T556	2366-2376	NN	denotes	expression
T548	2377-2383	NN	denotes	regime
T557	2384-2386	IN	denotes	of
T558	2387-2396	JJ	denotes	olfactory
T559	2397-2406	NNS	denotes	receptors
T560	2406-2407	.	denotes	.
T561	2407-2664	sentence	denotes	We provide 5' untranslated region sequences and candidate promoter regions for more than 300 olfactory receptors, valuable resources for computational regulatory motif searches and for designing olfactory receptor microarrays and other experimental probes.
T562	2408-2410	PRP	denotes	We
T563	2411-2418	VBP	denotes	provide
T564	2419-2420	CD	denotes	5
T566	2420-2421	SYM	denotes	'
T567	2422-2434	JJ	denotes	untranslated
T565	2435-2441	NN	denotes	region
T568	2442-2451	NNS	denotes	sequences
T569	2452-2455	CC	denotes	and
T570	2456-2465	NN	denotes	candidate
T572	2466-2474	NN	denotes	promoter
T571	2475-2482	NNS	denotes	regions
T573	2483-2486	IN	denotes	for
T574	2487-2491	JJR	denotes	more
T576	2492-2496	IN	denotes	than
T575	2497-2500	CD	denotes	300
T578	2501-2510	JJ	denotes	olfactory
T577	2511-2520	NNS	denotes	receptors
T579	2520-2522	,	denotes	,
T580	2522-2530	JJ	denotes	valuable
T581	2531-2540	NNS	denotes	resources
T582	2541-2544	IN	denotes	for
T583	2545-2558	JJ	denotes	computational
T585	2559-2569	JJ	denotes	regulatory
T584	2570-2575	NN	denotes	motif
T586	2576-2584	NNS	denotes	searches
T587	2585-2588	CC	denotes	and
T588	2589-2592	IN	denotes	for
T589	2593-2602	VBG	denotes	designing
T590	2603-2612	JJ	denotes	olfactory
T591	2613-2621	NN	denotes	receptor
T592	2622-2633	NNS	denotes	microarrays
T593	2634-2637	CC	denotes	and
T594	2638-2643	JJ	denotes	other
T596	2644-2656	JJ	denotes	experimental
T595	2657-2663	NNS	denotes	probes
T597	2663-2664	.	denotes	.
T1203	2677-2680	DT	denotes	The
T1204	2681-2692	NN	denotes	interaction
T1206	2693-2695	IN	denotes	of
T1207	2696-2705	JJ	denotes	olfactory
T1209	2706-2707	-LRB-	denotes	(
T1210	2707-2709	CC	denotes	or
T1211	2710-2717	NN	denotes	odorant
T1212	2717-2718	-RRB-	denotes	)
T1208	2719-2728	NNS	denotes	receptors
T1213	2729-2733	IN	denotes	with
T1214	2734-2739	PRP$	denotes	their
T1216	2740-2747	NN	denotes	odorant
T1215	2748-2755	NNS	denotes	ligands
T1205	2756-2758	VBZ	denotes	is
T1217	2759-2762	DT	denotes	the
T1219	2763-2768	JJ	denotes	first
T1218	2769-2773	NN	denotes	step
T1220	2774-2776	IN	denotes	in
T1221	2777-2778	DT	denotes	a
T1223	2779-2785	NN	denotes	signal
T1224	2786-2798	NN	denotes	transduction
T1222	2799-2806	NN	denotes	pathway
T1225	2807-2811	WDT	denotes	that
T1226	2812-2819	VBZ	denotes	results
T1227	2820-2822	IN	denotes	in
T1228	2823-2826	DT	denotes	the
T1229	2827-2837	NN	denotes	perception
T1230	2838-2840	IN	denotes	of
T1231	2841-2846	NN	denotes	smell
T1232	2846-2847	.	denotes	.
T1233	2847-2987	sentence	denotes	The olfactory receptor gene family is one of the largest in the mammalian genome, comprising about 1,500 members in the mouse genome [1,2].
T1234	2848-2851	DT	denotes	The
T1236	2852-2861	JJ	denotes	olfactory
T1237	2862-2870	NN	denotes	receptor
T1238	2871-2875	NN	denotes	gene
T1235	2876-2882	NN	denotes	family
T1239	2883-2885	VBZ	denotes	is
T1240	2886-2889	CD	denotes	one
T1241	2890-2892	IN	denotes	of
T1242	2893-2896	DT	denotes	the
T1243	2897-2904	JJS	denotes	largest
T1244	2905-2907	IN	denotes	in
T1245	2908-2911	DT	denotes	the
T1247	2912-2921	JJ	denotes	mammalian
T1246	2922-2928	NN	denotes	genome
T1248	2928-2930	,	denotes	,
T1249	2930-2940	VBG	denotes	comprising
T1250	2941-2946	IN	denotes	about
T1251	2947-2952	CD	denotes	1,500
T1252	2953-2960	NNS	denotes	members
T1253	2961-2963	IN	denotes	in
T1254	2964-2967	DT	denotes	the
T1256	2968-2973	NN	denotes	mouse
T1255	2974-2980	NN	denotes	genome
T1257	2981-2982	-LRB-	denotes	[
T1259	2982-2983	CD	denotes	1
T1260	2983-2984	,	denotes	,
T1258	2984-2985	CD	denotes	2
T1261	2985-2986	-RRB-	denotes	]
T1262	2986-2987	.	denotes	.
T1263	2987-3242	sentence	denotes	Olfactory receptors were originally identified in an elegant experiment based on the hypothesis that they would be seven-transmembrane-domain proteins encoded by a large, diverse gene family whose expression is restricted to the olfactory epithelium [3].
T1264	2988-2997	JJ	denotes	Olfactory
T1265	2998-3007	NNS	denotes	receptors
T1267	3008-3012	VBD	denotes	were
T1268	3013-3023	RB	denotes	originally
T1266	3024-3034	VBN	denotes	identified
T1269	3035-3037	IN	denotes	in
T1270	3038-3040	DT	denotes	an
T1272	3041-3048	JJ	denotes	elegant
T1271	3049-3059	NN	denotes	experiment
T1273	3060-3065	VBN	denotes	based
T1274	3066-3068	IN	denotes	on
T1275	3069-3072	DT	denotes	the
T1276	3073-3083	NN	denotes	hypothesis
T1277	3084-3088	IN	denotes	that
T1279	3089-3093	PRP	denotes	they
T1280	3094-3099	MD	denotes	would
T1278	3100-3102	VB	denotes	be
T1281	3103-3108	CD	denotes	seven
T1283	3108-3109	HYPH	denotes	-
T1282	3109-3122	NN	denotes	transmembrane
T1285	3122-3123	HYPH	denotes	-
T1284	3123-3129	NN	denotes	domain
T1286	3130-3138	NN	denotes	proteins
T1287	3139-3146	VBN	denotes	encoded
T1288	3147-3149	IN	denotes	by
T1289	3150-3151	DT	denotes	a
T1291	3152-3157	JJ	denotes	large
T1292	3157-3159	,	denotes	,
T1293	3159-3166	JJ	denotes	diverse
T1294	3167-3171	NN	denotes	gene
T1290	3172-3178	NN	denotes	family
T1295	3179-3184	WP$	denotes	whose
T1296	3185-3195	NN	denotes	expression
T1298	3196-3198	VBZ	denotes	is
T1297	3199-3209	VBN	denotes	restricted
T1299	3210-3212	IN	denotes	to
T1300	3213-3216	DT	denotes	the
T1302	3217-3226	JJ	denotes	olfactory
T1301	3227-3237	NN	denotes	epithelium
T1303	3238-3239	-LRB-	denotes	[
T1304	3239-3240	CD	denotes	3
T1305	3240-3241	-RRB-	denotes	]
T1306	3241-3242	.	denotes	.
T1307	3242-3425	sentence	denotes	Subsequent studies have shown that some of these receptors do indeed respond to odorants and can confer that responsivity when expressed in heterologous cell types (for example [4]).
T1308	3243-3253	JJ	denotes	Subsequent
T1309	3254-3261	NNS	denotes	studies
T1311	3262-3266	VBP	denotes	have
T1310	3267-3272	VBN	denotes	shown
T1312	3273-3277	IN	denotes	that
T1314	3278-3282	DT	denotes	some
T1315	3283-3285	IN	denotes	of
T1316	3286-3291	DT	denotes	these
T1317	3292-3301	NNS	denotes	receptors
T1318	3302-3304	VBP	denotes	do
T1319	3305-3311	RB	denotes	indeed
T1313	3312-3319	VB	denotes	respond
T1320	3320-3322	IN	denotes	to
T1321	3323-3331	NNS	denotes	odorants
T1322	3332-3335	CC	denotes	and
T1323	3336-3339	MD	denotes	can
T1324	3340-3346	VB	denotes	confer
T1325	3347-3351	DT	denotes	that
T1326	3352-3364	NN	denotes	responsivity
T1327	3365-3369	WRB	denotes	when
T1328	3370-3379	VBN	denotes	expressed
T1329	3380-3382	IN	denotes	in
T1330	3383-3395	JJ	denotes	heterologous
T1332	3396-3400	NN	denotes	cell
T1331	3401-3406	NNS	denotes	types
T1333	3407-3408	-LRB-	denotes	(
T1334	3408-3411	IN	denotes	for
T1336	3412-3419	NN	denotes	example
T1337	3420-3421	-LRB-	denotes	[
T1335	3421-3422	CD	denotes	4
T1338	3422-3423	-RRB-	denotes	]
T1339	3423-3424	-RRB-	denotes	)
T1340	3424-3425	.	denotes	.
T1341	3425-3557	sentence	denotes	Recent computational investigations have provided the almost complete human [5,6] and mouse [1,2] olfactory receptor-gene catalogs.
T1342	3426-3432	JJ	denotes	Recent
T1344	3433-3446	JJ	denotes	computational
T1343	3447-3461	NNS	denotes	investigations
T1346	3462-3466	VBP	denotes	have
T1345	3467-3475	VBN	denotes	provided
T1347	3476-3479	DT	denotes	the
T1349	3480-3486	RB	denotes	almost
T1350	3487-3495	JJ	denotes	complete
T1351	3496-3501	JJ	denotes	human
T1353	3502-3503	-LRB-	denotes	[
T1354	3503-3504	CD	denotes	5
T1355	3504-3505	,	denotes	,
T1352	3505-3506	CD	denotes	6
T1357	3506-3507	-RRB-	denotes	]
T1358	3508-3511	CC	denotes	and
T1359	3512-3517	NN	denotes	mouse
T1361	3518-3519	-LRB-	denotes	[
T1362	3519-3520	CD	denotes	1
T1363	3520-3521	,	denotes	,
T1360	3521-3522	CD	denotes	2
T1364	3522-3523	-RRB-	denotes	]
T1365	3524-3533	JJ	denotes	olfactory
T1366	3534-3542	NN	denotes	receptor
T1367	3542-3543	HYPH	denotes	-
T1356	3543-3547	NN	denotes	gene
T1348	3548-3556	NNS	denotes	catalogs
T1368	3556-3557	.	denotes	.
T1369	3557-3854	sentence	denotes	However, the assignment of most of these genes as olfactory receptors is based solely on similarity to one of a relatively small number of experimentally confirmed mouse olfactory receptors or, worse, on similarity to a gene that in turn was defined as an olfactory receptor solely by similarity.
T1370	3558-3565	RB	denotes	However
T1372	3565-3567	,	denotes	,
T1373	3567-3570	DT	denotes	the
T1374	3571-3581	NN	denotes	assignment
T1375	3582-3584	IN	denotes	of
T1376	3585-3589	JJS	denotes	most
T1377	3590-3592	IN	denotes	of
T1378	3593-3598	DT	denotes	these
T1379	3599-3604	NNS	denotes	genes
T1380	3605-3607	IN	denotes	as
T1381	3608-3617	JJ	denotes	olfactory
T1382	3618-3627	NNS	denotes	receptors
T1383	3628-3630	VBZ	denotes	is
T1371	3631-3636	VBN	denotes	based
T1384	3637-3643	RB	denotes	solely
T1385	3644-3646	IN	denotes	on
T1386	3647-3657	NN	denotes	similarity
T1387	3658-3660	IN	denotes	to
T1388	3661-3664	CD	denotes	one
T1389	3665-3667	IN	denotes	of
T1390	3668-3669	DT	denotes	a
T1392	3670-3680	RB	denotes	relatively
T1393	3681-3686	JJ	denotes	small
T1391	3687-3693	NN	denotes	number
T1394	3694-3696	IN	denotes	of
T1395	3697-3711	RB	denotes	experimentally
T1396	3712-3721	VBN	denotes	confirmed
T1398	3722-3727	NN	denotes	mouse
T1399	3728-3737	JJ	denotes	olfactory
T1397	3738-3747	NNS	denotes	receptors
T1400	3748-3750	CC	denotes	or
T1401	3750-3752	,	denotes	,
T1402	3752-3757	RBR	denotes	worse
T1404	3757-3759	,	denotes	,
T1403	3759-3761	IN	denotes	on
T1405	3762-3772	NN	denotes	similarity
T1406	3773-3775	IN	denotes	to
T1407	3776-3777	DT	denotes	a
T1408	3778-3782	NN	denotes	gene
T1409	3783-3787	WDT	denotes	that
T1411	3788-3790	IN	denotes	in
T1412	3791-3795	NN	denotes	turn
T1413	3796-3799	VBD	denotes	was
T1410	3800-3807	VBN	denotes	defined
T1414	3808-3810	IN	denotes	as
T1415	3811-3813	DT	denotes	an
T1417	3814-3823	JJ	denotes	olfactory
T1416	3824-3832	NN	denotes	receptor
T1418	3833-3839	RB	denotes	solely
T1419	3840-3842	IN	denotes	by
T1420	3843-3853	NN	denotes	similarity
T1421	3853-3854	.	denotes	.
T1422	3854-4157	sentence	denotes	While similarity-based genome annotation is a good initial method to identify genes and predict their function, in some cases it can be misleading, as genes of similar sequence can carry out different functions and be expressed in different tissues (for example, the sugar transporter gene family [7]).
T1423	3855-3860	IN	denotes	While
T1425	3861-3871	NN	denotes	similarity
T1427	3871-3872	HYPH	denotes	-
T1426	3872-3877	VBN	denotes	based
T1429	3878-3884	NN	denotes	genome
T1428	3885-3895	NN	denotes	annotation
T1424	3896-3898	VBZ	denotes	is
T1431	3899-3900	DT	denotes	a
T1433	3901-3905	JJ	denotes	good
T1434	3906-3913	JJ	denotes	initial
T1432	3914-3920	NN	denotes	method
T1435	3921-3923	TO	denotes	to
T1436	3924-3932	VB	denotes	identify
T1437	3933-3938	NNS	denotes	genes
T1438	3939-3942	CC	denotes	and
T1439	3943-3950	VBP	denotes	predict
T1440	3951-3956	PRP$	denotes	their
T1441	3957-3965	NN	denotes	function
T1442	3965-3967	,	denotes	,
T1443	3967-3969	IN	denotes	in
T1444	3970-3974	DT	denotes	some
T1445	3975-3980	NNS	denotes	cases
T1446	3981-3983	PRP	denotes	it
T1447	3984-3987	MD	denotes	can
T1430	3988-3990	VB	denotes	be
T1448	3991-4001	JJ	denotes	misleading
T1449	4001-4003	,	denotes	,
T1450	4003-4005	IN	denotes	as
T1452	4006-4011	NNS	denotes	genes
T1453	4012-4014	IN	denotes	of
T1454	4015-4022	JJ	denotes	similar
T1455	4023-4031	NN	denotes	sequence
T1456	4032-4035	MD	denotes	can
T1451	4036-4041	VB	denotes	carry
T1457	4042-4045	RP	denotes	out
T1458	4046-4055	JJ	denotes	different
T1459	4056-4065	NNS	denotes	functions
T1460	4066-4069	CC	denotes	and
T1461	4070-4072	VB	denotes	be
T1462	4073-4082	VBN	denotes	expressed
T1463	4083-4085	IN	denotes	in
T1464	4086-4095	JJ	denotes	different
T1465	4096-4103	NNS	denotes	tissues
T1466	4104-4105	-LRB-	denotes	(
T1467	4105-4108	IN	denotes	for
T1469	4109-4116	NN	denotes	example
T1470	4116-4118	,	denotes	,
T1471	4118-4121	DT	denotes	the
T1472	4122-4127	NN	denotes	sugar
T1473	4128-4139	NN	denotes	transporter
T1474	4140-4144	NN	denotes	gene
T1468	4145-4151	NN	denotes	family
T1475	4152-4153	-LRB-	denotes	[
T1476	4153-4154	CD	denotes	7
T1477	4154-4155	-RRB-	denotes	]
T1478	4155-4156	-RRB-	denotes	)
T1479	4156-4157	.	denotes	.
T1480	4157-4380	sentence	denotes	A small subset of olfactory receptors appears to be expressed in non-olfactory tissues, principally the testis [8], but also taste tissues [9], prostate [10], erythroid cells [11], notochord [12] and perhaps other tissues.
T1481	4158-4159	DT	denotes	A
T1483	4160-4165	JJ	denotes	small
T1482	4166-4172	NN	denotes	subset
T1485	4173-4175	IN	denotes	of
T1486	4176-4185	JJ	denotes	olfactory
T1487	4186-4195	NNS	denotes	receptors
T1484	4196-4203	VBZ	denotes	appears
T1488	4204-4206	TO	denotes	to
T1490	4207-4209	VB	denotes	be
T1489	4210-4219	VBN	denotes	expressed
T1491	4220-4222	IN	denotes	in
T1492	4223-4236	JJ	denotes	non-olfactory
T1493	4237-4244	NNS	denotes	tissues
T1494	4244-4246	,	denotes	,
T1495	4246-4257	RB	denotes	principally
T1497	4258-4261	DT	denotes	the
T1496	4262-4268	NN	denotes	testis
T1498	4269-4270	-LRB-	denotes	[
T1499	4270-4271	CD	denotes	8
T1500	4271-4272	-RRB-	denotes	]
T1501	4272-4274	,	denotes	,
T1502	4274-4277	CC	denotes	but
T1503	4278-4282	RB	denotes	also
T1504	4283-4288	NN	denotes	taste
T1505	4289-4296	NNS	denotes	tissues
T1506	4297-4298	-LRB-	denotes	[
T1507	4298-4299	CD	denotes	9
T1508	4299-4300	-RRB-	denotes	]
T1509	4300-4302	,	denotes	,
T1510	4302-4310	NN	denotes	prostate
T1511	4311-4312	-LRB-	denotes	[
T1512	4312-4314	CD	denotes	10
T1513	4314-4315	-RRB-	denotes	]
T1514	4315-4317	,	denotes	,
T1515	4317-4326	JJ	denotes	erythroid
T1516	4327-4332	NNS	denotes	cells
T1517	4333-4334	-LRB-	denotes	[
T1518	4334-4336	CD	denotes	11
T1519	4336-4337	-RRB-	denotes	]
T1520	4337-4339	,	denotes	,
T1521	4339-4348	NN	denotes	notochord
T1522	4349-4350	-LRB-	denotes	[
T1523	4350-4352	CD	denotes	12
T1524	4352-4353	-RRB-	denotes	]
T1525	4354-4357	CC	denotes	and
T1526	4358-4365	RB	denotes	perhaps
T1528	4366-4371	JJ	denotes	other
T1527	4372-4379	NNS	denotes	tissues
T1529	4379-4380	.	denotes	.
T1530	4380-4526	sentence	denotes	Expression in the testis has led some investigators to suggest that a subset of olfactory receptors may function as spermatid chemoreceptors [8].
T1531	4381-4391	NN	denotes	Expression
T1533	4392-4394	IN	denotes	in
T1534	4395-4398	DT	denotes	the
T1535	4399-4405	NN	denotes	testis
T1536	4406-4409	VBZ	denotes	has
T1532	4410-4413	VBN	denotes	led
T1537	4414-4418	DT	denotes	some
T1538	4419-4432	NNS	denotes	investigators
T1539	4433-4435	TO	denotes	to
T1540	4436-4443	VB	denotes	suggest
T1541	4444-4448	IN	denotes	that
T1543	4449-4450	DT	denotes	a
T1544	4451-4457	NN	denotes	subset
T1545	4458-4460	IN	denotes	of
T1546	4461-4470	JJ	denotes	olfactory
T1547	4471-4480	NNS	denotes	receptors
T1548	4481-4484	MD	denotes	may
T1542	4485-4493	VB	denotes	function
T1549	4494-4496	IN	denotes	as
T1550	4497-4506	JJ	denotes	spermatid
T1551	4507-4521	NNS	denotes	chemoreceptors
T1552	4522-4523	-LRB-	denotes	[
T1553	4523-4524	CD	denotes	8
T1554	4524-4525	-RRB-	denotes	]
T1555	4525-4526	.	denotes	.
T1556	4526-4654	sentence	denotes	Recent studies of one human testis-expressed olfactory receptor indicate that it does indeed function in sperm chemotaxis [13].
T1557	4527-4533	JJ	denotes	Recent
T1558	4534-4541	NNS	denotes	studies
T1560	4542-4544	IN	denotes	of
T1561	4545-4548	CD	denotes	one
T1563	4549-4554	JJ	denotes	human
T1564	4555-4561	NN	denotes	testis
T1566	4561-4562	HYPH	denotes	-
T1565	4562-4571	VBN	denotes	expressed
T1567	4572-4581	JJ	denotes	olfactory
T1562	4582-4590	NN	denotes	receptor
T1559	4591-4599	VBP	denotes	indicate
T1568	4600-4604	IN	denotes	that
T1570	4605-4607	PRP	denotes	it
T1571	4608-4612	VBZ	denotes	does
T1572	4613-4619	RB	denotes	indeed
T1569	4620-4628	VB	denotes	function
T1573	4629-4631	IN	denotes	in
T1574	4632-4637	NN	denotes	sperm
T1575	4638-4648	NN	denotes	chemotaxis
T1576	4649-4650	-LRB-	denotes	[
T1577	4650-4652	CD	denotes	13
T1578	4652-4653	-RRB-	denotes	]
T1579	4653-4654	.	denotes	.
T1580	4654-4956	sentence	denotes	Due to the paucity of experimental evidence of the olfactory function of most genes in the family and suggestions of extra-olfactory roles, we embarked on an olfactory receptor expressed sequence tag (EST) project to confirm olfactory epithelial expression of hundreds of mouse odorant receptor genes.
T1581	4655-4658	IN	denotes	Due
T1583	4659-4661	IN	denotes	to
T1584	4662-4665	DT	denotes	the
T1585	4666-4673	NN	denotes	paucity
T1586	4674-4676	IN	denotes	of
T1587	4677-4689	JJ	denotes	experimental
T1588	4690-4698	NN	denotes	evidence
T1589	4699-4701	IN	denotes	of
T1590	4702-4705	DT	denotes	the
T1592	4706-4715	JJ	denotes	olfactory
T1591	4716-4724	NN	denotes	function
T1593	4725-4727	IN	denotes	of
T1594	4728-4732	JJS	denotes	most
T1595	4733-4738	NNS	denotes	genes
T1596	4739-4741	IN	denotes	in
T1597	4742-4745	DT	denotes	the
T1598	4746-4752	NN	denotes	family
T1599	4753-4756	CC	denotes	and
T1600	4757-4768	NNS	denotes	suggestions
T1601	4769-4771	IN	denotes	of
T1602	4772-4787	JJ	denotes	extra-olfactory
T1603	4788-4793	NNS	denotes	roles
T1604	4793-4795	,	denotes	,
T1605	4795-4797	PRP	denotes	we
T1582	4798-4806	VBD	denotes	embarked
T1606	4807-4809	IN	denotes	on
T1607	4810-4812	DT	denotes	an
T1609	4813-4822	JJ	denotes	olfactory
T1610	4823-4831	NN	denotes	receptor
T1611	4832-4841	VBN	denotes	expressed
T1613	4842-4850	NN	denotes	sequence
T1612	4851-4854	NN	denotes	tag
T1614	4855-4856	-LRB-	denotes	(
T1615	4856-4859	NN	denotes	EST
T1616	4859-4860	-RRB-	denotes	)
T1608	4861-4868	NN	denotes	project
T1617	4869-4871	TO	denotes	to
T1618	4872-4879	VB	denotes	confirm
T1619	4880-4889	JJ	denotes	olfactory
T1621	4890-4900	JJ	denotes	epithelial
T1620	4901-4911	NN	denotes	expression
T1622	4912-4914	IN	denotes	of
T1623	4915-4923	NNS	denotes	hundreds
T1624	4924-4926	IN	denotes	of
T1625	4927-4932	NN	denotes	mouse
T1627	4933-4940	NN	denotes	odorant
T1628	4941-4949	NN	denotes	receptor
T1626	4950-4955	NNS	denotes	genes
T1629	4955-4956	.	denotes	.
T1630	4956-5064	sentence	denotes	Within the olfactory epithelium, individual olfactory receptor genes show an intriguing expression pattern.
T1631	4957-4963	IN	denotes	Within
T1633	4964-4967	DT	denotes	the
T1635	4968-4977	JJ	denotes	olfactory
T1634	4978-4988	NN	denotes	epithelium
T1636	4988-4990	,	denotes	,
T1637	4990-5000	JJ	denotes	individual
T1639	5001-5010	JJ	denotes	olfactory
T1640	5011-5019	NN	denotes	receptor
T1638	5020-5025	NNS	denotes	genes
T1632	5026-5030	VBP	denotes	show
T1641	5031-5033	DT	denotes	an
T1643	5034-5044	JJ	denotes	intriguing
T1644	5045-5055	NN	denotes	expression
T1642	5056-5063	NN	denotes	pattern
T1645	5063-5064	.	denotes	.
T1646	5064-5170	sentence	denotes	Each olfactory receptor is expressed in a subset of cells in one of four zones of the epithelium [14,15].
T1647	5065-5069	DT	denotes	Each
T1649	5070-5079	JJ	denotes	olfactory
T1648	5080-5088	NN	denotes	receptor
T1651	5089-5091	VBZ	denotes	is
T1650	5092-5101	VBN	denotes	expressed
T1652	5102-5104	IN	denotes	in
T1653	5105-5106	DT	denotes	a
T1654	5107-5113	NN	denotes	subset
T1655	5114-5116	IN	denotes	of
T1656	5117-5122	NNS	denotes	cells
T1657	5123-5125	IN	denotes	in
T1658	5126-5129	CD	denotes	one
T1659	5130-5132	IN	denotes	of
T1660	5133-5137	CD	denotes	four
T1661	5138-5143	NNS	denotes	zones
T1662	5144-5146	IN	denotes	of
T1663	5147-5150	DT	denotes	the
T1664	5151-5161	NN	denotes	epithelium
T1665	5162-5163	-LRB-	denotes	[
T1667	5163-5165	CD	denotes	14
T1668	5165-5166	,	denotes	,
T1666	5166-5168	CD	denotes	15
T1669	5168-5169	-RRB-	denotes	]
T1670	5169-5170	.	denotes	.
T1671	5170-5357	sentence	denotes	Furthermore, each olfactory neuron expresses only one allele [16] of a single olfactory receptor gene [17,18], and the remaining approximately 1,499 genes are transcriptionally inactive.
T1672	5171-5182	RB	denotes	Furthermore
T1674	5182-5184	,	denotes	,
T1675	5184-5188	DT	denotes	each
T1677	5189-5198	JJ	denotes	olfactory
T1676	5199-5205	NN	denotes	neuron
T1673	5206-5215	VBZ	denotes	expresses
T1678	5216-5220	RB	denotes	only
T1680	5221-5224	CD	denotes	one
T1679	5225-5231	NN	denotes	allele
T1681	5232-5233	-LRB-	denotes	[
T1682	5233-5235	CD	denotes	16
T1683	5235-5236	-RRB-	denotes	]
T1684	5237-5239	IN	denotes	of
T1685	5240-5241	DT	denotes	a
T1687	5242-5248	JJ	denotes	single
T1688	5249-5258	JJ	denotes	olfactory
T1689	5259-5267	NN	denotes	receptor
T1686	5268-5272	NN	denotes	gene
T1690	5273-5274	-LRB-	denotes	[
T1692	5274-5276	CD	denotes	17
T1693	5276-5277	,	denotes	,
T1691	5277-5279	CD	denotes	18
T1694	5279-5280	-RRB-	denotes	]
T1695	5280-5282	,	denotes	,
T1696	5282-5285	CC	denotes	and
T1697	5286-5289	DT	denotes	the
T1699	5290-5299	NN	denotes	remaining
T1700	5300-5313	RB	denotes	approximately
T1701	5314-5319	CD	denotes	1,499
T1698	5320-5325	NNS	denotes	genes
T1702	5326-5329	VBP	denotes	are
T1703	5330-5347	RB	denotes	transcriptionally
T1704	5348-5356	JJ	denotes	inactive
T1705	5356-5357	.	denotes	.
T1706	5357-5465	sentence	denotes	While the mechanism ensuring singular expression is unknown, many hypotheses have been proposed [14,16,19].
T1707	5358-5363	IN	denotes	While
T1709	5364-5367	DT	denotes	the
T1710	5368-5377	NN	denotes	mechanism
T1711	5378-5386	VBG	denotes	ensuring
T1712	5387-5395	JJ	denotes	singular
T1713	5396-5406	NN	denotes	expression
T1708	5407-5409	VBZ	denotes	is
T1715	5410-5417	JJ	denotes	unknown
T1716	5417-5419	,	denotes	,
T1717	5419-5423	JJ	denotes	many
T1718	5424-5434	NNS	denotes	hypotheses
T1719	5435-5439	VBP	denotes	have
T1720	5440-5444	VBN	denotes	been
T1714	5445-5453	VBN	denotes	proposed
T1721	5454-5455	-LRB-	denotes	[
T1723	5455-5457	CD	denotes	14
T1724	5457-5458	,	denotes	,
T1725	5458-5460	CD	denotes	16
T1726	5460-5461	,	denotes	,
T1722	5461-5463	CD	denotes	19
T1727	5463-5464	-RRB-	denotes	]
T1728	5464-5465	.	denotes	.
T1729	5465-5694	sentence	denotes	In one model, somatic DNA recombination would bring one olfactory receptor gene into a transcriptionally active genomic configuration, as observed for the yeast mating type locus [20] and the mammalian immunoglobulin genes [21].
T1730	5466-5468	IN	denotes	In
T1732	5469-5472	CD	denotes	one
T1733	5473-5478	NN	denotes	model
T1734	5478-5480	,	denotes	,
T1735	5480-5487	JJ	denotes	somatic
T1737	5488-5491	NN	denotes	DNA
T1736	5492-5505	NN	denotes	recombination
T1738	5506-5511	MD	denotes	would
T1731	5512-5517	VB	denotes	bring
T1739	5518-5521	CD	denotes	one
T1741	5522-5531	JJ	denotes	olfactory
T1742	5532-5540	NN	denotes	receptor
T1740	5541-5545	NN	denotes	gene
T1743	5546-5550	IN	denotes	into
T1744	5551-5552	DT	denotes	a
T1746	5553-5570	RB	denotes	transcriptionally
T1747	5571-5577	JJ	denotes	active
T1748	5578-5585	JJ	denotes	genomic
T1745	5586-5599	NN	denotes	configuration
T1749	5599-5601	,	denotes	,
T1750	5601-5603	IN	denotes	as
T1751	5604-5612	VBN	denotes	observed
T1752	5613-5616	IN	denotes	for
T1753	5617-5620	DT	denotes	the
T1755	5621-5626	NN	denotes	yeast
T1756	5627-5633	NN	denotes	mating
T1757	5634-5638	NN	denotes	type
T1754	5639-5644	NN	denotes	locus
T1758	5645-5646	-LRB-	denotes	[
T1759	5646-5648	CD	denotes	20
T1760	5648-5649	-RRB-	denotes	]
T1761	5650-5653	CC	denotes	and
T1762	5654-5657	DT	denotes	the
T1764	5658-5667	JJ	denotes	mammalian
T1765	5668-5682	NN	denotes	immunoglobulin
T1763	5683-5688	NNS	denotes	genes
T1766	5689-5690	-LRB-	denotes	[
T1767	5690-5692	CD	denotes	21
T1768	5692-5693	-RRB-	denotes	]
T1769	5693-5694	.	denotes	.
T1770	5694-5812	sentence	denotes	Alternatively, a second model invokes a combinatorial code of transcription factor binding sites unique to each gene.
T1771	5695-5708	RB	denotes	Alternatively
T1773	5708-5710	,	denotes	,
T1774	5710-5711	DT	denotes	a
T1776	5712-5718	JJ	denotes	second
T1775	5719-5724	NN	denotes	model
T1772	5725-5732	VBZ	denotes	invokes
T1777	5733-5734	DT	denotes	a
T1779	5735-5748	JJ	denotes	combinatorial
T1778	5749-5753	NN	denotes	code
T1780	5754-5756	IN	denotes	of
T1781	5757-5770	NN	denotes	transcription
T1782	5771-5777	NN	denotes	factor
T1784	5778-5785	NN	denotes	binding
T1783	5786-5791	NNS	denotes	sites
T1785	5792-5798	JJ	denotes	unique
T1786	5799-5801	IN	denotes	to
T1787	5802-5806	DT	denotes	each
T1788	5807-5811	NN	denotes	gene
T1789	5811-5812	.	denotes	.
T1790	5812-5950	sentence	denotes	This is unlikely, however, as even olfactory receptor transgenes with identical upstream regions are expressed in different neurons [18].
T1791	5813-5817	DT	denotes	This
T1792	5818-5820	VBZ	denotes	is
T1793	5821-5829	JJ	denotes	unlikely
T1794	5829-5831	,	denotes	,
T1795	5831-5838	RB	denotes	however
T1796	5838-5840	,	denotes	,
T1797	5840-5842	IN	denotes	as
T1799	5843-5847	RB	denotes	even
T1801	5848-5857	JJ	denotes	olfactory
T1802	5858-5866	NN	denotes	receptor
T1800	5867-5877	NNS	denotes	transgenes
T1803	5878-5882	IN	denotes	with
T1804	5883-5892	JJ	denotes	identical
T1806	5893-5901	JJ	denotes	upstream
T1805	5902-5909	NNS	denotes	regions
T1807	5910-5913	VBP	denotes	are
T1798	5914-5923	VBN	denotes	expressed
T1808	5924-5926	IN	denotes	in
T1809	5927-5936	JJ	denotes	different
T1810	5937-5944	NNS	denotes	neurons
T1811	5945-5946	-LRB-	denotes	[
T1812	5946-5948	CD	denotes	18
T1813	5948-5949	-RRB-	denotes	]
T1814	5949-5950	.	denotes	.
T1815	5950-6326	sentence	denotes	In a third model, there would be a limiting quantity of transcription factors - the cell might contain a single transcriptional 'machine' that is capable of accommodating the promoter of only one olfactory receptor gene, similar to the expression site body used by African trypanosomes to ensure singular expression of only one set of variant surface glycoprotein genes [22].
T1816	5951-5953	IN	denotes	In
T1818	5954-5955	DT	denotes	a
T1820	5956-5961	JJ	denotes	third
T1819	5962-5967	NN	denotes	model
T1821	5967-5969	,	denotes	,
T1822	5969-5974	EX	denotes	there
T1824	5975-5980	MD	denotes	would
T1823	5981-5983	VB	denotes	be
T1825	5984-5985	DT	denotes	a
T1827	5986-5994	VBG	denotes	limiting
T1826	5995-6003	NN	denotes	quantity
T1828	6004-6006	IN	denotes	of
T1829	6007-6020	NN	denotes	transcription
T1830	6021-6028	NNS	denotes	factors
T1831	6029-6030	:	denotes	-
T1832	6031-6034	DT	denotes	the
T1833	6035-6039	NN	denotes	cell
T1834	6040-6045	MD	denotes	might
T1817	6046-6053	VB	denotes	contain
T1835	6054-6055	DT	denotes	a
T1837	6056-6062	JJ	denotes	single
T1838	6063-6078	JJ	denotes	transcriptional
T1839	6079-6080	``	denotes	'
T1836	6080-6087	NN	denotes	machine
T1840	6087-6088	''	denotes	'
T1841	6089-6093	WDT	denotes	that
T1842	6094-6096	VBZ	denotes	is
T1843	6097-6104	JJ	denotes	capable
T1844	6105-6107	IN	denotes	of
T1845	6108-6121	VBG	denotes	accommodating
T1846	6122-6125	DT	denotes	the
T1847	6126-6134	NN	denotes	promoter
T1848	6135-6137	IN	denotes	of
T1849	6138-6142	RB	denotes	only
T1851	6143-6146	CD	denotes	one
T1852	6147-6156	JJ	denotes	olfactory
T1853	6157-6165	NN	denotes	receptor
T1850	6166-6170	NN	denotes	gene
T1854	6170-6172	,	denotes	,
T1855	6172-6179	JJ	denotes	similar
T1856	6180-6182	IN	denotes	to
T1857	6183-6186	DT	denotes	the
T1859	6187-6197	NN	denotes	expression
T1860	6198-6202	NN	denotes	site
T1858	6203-6207	NN	denotes	body
T1861	6208-6212	VBN	denotes	used
T1862	6213-6215	IN	denotes	by
T1863	6216-6223	JJ	denotes	African
T1864	6224-6236	NNPS	denotes	trypanosomes
T1865	6237-6239	TO	denotes	to
T1866	6240-6246	VB	denotes	ensure
T1867	6247-6255	JJ	denotes	singular
T1868	6256-6266	NN	denotes	expression
T1869	6267-6269	IN	denotes	of
T1870	6270-6274	RB	denotes	only
T1872	6275-6278	CD	denotes	one
T1871	6279-6282	NN	denotes	set
T1873	6283-6285	IN	denotes	of
T1874	6286-6293	JJ	denotes	variant
T1876	6294-6301	NN	denotes	surface
T1875	6302-6314	NN	denotes	glycoprotein
T1877	6315-6320	NNS	denotes	genes
T1878	6321-6322	-LRB-	denotes	[
T1879	6322-6324	CD	denotes	22
T1880	6324-6325	-RRB-	denotes	]
T1881	6325-6326	.	denotes	.
T1882	6326-6572	sentence	denotes	Finally, in a fourth model, transcriptional activity at one stochastically chosen olfactory receptor allele might send negative feedback to repress activity of all other olfactory receptors and/or positive feedback to enhance its own expression.
T1883	6327-6334	RB	denotes	Finally
T1885	6334-6336	,	denotes	,
T1886	6336-6338	IN	denotes	in
T1887	6339-6340	DT	denotes	a
T1889	6341-6347	JJ	denotes	fourth
T1888	6348-6353	NN	denotes	model
T1890	6353-6355	,	denotes	,
T1891	6355-6370	JJ	denotes	transcriptional
T1892	6371-6379	NN	denotes	activity
T1893	6380-6382	IN	denotes	at
T1894	6383-6386	CD	denotes	one
T1896	6387-6401	RB	denotes	stochastically
T1897	6402-6408	VBN	denotes	chosen
T1898	6409-6418	JJ	denotes	olfactory
T1899	6419-6427	NN	denotes	receptor
T1895	6428-6434	NN	denotes	allele
T1900	6435-6440	MD	denotes	might
T1884	6441-6445	VB	denotes	send
T1901	6446-6454	JJ	denotes	negative
T1902	6455-6463	NN	denotes	feedback
T1903	6464-6466	TO	denotes	to
T1904	6467-6474	VB	denotes	repress
T1905	6475-6483	NN	denotes	activity
T1906	6484-6486	IN	denotes	of
T1907	6487-6490	DT	denotes	all
T1909	6491-6496	JJ	denotes	other
T1910	6497-6506	JJ	denotes	olfactory
T1908	6507-6516	NNS	denotes	receptors
T1911	6517-6520	CC	denotes	and
T1912	6520-6521	HYPH	denotes	/
T1913	6521-6523	CC	denotes	or
T1914	6524-6532	JJ	denotes	positive
T1915	6533-6541	NN	denotes	feedback
T1916	6542-6544	TO	denotes	to
T1917	6545-6552	VB	denotes	enhance
T1918	6553-6556	PRP$	denotes	its
T1920	6557-6560	JJ	denotes	own
T1919	6561-6571	NN	denotes	expression
T1921	6571-6572	.	denotes	.
T1922	6572-6778	sentence	denotes	In the latter three models, some or all olfactory receptor genes might share transcription factor binding motifs, and in the first model, olfactory receptor genes might share a common recombination signal.
T1923	6573-6575	IN	denotes	In
T1925	6576-6579	DT	denotes	the
T1927	6580-6586	JJ	denotes	latter
T1928	6587-6592	CD	denotes	three
T1926	6593-6599	NNS	denotes	models
T1929	6599-6601	,	denotes	,
T1930	6601-6605	DT	denotes	some
T1931	6606-6608	CC	denotes	or
T1932	6609-6612	DT	denotes	all
T1934	6613-6622	JJ	denotes	olfactory
T1935	6623-6631	NN	denotes	receptor
T1933	6632-6637	NNS	denotes	genes
T1936	6638-6643	MD	denotes	might
T1924	6644-6649	VB	denotes	share
T1937	6650-6663	NN	denotes	transcription
T1938	6664-6670	NN	denotes	factor
T1939	6671-6678	NN	denotes	binding
T1940	6679-6685	NNS	denotes	motifs
T1941	6685-6687	,	denotes	,
T1942	6687-6690	CC	denotes	and
T1943	6691-6693	IN	denotes	in
T1945	6694-6697	DT	denotes	the
T1947	6698-6703	JJ	denotes	first
T1946	6704-6709	NN	denotes	model
T1948	6709-6711	,	denotes	,
T1949	6711-6720	JJ	denotes	olfactory
T1950	6721-6729	NN	denotes	receptor
T1951	6730-6735	NNS	denotes	genes
T1952	6736-6741	MD	denotes	might
T1944	6742-6747	VB	denotes	share
T1953	6748-6749	DT	denotes	a
T1955	6750-6756	JJ	denotes	common
T1956	6757-6770	NN	denotes	recombination
T1954	6771-6777	NN	denotes	signal
T1957	6777-6778	.	denotes	.
T1958	6778-6975	sentence	denotes	In order to perform computational and experimental searches for such signals, it is important to have a better idea of the transcriptional start site of a large number of olfactory receptor genes.
T1959	6779-6781	IN	denotes	In
T1961	6782-6787	NN	denotes	order
T1962	6788-6790	TO	denotes	to
T1963	6791-6798	VB	denotes	perform
T1964	6799-6812	JJ	denotes	computational
T1966	6813-6816	CC	denotes	and
T1967	6817-6829	JJ	denotes	experimental
T1965	6830-6838	NNS	denotes	searches
T1968	6839-6842	IN	denotes	for
T1969	6843-6847	JJ	denotes	such
T1970	6848-6855	NNS	denotes	signals
T1971	6855-6857	,	denotes	,
T1972	6857-6859	PRP	denotes	it
T1960	6860-6862	VBZ	denotes	is
T1973	6863-6872	JJ	denotes	important
T1974	6873-6875	TO	denotes	to
T1975	6876-6880	VB	denotes	have
T1976	6881-6882	DT	denotes	a
T1978	6883-6889	JJR	denotes	better
T1977	6890-6894	NN	denotes	idea
T1979	6895-6897	IN	denotes	of
T1980	6898-6901	DT	denotes	the
T1982	6902-6917	JJ	denotes	transcriptional
T1983	6918-6923	NN	denotes	start
T1981	6924-6928	NN	denotes	site
T1984	6929-6931	IN	denotes	of
T1985	6932-6933	DT	denotes	a
T1987	6934-6939	JJ	denotes	large
T1986	6940-6946	NN	denotes	number
T1988	6947-6949	IN	denotes	of
T1989	6950-6959	JJ	denotes	olfactory
T1990	6960-6968	NN	denotes	receptor
T1991	6969-6974	NNS	denotes	genes
T1992	6974-6975	.	denotes	.
T1993	6975-7139	sentence	denotes	Our olfactory receptor EST collection provides 5' untranslated region (UTR) sequences for many genes and, therefore, a large dataset of candidate promoter regions.
T1994	6976-6979	PRP$	denotes	Our
T1996	6980-6989	JJ	denotes	olfactory
T1997	6990-6998	NN	denotes	receptor
T1998	6999-7002	NN	denotes	EST
T1995	7003-7013	NN	denotes	collection
T1999	7014-7022	VBZ	denotes	provides
T2000	7023-7024	CD	denotes	5
T2002	7024-7025	SYM	denotes	'
T2003	7026-7038	JJ	denotes	untranslated
T2004	7039-7045	NN	denotes	region
T2005	7046-7047	-LRB-	denotes	(
T2006	7047-7050	NN	denotes	UTR
T2007	7050-7051	-RRB-	denotes	)
T2001	7052-7061	NNS	denotes	sequences
T2008	7062-7065	IN	denotes	for
T2009	7066-7070	JJ	denotes	many
T2010	7071-7076	NNS	denotes	genes
T2011	7077-7080	CC	denotes	and
T2012	7080-7082	,	denotes	,
T2013	7082-7091	RB	denotes	therefore
T2015	7091-7093	,	denotes	,
T2016	7093-7094	DT	denotes	a
T2017	7095-7100	JJ	denotes	large
T2014	7101-7108	NN	denotes	dataset
T2018	7109-7111	IN	denotes	of
T2019	7112-7121	NN	denotes	candidate
T2020	7122-7130	NN	denotes	promoter
T2021	7131-7138	NNS	denotes	regions
T2022	7138-7139	.	denotes	.
T2023	7139-7281	sentence	denotes	Olfactory receptor genes have an intronless coding region, simplifying both computational and experimental olfactory receptor identification.
T2024	7140-7149	JJ	denotes	Olfactory
T2025	7150-7158	NN	denotes	receptor
T2026	7159-7164	NNS	denotes	genes
T2027	7165-7169	VBP	denotes	have
T2028	7170-7172	DT	denotes	an
T2030	7173-7183	JJ	denotes	intronless
T2031	7184-7190	VBG	denotes	coding
T2029	7191-7197	NN	denotes	region
T2032	7197-7199	,	denotes	,
T2033	7199-7210	VBG	denotes	simplifying
T2034	7211-7215	CC	denotes	both
T2035	7216-7229	JJ	denotes	computational
T2037	7230-7233	CC	denotes	and
T2038	7234-7246	JJ	denotes	experimental
T2039	7247-7256	JJ	denotes	olfactory
T2040	7257-7265	NN	denotes	receptor
T2036	7266-7280	NN	denotes	identification
T2041	7280-7281	.	denotes	.
T2042	7281-7360	sentence	denotes	For a small number of olfactory receptors, gene structure has been determined.
T2043	7282-7285	IN	denotes	For
T2045	7286-7287	DT	denotes	a
T2047	7288-7293	JJ	denotes	small
T2046	7294-7300	NN	denotes	number
T2048	7301-7303	IN	denotes	of
T2049	7304-7313	JJ	denotes	olfactory
T2050	7314-7323	NNS	denotes	receptors
T2051	7323-7325	,	denotes	,
T2052	7325-7329	NN	denotes	gene
T2053	7330-7339	NN	denotes	structure
T2054	7340-7343	VBZ	denotes	has
T2055	7344-7348	VBN	denotes	been
T2044	7349-7359	VBN	denotes	determined
T2056	7359-7360	.	denotes	.
T2057	7360-7472	sentence	denotes	Additional 5' untranslated exons lie upstream of the coding region and can be alternatively spliced [19,23-26].
T2058	7361-7371	JJ	denotes	Additional
T2060	7372-7373	CD	denotes	5
T2061	7373-7374	SYM	denotes	'
T2062	7375-7387	JJ	denotes	untranslated
T2059	7388-7393	NNS	denotes	exons
T2063	7394-7397	VBP	denotes	lie
T2064	7398-7406	RB	denotes	upstream
T2065	7407-7409	IN	denotes	of
T2066	7410-7413	DT	denotes	the
T2068	7414-7420	VBG	denotes	coding
T2067	7421-7427	NN	denotes	region
T2069	7428-7431	CC	denotes	and
T2070	7432-7435	MD	denotes	can
T2072	7436-7438	VB	denotes	be
T2073	7439-7452	RB	denotes	alternatively
T2071	7453-7460	VBN	denotes	spliced
T2074	7461-7462	-LRB-	denotes	[
T2076	7462-7464	CD	denotes	19
T2077	7464-7465	,	denotes	,
T2075	7465-7467	CD	denotes	23
T2078	7467-7468	SYM	denotes	-
T2079	7468-7470	CD	denotes	26
T2080	7470-7471	-RRB-	denotes	]
T2081	7471-7472	.	denotes	.
T2082	7472-7524	sentence	denotes	The 3' untranslated region is typically intronless.
T2083	7473-7476	DT	denotes	The
T2085	7477-7478	CD	denotes	3
T2086	7478-7479	SYM	denotes	'
T2087	7480-7492	JJ	denotes	untranslated
T2084	7493-7499	NN	denotes	region
T2088	7500-7502	VBZ	denotes	is
T2089	7503-7512	RB	denotes	typically
T2090	7513-7523	JJ	denotes	intronless
T2091	7523-7524	.	denotes	.
T2092	7524-7657	sentence	denotes	Exceptions to this stereotyped structure have been described for some human olfactory receptors, but are thought to be rare [25-27].
T2093	7525-7535	NNS	denotes	Exceptions
T2095	7536-7538	IN	denotes	to
T2096	7539-7543	DT	denotes	this
T2098	7544-7555	JJ	denotes	stereotyped
T2097	7556-7565	NN	denotes	structure
T2099	7566-7570	VBP	denotes	have
T2100	7571-7575	VBN	denotes	been
T2094	7576-7585	VBN	denotes	described
T2101	7586-7589	IN	denotes	for
T2102	7590-7594	DT	denotes	some
T2104	7595-7600	JJ	denotes	human
T2105	7601-7610	JJ	denotes	olfactory
T2103	7611-7620	NNS	denotes	receptors
T2106	7620-7622	,	denotes	,
T2107	7622-7625	CC	denotes	but
T2108	7626-7629	VBP	denotes	are
T2109	7630-7637	VBN	denotes	thought
T2110	7638-7640	TO	denotes	to
T2111	7641-7643	VB	denotes	be
T2112	7644-7648	JJ	denotes	rare
T2113	7649-7650	-LRB-	denotes	[
T2114	7650-7652	CD	denotes	25
T2115	7652-7653	SYM	denotes	-
T2116	7653-7655	CD	denotes	27
T2117	7655-7656	-RRB-	denotes	]
T2118	7656-7657	.	denotes	.
T2119	7657-7781	sentence	denotes	cDNA identification and RACE data have been used to determine gene structure for about 30 genes, see, for example, [19,23].
T2120	7658-7662	NN	denotes	cDNA
T2121	7663-7677	NN	denotes	identification
T2123	7678-7681	CC	denotes	and
T2124	7682-7686	NN	denotes	RACE
T2125	7687-7691	NNS	denotes	data
T2126	7692-7696	VBP	denotes	have
T2127	7697-7701	VBN	denotes	been
T2122	7702-7706	VBN	denotes	used
T2128	7707-7709	TO	denotes	to
T2129	7710-7719	VB	denotes	determine
T2130	7720-7724	NN	denotes	gene
T2131	7725-7734	NN	denotes	structure
T2132	7735-7738	IN	denotes	for
T2133	7739-7744	RB	denotes	about
T2134	7745-7747	CD	denotes	30
T2135	7748-7753	NNS	denotes	genes
T2136	7753-7755	,	denotes	,
T2137	7755-7758	VB	denotes	see
T2138	7758-7760	,	denotes	,
T2139	7760-7763	IN	denotes	for
T2141	7764-7771	NN	denotes	example
T2142	7771-7773	,	denotes	,
T2143	7773-7774	-LRB-	denotes	[
T2144	7774-7776	CD	denotes	19
T2145	7776-7777	,	denotes	,
T2140	7777-7779	CD	denotes	23
T2146	7779-7780	-RRB-	denotes	]
T2147	7780-7781	.	denotes	.
T2148	7781-7933	sentence	denotes	However, computational prediction of the location of 5' upstream exons and the extent of the 3' UTR from genomic sequence has been extremely difficult.
T2149	7782-7789	RB	denotes	However
T2151	7789-7791	,	denotes	,
T2152	7791-7804	JJ	denotes	computational
T2153	7805-7815	NN	denotes	prediction
T2154	7816-7818	IN	denotes	of
T2155	7819-7822	DT	denotes	the
T2156	7823-7831	NN	denotes	location
T2157	7832-7834	IN	denotes	of
T2158	7835-7836	CD	denotes	5
T2160	7836-7837	SYM	denotes	'
T2161	7838-7846	JJ	denotes	upstream
T2159	7847-7852	NNS	denotes	exons
T2162	7853-7856	CC	denotes	and
T2163	7857-7860	DT	denotes	the
T2164	7861-7867	NN	denotes	extent
T2165	7868-7870	IN	denotes	of
T2166	7871-7874	DT	denotes	the
T2168	7875-7876	CD	denotes	3
T2169	7876-7877	SYM	denotes	'
T2167	7878-7881	NN	denotes	UTR
T2170	7882-7886	IN	denotes	from
T2171	7887-7894	JJ	denotes	genomic
T2172	7895-7903	NN	denotes	sequence
T2173	7904-7907	VBZ	denotes	has
T2150	7908-7912	VBN	denotes	been
T2174	7913-7922	RB	denotes	extremely
T2175	7923-7932	JJ	denotes	difficult
T2176	7932-7933	.	denotes	.
T2177	7933-8107	sentence	denotes	A combination of splice site predictions and similarity to other olfactory receptors has allowed some investigators to predict 5' exon locations for around 15 genes [25,28].
T2178	7934-7935	DT	denotes	A
T2179	7936-7947	NN	denotes	combination
T2181	7948-7950	IN	denotes	of
T2182	7951-7957	NN	denotes	splice
T2183	7958-7962	NN	denotes	site
T2184	7963-7974	NNS	denotes	predictions
T2185	7975-7978	CC	denotes	and
T2186	7979-7989	NN	denotes	similarity
T2187	7990-7992	IN	denotes	to
T2188	7993-7998	JJ	denotes	other
T2190	7999-8008	JJ	denotes	olfactory
T2189	8009-8018	NNS	denotes	receptors
T2191	8019-8022	VBZ	denotes	has
T2180	8023-8030	VBN	denotes	allowed
T2192	8031-8035	DT	denotes	some
T2193	8036-8049	NNS	denotes	investigators
T2195	8050-8052	TO	denotes	to
T2194	8053-8060	VB	denotes	predict
T2196	8061-8062	CD	denotes	5
T2198	8062-8063	SYM	denotes	'
T2197	8064-8068	NN	denotes	exon
T2199	8069-8078	NNS	denotes	locations
T2200	8079-8082	IN	denotes	for
T2201	8083-8089	RB	denotes	around
T2202	8090-8092	CD	denotes	15
T2203	8093-8098	NNS	denotes	genes
T2204	8099-8100	-LRB-	denotes	[
T2206	8100-8102	CD	denotes	25
T2207	8102-8103	,	denotes	,
T2205	8103-8105	CD	denotes	28
T2208	8105-8106	-RRB-	denotes	]
T2209	8106-8107	.	denotes	.
T2210	8107-8195	sentence	denotes	Experimental validation shows that some, but not all, predictions are accurate [24,25].
T2211	8108-8120	JJ	denotes	Experimental
T2212	8121-8131	NN	denotes	validation
T2213	8132-8137	VBZ	denotes	shows
T2214	8138-8142	IN	denotes	that
T2216	8143-8147	DT	denotes	some
T2217	8147-8149	,	denotes	,
T2218	8149-8152	CC	denotes	but
T2219	8153-8156	RB	denotes	not
T2220	8157-8160	DT	denotes	all
T2221	8160-8162	,	denotes	,
T2222	8162-8173	NNS	denotes	predictions
T2215	8174-8177	VBP	denotes	are
T2223	8178-8186	JJ	denotes	accurate
T2224	8187-8188	-LRB-	denotes	[
T2226	8188-8190	CD	denotes	24
T2227	8190-8191	,	denotes	,
T2225	8191-8193	CD	denotes	25
T2228	8193-8194	-RRB-	denotes	]
T2229	8194-8195	.	denotes	.
T2230	8195-8303	sentence	denotes	The total number of olfactory receptors for which gene structure is known is vastly increased by our study.
T2231	8196-8199	DT	denotes	The
T2233	8200-8205	JJ	denotes	total
T2232	8206-8212	NN	denotes	number
T2235	8213-8215	IN	denotes	of
T2236	8216-8225	JJ	denotes	olfactory
T2237	8226-8235	NNS	denotes	receptors
T2238	8236-8239	IN	denotes	for
T2240	8240-8245	WDT	denotes	which
T2241	8246-8250	NN	denotes	gene
T2242	8251-8260	NN	denotes	structure
T2243	8261-8263	VBZ	denotes	is
T2239	8264-8269	VBN	denotes	known
T2244	8270-8272	VBZ	denotes	is
T2245	8273-8279	RB	denotes	vastly
T2234	8280-8289	VBN	denotes	increased
T2246	8290-8292	IN	denotes	by
T2247	8293-8296	PRP$	denotes	our
T2248	8297-8302	NN	denotes	study
T2249	8302-8303	.	denotes	.
T2250	8303-8419	sentence	denotes	In this report, we describe the isolation and analysis of over 1,200 cDNAs representing 419 odorant receptor genes.
T2251	8304-8306	IN	denotes	In
T2253	8307-8311	DT	denotes	this
T2254	8312-8318	NN	denotes	report
T2255	8318-8320	,	denotes	,
T2256	8320-8322	PRP	denotes	we
T2252	8323-8331	VBP	denotes	describe
T2257	8332-8335	DT	denotes	the
T2258	8336-8345	NN	denotes	isolation
T2259	8346-8349	CC	denotes	and
T2260	8350-8358	NN	denotes	analysis
T2261	8359-8361	IN	denotes	of
T2262	8362-8366	IN	denotes	over
T2263	8367-8372	CD	denotes	1,200
T2264	8373-8378	NNS	denotes	cDNAs
T2265	8379-8391	VBG	denotes	representing
T2266	8392-8395	CD	denotes	419
T2268	8396-8403	JJ	denotes	odorant
T2269	8404-8412	NN	denotes	receptor
T2267	8413-8418	NNS	denotes	genes
T2270	8418-8419	.	denotes	.
T2271	8419-8568	sentence	denotes	We screened a mouse olfactory epithelium library with degenerate olfactory receptor probes and obtained 5' end sequences (ESTs) from purified cDNAs.
T2272	8420-8422	PRP	denotes	We
T2273	8423-8431	VBD	denotes	screened
T2274	8432-8433	DT	denotes	a
T2276	8434-8439	NN	denotes	mouse
T2278	8440-8449	JJ	denotes	olfactory
T2277	8450-8460	NN	denotes	epithelium
T2275	8461-8468	NN	denotes	library
T2279	8469-8473	IN	denotes	with
T2280	8474-8484	JJ	denotes	degenerate
T2282	8485-8494	JJ	denotes	olfactory
T2283	8495-8503	NN	denotes	receptor
T2281	8504-8510	NNS	denotes	probes
T2284	8511-8514	CC	denotes	and
T2285	8515-8523	VBN	denotes	obtained
T2286	8524-8525	CD	denotes	5
T2287	8525-8526	SYM	denotes	'
T2288	8527-8530	NN	denotes	end
T2289	8531-8540	NNS	denotes	sequences
T2290	8541-8542	-LRB-	denotes	(
T2291	8542-8546	NNS	denotes	ESTs
T2292	8546-8547	-RRB-	denotes	)
T2293	8548-8552	IN	denotes	from
T2294	8553-8561	VBN	denotes	purified
T2295	8562-8567	NNS	denotes	cDNAs
T2296	8567-8568	.	denotes	.
T2297	8568-8834	sentence	denotes	These clones confirm olfactory expression of over 400 olfactory receptors, provide their gene structure, demonstrate that not all olfactory receptors are expressed at the same level and show that most olfactory receptor genes have multiple transcriptional isoforms.
T2298	8569-8574	DT	denotes	These
T2299	8575-8581	NNS	denotes	clones
T2300	8582-8589	VBP	denotes	confirm
T2301	8590-8599	JJ	denotes	olfactory
T2302	8600-8610	NN	denotes	expression
T2303	8611-8613	IN	denotes	of
T2304	8614-8618	IN	denotes	over
T2305	8619-8622	CD	denotes	400
T2307	8623-8632	JJ	denotes	olfactory
T2306	8633-8642	NNS	denotes	receptors
T2308	8642-8644	,	denotes	,
T2309	8644-8651	VBP	denotes	provide
T2310	8652-8657	PRP$	denotes	their
T2312	8658-8662	NN	denotes	gene
T2311	8663-8672	NN	denotes	structure
T2313	8672-8674	,	denotes	,
T2314	8674-8685	VBP	denotes	demonstrate
T2315	8686-8690	IN	denotes	that
T2317	8691-8694	RB	denotes	not
T2319	8695-8698	DT	denotes	all
T2320	8699-8708	JJ	denotes	olfactory
T2318	8709-8718	NNS	denotes	receptors
T2321	8719-8722	VBP	denotes	are
T2316	8723-8732	VBN	denotes	expressed
T2322	8733-8735	IN	denotes	at
T2323	8736-8739	DT	denotes	the
T2325	8740-8744	JJ	denotes	same
T2324	8745-8750	NN	denotes	level
T2326	8751-8754	CC	denotes	and
T2327	8755-8759	VBP	denotes	show
T2328	8760-8764	IN	denotes	that
T2330	8765-8769	JJS	denotes	most
T2332	8770-8779	JJ	denotes	olfactory
T2333	8780-8788	NN	denotes	receptor
T2331	8789-8794	NNS	denotes	genes
T2329	8795-8799	VBP	denotes	have
T2334	8800-8808	JJ	denotes	multiple
T2336	8809-8824	JJ	denotes	transcriptional
T2335	8825-8833	NNS	denotes	isoforms
T2337	8833-8834	.	denotes	.
T2624	8845-8847	RB	denotes	At
T2626	8848-8853	RBS	denotes	least
T2625	8854-8857	CD	denotes	419
T2628	8858-8863	NN	denotes	mouse
T2629	8864-8873	JJ	denotes	olfactory
T2630	8874-8882	NN	denotes	receptor
T2627	8883-8888	NNS	denotes	genes
T2632	8889-8892	VBP	denotes	are
T2631	8893-8902	VBN	denotes	expressed
T2633	8903-8905	IN	denotes	in
T2634	8906-8909	DT	denotes	the
T2636	8910-8919	JJ	denotes	olfactory
T2635	8920-8930	NN	denotes	epithelium
T2637	8930-9087	sentence	denotes	We have isolated 1,264 olfactory receptor cDNA clones, which together confirm the olfactory epithelial expression of 419 annotated olfactory receptor genes.
T2638	8931-8933	PRP	denotes	We
T2640	8934-8938	VBP	denotes	have
T2639	8939-8947	VBN	denotes	isolated
T2641	8948-8953	CD	denotes	1,264
T2643	8954-8963	JJ	denotes	olfactory
T2644	8964-8972	NN	denotes	receptor
T2645	8973-8977	NN	denotes	cDNA
T2642	8978-8984	NNS	denotes	clones
T2646	8984-8986	,	denotes	,
T2647	8986-8991	WDT	denotes	which
T2649	8992-9000	RB	denotes	together
T2648	9001-9008	VBP	denotes	confirm
T2650	9009-9012	DT	denotes	the
T2652	9013-9022	JJ	denotes	olfactory
T2653	9023-9033	JJ	denotes	epithelial
T2651	9034-9044	NN	denotes	expression
T2654	9045-9047	IN	denotes	of
T2655	9048-9051	CD	denotes	419
T2657	9052-9061	VBN	denotes	annotated
T2658	9062-9071	JJ	denotes	olfactory
T2659	9072-9080	NN	denotes	receptor
T2656	9081-9086	NNS	denotes	genes
T2660	9086-9087	.	denotes	.
T2661	9087-9345	sentence	denotes	We used low-stringency hybridization with degenerate olfactory receptor DNA probes to screen around 4,100,000 plaque-forming units (pfu) of an adult mouse olfactory epithelium cDNA library and around 640,000 pfu of an embryonic olfactory epithelium library.
T2662	9088-9090	PRP	denotes	We
T2663	9091-9095	VBD	denotes	used
T2664	9096-9099	JJ	denotes	low
T2666	9099-9100	HYPH	denotes	-
T2665	9100-9110	NN	denotes	stringency
T2667	9111-9124	NN	denotes	hybridization
T2668	9125-9129	IN	denotes	with
T2669	9130-9140	JJ	denotes	degenerate
T2671	9141-9150	JJ	denotes	olfactory
T2672	9151-9159	NN	denotes	receptor
T2673	9160-9163	NN	denotes	DNA
T2670	9164-9170	NNS	denotes	probes
T2674	9171-9173	TO	denotes	to
T2675	9174-9180	VB	denotes	screen
T2676	9181-9187	RB	denotes	around
T2677	9188-9197	CD	denotes	4,100,000
T2679	9198-9204	NN	denotes	plaque
T2681	9204-9205	HYPH	denotes	-
T2680	9205-9212	VBG	denotes	forming
T2678	9213-9218	NNS	denotes	units
T2682	9219-9220	-LRB-	denotes	(
T2683	9220-9223	NNS	denotes	pfu
T2684	9223-9224	-RRB-	denotes	)
T2685	9225-9227	IN	denotes	of
T2686	9228-9230	DT	denotes	an
T2688	9231-9236	JJ	denotes	adult
T2689	9237-9242	NN	denotes	mouse
T2691	9243-9252	JJ	denotes	olfactory
T2692	9253-9263	NN	denotes	epithelium
T2690	9264-9268	NN	denotes	cDNA
T2687	9269-9276	NN	denotes	library
T2693	9277-9280	CC	denotes	and
T2694	9281-9287	RB	denotes	around
T2695	9288-9295	CD	denotes	640,000
T2696	9296-9299	NNS	denotes	pfu
T2697	9300-9302	IN	denotes	of
T2698	9303-9305	DT	denotes	an
T2700	9306-9315	JJ	denotes	embryonic
T2702	9316-9325	JJ	denotes	olfactory
T2701	9326-9336	NN	denotes	epithelium
T2699	9337-9344	NN	denotes	library
T2703	9344-9345	.	denotes	.
T2704	9345-9461	sentence	denotes	We obtained sequences from 1,715 hybridization-positive cDNAs following secondary screens to isolate single clones.
T2705	9346-9348	PRP	denotes	We
T2706	9349-9357	VBD	denotes	obtained
T2707	9358-9367	NNS	denotes	sequences
T2708	9368-9372	IN	denotes	from
T2709	9373-9378	CD	denotes	1,715
T2711	9379-9392	NN	denotes	hybridization
T2713	9392-9393	HYPH	denotes	-
T2712	9393-9401	JJ	denotes	positive
T2710	9402-9407	NNS	denotes	cDNAs
T2714	9408-9417	VBG	denotes	following
T2715	9418-9427	JJ	denotes	secondary
T2716	9428-9435	NNS	denotes	screens
T2717	9436-9438	TO	denotes	to
T2718	9439-9446	VB	denotes	isolate
T2719	9447-9453	JJ	denotes	single
T2720	9454-9460	NNS	denotes	clones
T2721	9460-9461	.	denotes	.
T2722	9461-9533	sentence	denotes	Of these clones, 1,264 yielded olfactory receptor-containing sequences.
T2723	9462-9464	IN	denotes	Of
T2725	9465-9470	DT	denotes	these
T2726	9471-9477	NNS	denotes	clones
T2727	9477-9479	,	denotes	,
T2728	9479-9484	CD	denotes	1,264
T2724	9485-9492	VBD	denotes	yielded
T2729	9493-9502	JJ	denotes	olfactory
T2730	9503-9511	NN	denotes	receptor
T2732	9511-9512	HYPH	denotes	-
T2731	9512-9522	VBG	denotes	containing
T2733	9523-9532	NNS	denotes	sequences
T2734	9532-9533	.	denotes	.
T2735	9533-9647	sentence	denotes	The 26% false-positive rate is a consequence of using low-stringency hybridization to obtain maximal sensitivity.
T2736	9534-9537	DT	denotes	The
T2738	9538-9540	CD	denotes	26
T2739	9540-9541	NN	denotes	%
T2740	9542-9547	JJ	denotes	false
T2742	9547-9548	HYPH	denotes	-
T2741	9548-9556	JJ	denotes	positive
T2737	9557-9561	NN	denotes	rate
T2743	9562-9564	VBZ	denotes	is
T2744	9565-9566	DT	denotes	a
T2745	9567-9578	NN	denotes	consequence
T2746	9579-9581	IN	denotes	of
T2747	9582-9587	VBG	denotes	using
T2748	9588-9591	JJ	denotes	low
T2750	9591-9592	HYPH	denotes	-
T2749	9592-9602	NN	denotes	stringency
T2751	9603-9616	NN	denotes	hybridization
T2752	9617-9619	TO	denotes	to
T2753	9620-9626	VB	denotes	obtain
T2754	9627-9634	JJ	denotes	maximal
T2755	9635-9646	NN	denotes	sensitivity
T2756	9646-9647	.	denotes	.
T2757	9647-9955	sentence	denotes	Continuing the screen would have resulted in cDNAs from additional olfactory receptors, but we reached a point of limiting returns: our final batch of 45 olfactory receptor-positive sequences represented 33 different olfactory receptors, of which only four had not been encountered previously in our screen.
T2758	9648-9658	VBG	denotes	Continuing
T2760	9659-9662	DT	denotes	the
T2761	9663-9669	NN	denotes	screen
T2762	9670-9675	MD	denotes	would
T2763	9676-9680	VB	denotes	have
T2759	9681-9689	VBN	denotes	resulted
T2765	9690-9692	IN	denotes	in
T2766	9693-9698	NNS	denotes	cDNAs
T2767	9699-9703	IN	denotes	from
T2768	9704-9714	JJ	denotes	additional
T2770	9715-9724	JJ	denotes	olfactory
T2769	9725-9734	NNS	denotes	receptors
T2771	9734-9736	,	denotes	,
T2772	9736-9739	CC	denotes	but
T2773	9740-9742	PRP	denotes	we
T2774	9743-9750	VBD	denotes	reached
T2775	9751-9752	DT	denotes	a
T2776	9753-9758	NN	denotes	point
T2777	9759-9761	IN	denotes	of
T2778	9762-9770	VBG	denotes	limiting
T2779	9771-9778	NNS	denotes	returns
T2780	9778-9780	:	denotes	:
T2781	9780-9783	PRP$	denotes	our
T2783	9784-9789	JJ	denotes	final
T2782	9790-9795	NN	denotes	batch
T2784	9796-9798	IN	denotes	of
T2785	9799-9801	CD	denotes	45
T2787	9802-9811	JJ	denotes	olfactory
T2788	9812-9820	NN	denotes	receptor
T2790	9820-9821	HYPH	denotes	-
T2789	9821-9829	JJ	denotes	positive
T2786	9830-9839	NNS	denotes	sequences
T2764	9840-9851	VBD	denotes	represented
T2791	9852-9854	CD	denotes	33
T2793	9855-9864	JJ	denotes	different
T2794	9865-9874	JJ	denotes	olfactory
T2792	9875-9884	NNS	denotes	receptors
T2795	9884-9886	,	denotes	,
T2796	9886-9888	IN	denotes	of
T2798	9889-9894	WDT	denotes	which
T2799	9895-9899	RB	denotes	only
T2800	9900-9904	CD	denotes	four
T2801	9905-9908	VBD	denotes	had
T2802	9909-9912	RB	denotes	not
T2803	9913-9917	VBN	denotes	been
T2797	9918-9929	VBN	denotes	encountered
T2804	9930-9940	RB	denotes	previously
T2805	9941-9943	IN	denotes	in
T2806	9944-9947	PRP$	denotes	our
T2807	9948-9954	NN	denotes	screen
T2808	9954-9955	.	denotes	.
T2809	9955-10019	sentence	denotes	Sequence analysis shows that the libraries are of high quality.
T2810	9956-9964	NN	denotes	Sequence
T2811	9965-9973	NN	denotes	analysis
T2812	9974-9979	VBZ	denotes	shows
T2813	9980-9984	IN	denotes	that
T2815	9985-9988	DT	denotes	the
T2816	9989-9998	NNS	denotes	libraries
T2814	9999-10002	VBP	denotes	are
T2817	10003-10005	IN	denotes	of
T2818	10006-10010	JJ	denotes	high
T2819	10011-10018	NN	denotes	quality
T2820	10018-10019	.	denotes	.
T2821	10019-10173	sentence	denotes	Firstly, directional cloning was successful: only eight out of 1,430 cDNA sequences with any protein homology matched that protein on the reverse strand.
T2822	10020-10027	RB	denotes	Firstly
T2824	10027-10029	,	denotes	,
T2825	10029-10040	JJ	denotes	directional
T2826	10041-10048	NN	denotes	cloning
T2827	10049-10052	VBD	denotes	was
T2828	10053-10063	JJ	denotes	successful
T2829	10063-10065	:	denotes	:
T2830	10065-10069	RB	denotes	only
T2832	10070-10075	CD	denotes	eight
T2833	10076-10079	IN	denotes	out
T2834	10080-10082	IN	denotes	of
T2831	10083-10088	CD	denotes	1,430
T2836	10089-10093	NN	denotes	cDNA
T2835	10094-10103	NNS	denotes	sequences
T2837	10104-10108	IN	denotes	with
T2838	10109-10112	DT	denotes	any
T2840	10113-10120	NN	denotes	protein
T2839	10121-10129	NN	denotes	homology
T2823	10130-10137	VBD	denotes	matched
T2841	10138-10142	DT	denotes	that
T2842	10143-10150	NN	denotes	protein
T2843	10151-10153	IN	denotes	on
T2844	10154-10157	DT	denotes	the
T2846	10158-10165	JJ	denotes	reverse
T2845	10166-10172	NN	denotes	strand
T2847	10172-10173	.	denotes	.
T2848	10173-10485	sentence	denotes	Secondly, genomic contamination is rare: when the 83 olfactory receptor-containing sequences that had a 5' UTR of 400 bp or longer were aligned to genomic sequence, 80 spliced across at least one intron, leaving a maximum of three clones (3.6%) that potentially represent genomic contamination of the libraries.
T2849	10174-10182	RB	denotes	Secondly
T2851	10182-10184	,	denotes	,
T2852	10184-10191	JJ	denotes	genomic
T2853	10192-10205	NN	denotes	contamination
T2854	10206-10208	VBZ	denotes	is
T2855	10209-10213	JJ	denotes	rare
T2856	10213-10215	:	denotes	:
T2857	10215-10219	WRB	denotes	when
T2859	10220-10223	DT	denotes	the
T2861	10224-10226	CD	denotes	83
T2862	10227-10236	JJ	denotes	olfactory
T2863	10237-10245	NN	denotes	receptor
T2865	10245-10246	HYPH	denotes	-
T2864	10246-10256	VBG	denotes	containing
T2860	10257-10266	NNS	denotes	sequences
T2866	10267-10271	WDT	denotes	that
T2867	10272-10275	VBD	denotes	had
T2868	10276-10277	DT	denotes	a
T2870	10278-10279	CD	denotes	5
T2871	10279-10280	SYM	denotes	'
T2869	10281-10284	NN	denotes	UTR
T2872	10285-10287	IN	denotes	of
T2873	10288-10291	CD	denotes	400
T2874	10292-10294	NNS	denotes	bp
T2875	10295-10297	CC	denotes	or
T2876	10298-10304	JJR	denotes	longer
T2877	10305-10309	VBD	denotes	were
T2858	10310-10317	VBN	denotes	aligned
T2878	10318-10320	IN	denotes	to
T2879	10321-10328	JJ	denotes	genomic
T2880	10329-10337	NN	denotes	sequence
T2881	10337-10339	,	denotes	,
T2882	10339-10341	CD	denotes	80
T2850	10342-10349	VBN	denotes	spliced
T2883	10350-10356	IN	denotes	across
T2884	10357-10359	RB	denotes	at
T2886	10360-10365	RBS	denotes	least
T2885	10366-10369	CD	denotes	one
T2887	10370-10376	NN	denotes	intron
T2888	10376-10378	,	denotes	,
T2889	10378-10385	VBG	denotes	leaving
T2890	10386-10387	DT	denotes	a
T2891	10388-10395	NN	denotes	maximum
T2892	10396-10398	IN	denotes	of
T2893	10399-10404	CD	denotes	three
T2894	10405-10411	NNS	denotes	clones
T2895	10412-10413	-LRB-	denotes	(
T2897	10413-10416	CD	denotes	3.6
T2896	10416-10417	NN	denotes	%
T2898	10417-10418	-RRB-	denotes	)
T2899	10419-10423	WDT	denotes	that
T2901	10424-10435	RB	denotes	potentially
T2900	10436-10445	VBP	denotes	represent
T2902	10446-10453	JJ	denotes	genomic
T2903	10454-10467	NN	denotes	contamination
T2904	10468-10470	IN	denotes	of
T2905	10471-10474	DT	denotes	the
T2906	10475-10484	NNS	denotes	libraries
T2907	10484-10485	.	denotes	.
T2908	10485-10706	sentence	denotes	Thirdly, most clones are of a reasonable length: although we did not determine whether clones are full-length, 881 of 1,264 (70%) olfactory receptor cDNAs contain the gene's start codon and at least some 5' UTR sequence.
T2909	10486-10493	RB	denotes	Thirdly
T2911	10493-10495	,	denotes	,
T2912	10495-10499	JJS	denotes	most
T2913	10500-10506	NNS	denotes	clones
T2914	10507-10510	VBP	denotes	are
T2915	10511-10513	IN	denotes	of
T2916	10514-10515	DT	denotes	a
T2918	10516-10526	JJ	denotes	reasonable
T2917	10527-10533	NN	denotes	length
T2919	10533-10535	:	denotes	:
T2920	10535-10543	IN	denotes	although
T2922	10544-10546	PRP	denotes	we
T2923	10547-10550	VBD	denotes	did
T2924	10551-10554	RB	denotes	not
T2921	10555-10564	VB	denotes	determine
T2925	10565-10572	IN	denotes	whether
T2927	10573-10579	NNS	denotes	clones
T2926	10580-10583	VBP	denotes	are
T2928	10584-10588	JJ	denotes	full
T2930	10588-10589	HYPH	denotes	-
T2929	10589-10595	NN	denotes	length
T2931	10595-10597	,	denotes	,
T2932	10597-10600	CD	denotes	881
T2934	10601-10603	IN	denotes	of
T2933	10604-10609	CD	denotes	1,264
T2936	10610-10611	-LRB-	denotes	(
T2938	10611-10613	CD	denotes	70
T2937	10613-10614	NN	denotes	%
T2939	10614-10615	-RRB-	denotes	)
T2940	10616-10625	JJ	denotes	olfactory
T2941	10626-10634	NN	denotes	receptor
T2935	10635-10640	NNS	denotes	cDNAs
T2910	10641-10648	VBP	denotes	contain
T2942	10649-10652	DT	denotes	the
T2943	10653-10657	NN	denotes	gene
T2945	10657-10659	POS	denotes	's
T2946	10660-10665	NN	denotes	start
T2944	10666-10671	NN	denotes	codon
T2947	10672-10675	CC	denotes	and
T2948	10676-10678	RB	denotes	at
T2949	10679-10684	RBS	denotes	least
T2951	10685-10689	DT	denotes	some
T2952	10690-10691	CD	denotes	5
T2954	10691-10692	SYM	denotes	'
T2953	10693-10696	NN	denotes	UTR
T2950	10697-10705	NN	denotes	sequence
T2955	10705-10706	.	denotes	.
T2956	10706-10893	sentence	denotes	In order to match cDNAs to their genomic counterparts, first we updated our catalog of mouse olfactory receptor genes [1] based on Celera's most recent genome assembly (Release 13) [29].
T2957	10707-10709	IN	denotes	In
T2959	10710-10715	NN	denotes	order
T2960	10716-10718	TO	denotes	to
T2961	10719-10724	VB	denotes	match
T2962	10725-10730	NNS	denotes	cDNAs
T2963	10731-10733	IN	denotes	to
T2964	10734-10739	PRP$	denotes	their
T2966	10740-10747	JJ	denotes	genomic
T2965	10748-10760	NNS	denotes	counterparts
T2967	10760-10762	,	denotes	,
T2968	10762-10767	RB	denotes	first
T2969	10768-10770	PRP	denotes	we
T2958	10771-10778	VBD	denotes	updated
T2970	10779-10782	PRP$	denotes	our
T2971	10783-10790	NN	denotes	catalog
T2972	10791-10793	IN	denotes	of
T2973	10794-10799	NN	denotes	mouse
T2975	10800-10809	JJ	denotes	olfactory
T2976	10810-10818	NN	denotes	receptor
T2974	10819-10824	NNS	denotes	genes
T2977	10825-10826	-LRB-	denotes	[
T2978	10826-10827	CD	denotes	1
T2979	10827-10828	-RRB-	denotes	]
T2980	10829-10834	VBN	denotes	based
T2981	10835-10837	IN	denotes	on
T2982	10838-10844	NNP	denotes	Celera
T2984	10844-10846	POS	denotes	's
T2985	10847-10851	RBS	denotes	most
T2986	10852-10858	JJ	denotes	recent
T2987	10859-10865	NN	denotes	genome
T2983	10866-10874	NN	denotes	assembly
T2988	10875-10876	-LRB-	denotes	(
T2989	10876-10883	NN	denotes	Release
T2990	10884-10886	CD	denotes	13
T2991	10886-10887	-RRB-	denotes	)
T2992	10888-10889	-LRB-	denotes	[
T2993	10889-10891	CD	denotes	29
T2994	10891-10892	-RRB-	denotes	]
T2995	10892-10893	.	denotes	.
T2996	10893-10998	sentence	denotes	Previous reports of the mouse olfactory receptor repertoire [1,2] were based on the Release 12 assembly.
T2997	10894-10902	JJ	denotes	Previous
T2998	10903-10910	NNS	denotes	reports
T3000	10911-10913	IN	denotes	of
T3001	10914-10917	DT	denotes	the
T3003	10918-10923	NN	denotes	mouse
T3005	10924-10933	JJ	denotes	olfactory
T3004	10934-10942	NN	denotes	receptor
T3002	10943-10953	NN	denotes	repertoire
T3006	10954-10955	-LRB-	denotes	[
T3008	10955-10956	CD	denotes	1
T3009	10956-10957	,	denotes	,
T3007	10957-10958	CD	denotes	2
T3010	10958-10959	-RRB-	denotes	]
T3011	10960-10964	VBD	denotes	were
T2999	10965-10970	VBN	denotes	based
T3012	10971-10973	IN	denotes	on
T3013	10974-10977	DT	denotes	the
T3015	10978-10985	NN	denotes	Release
T3016	10986-10988	CD	denotes	12
T3014	10989-10997	NN	denotes	assembly
T3017	10997-10998	.	denotes	.
T3018	10998-11102	sentence	denotes	Release 13 consists of fewer, longer scaffold sequences containing fewer, smaller gaps than Release 12.
T3019	10999-11006	NN	denotes	Release
T3021	11007-11009	CD	denotes	13
T3020	11010-11018	VBZ	denotes	consists
T3022	11019-11021	IN	denotes	of
T3023	11022-11027	JJR	denotes	fewer
T3025	11027-11029	,	denotes	,
T3024	11029-11035	JJR	denotes	longer
T3027	11036-11044	NN	denotes	scaffold
T3026	11045-11054	NNS	denotes	sequences
T3028	11055-11065	VBG	denotes	containing
T3029	11066-11071	JJR	denotes	fewer
T3031	11071-11073	,	denotes	,
T3030	11073-11080	JJR	denotes	smaller
T3032	11081-11085	NNS	denotes	gaps
T3033	11086-11090	IN	denotes	than
T3034	11091-11098	NN	denotes	Release
T3035	11099-11101	CD	denotes	12
T3036	11101-11102	.	denotes	.
T3037	11102-11437	sentence	denotes	Using the BLAST-based methods detailed previously [1], we identified 1,490 olfactory receptor sequences in the new assembly, including 1,107 intact olfactory receptor genes (compared to 866 intact olfactory receptors in the old assembly) reflecting the reduced sequence error rate and increased coverage of the new assembly (Table 1).
T3038	11103-11108	VBG	denotes	Using
T3040	11109-11112	DT	denotes	the
T3042	11113-11118	NN	denotes	BLAST
T3044	11118-11119	HYPH	denotes	-
T3043	11119-11124	VBN	denotes	based
T3041	11125-11132	NNS	denotes	methods
T3045	11133-11141	VBN	denotes	detailed
T3046	11142-11152	RB	denotes	previously
T3047	11153-11154	-LRB-	denotes	[
T3048	11154-11155	CD	denotes	1
T3049	11155-11156	-RRB-	denotes	]
T3050	11156-11158	,	denotes	,
T3051	11158-11160	PRP	denotes	we
T3039	11161-11171	VBD	denotes	identified
T3052	11172-11177	CD	denotes	1,490
T3054	11178-11187	JJ	denotes	olfactory
T3055	11188-11196	NN	denotes	receptor
T3053	11197-11206	NNS	denotes	sequences
T3056	11207-11209	IN	denotes	in
T3057	11210-11213	DT	denotes	the
T3059	11214-11217	JJ	denotes	new
T3058	11218-11226	NN	denotes	assembly
T3060	11226-11228	,	denotes	,
T3061	11228-11237	VBG	denotes	including
T3062	11238-11243	CD	denotes	1,107
T3064	11244-11250	JJ	denotes	intact
T3065	11251-11260	JJ	denotes	olfactory
T3066	11261-11269	NN	denotes	receptor
T3063	11270-11275	NNS	denotes	genes
T3067	11276-11277	-LRB-	denotes	(
T3068	11277-11285	VBN	denotes	compared
T3069	11286-11288	IN	denotes	to
T3070	11289-11292	CD	denotes	866
T3072	11293-11299	JJ	denotes	intact
T3073	11300-11309	JJ	denotes	olfactory
T3071	11310-11319	NNS	denotes	receptors
T3074	11320-11322	IN	denotes	in
T3075	11323-11326	DT	denotes	the
T3077	11327-11330	JJ	denotes	old
T3076	11331-11339	NN	denotes	assembly
T3078	11339-11340	-RRB-	denotes	)
T3079	11341-11351	VBG	denotes	reflecting
T3080	11352-11355	DT	denotes	the
T3082	11356-11363	VBN	denotes	reduced
T3083	11364-11372	NN	denotes	sequence
T3084	11373-11378	NN	denotes	error
T3081	11379-11383	NN	denotes	rate
T3085	11384-11387	CC	denotes	and
T3086	11388-11397	VBN	denotes	increased
T3087	11398-11406	NN	denotes	coverage
T3088	11407-11409	IN	denotes	of
T3089	11410-11413	DT	denotes	the
T3091	11414-11417	JJ	denotes	new
T3090	11418-11426	NN	denotes	assembly
T3092	11427-11428	-LRB-	denotes	(
T3093	11428-11433	NN	denotes	Table
T3094	11434-11435	CD	denotes	1
T3095	11435-11436	-RRB-	denotes	)
T3096	11436-11437	.	denotes	.
T3097	11437-11657	sentence	denotes	We created a local database of genomic sequences including all olfactory receptor loci and 0.5 Mb flanking sequences (if available) and compared each cDNA sequence to this 'olfactory subgenome' database using sim4 [30].
T3098	11438-11440	PRP	denotes	We
T3099	11441-11448	VBD	denotes	created
T3100	11449-11450	DT	denotes	a
T3102	11451-11456	JJ	denotes	local
T3101	11457-11465	NN	denotes	database
T3103	11466-11468	IN	denotes	of
T3104	11469-11476	JJ	denotes	genomic
T3105	11477-11486	NNS	denotes	sequences
T3106	11487-11496	VBG	denotes	including
T3107	11497-11500	DT	denotes	all
T3109	11501-11510	JJ	denotes	olfactory
T3110	11511-11519	NN	denotes	receptor
T3108	11520-11524	NNS	denotes	loci
T3111	11525-11528	CC	denotes	and
T3112	11529-11532	CD	denotes	0.5
T3113	11533-11535	NN	denotes	Mb
T3115	11536-11544	NN	denotes	flanking
T3114	11545-11554	NNS	denotes	sequences
T3116	11555-11556	-LRB-	denotes	(
T3118	11556-11558	IN	denotes	if
T3117	11559-11568	JJ	denotes	available
T3119	11568-11569	-RRB-	denotes	)
T3120	11570-11573	CC	denotes	and
T3121	11574-11582	VBN	denotes	compared
T3122	11583-11587	DT	denotes	each
T3124	11588-11592	NN	denotes	cDNA
T3123	11593-11601	NN	denotes	sequence
T3125	11602-11604	IN	denotes	to
T3126	11605-11609	DT	denotes	this
T3128	11610-11611	``	denotes	'
T3129	11611-11620	JJ	denotes	olfactory
T3130	11621-11630	NN	denotes	subgenome
T3131	11630-11631	''	denotes	'
T3127	11632-11640	NN	denotes	database
T3132	11641-11646	VBG	denotes	using
T3133	11647-11651	NN	denotes	sim4
T3134	11652-11653	-LRB-	denotes	[
T3135	11653-11655	CD	denotes	30
T3136	11655-11656	-RRB-	denotes	]
T3137	11656-11657	.	denotes	.
T3138	11657-11814	sentence	denotes	cDNAs were assigned to individual genes based on their best match to an olfactory receptor coding region or its upstream region (see Materials and methods).
T3139	11658-11663	NNS	denotes	cDNAs
T3141	11664-11668	VBD	denotes	were
T3140	11669-11677	VBN	denotes	assigned
T3142	11678-11680	IN	denotes	to
T3143	11681-11691	JJ	denotes	individual
T3144	11692-11697	NNS	denotes	genes
T3145	11698-11703	VBN	denotes	based
T3146	11704-11706	IN	denotes	on
T3147	11707-11712	PRP$	denotes	their
T3149	11713-11717	JJS	denotes	best
T3148	11718-11723	NN	denotes	match
T3150	11724-11726	IN	denotes	to
T3151	11727-11729	DT	denotes	an
T3153	11730-11739	JJ	denotes	olfactory
T3154	11740-11748	NN	denotes	receptor
T3155	11749-11755	NN	denotes	coding
T3152	11756-11762	NN	denotes	region
T3156	11763-11765	CC	denotes	or
T3157	11766-11769	PRP$	denotes	its
T3159	11770-11778	JJ	denotes	upstream
T3158	11779-11785	NN	denotes	region
T3160	11786-11787	-LRB-	denotes	(
T3161	11787-11790	VB	denotes	see
T3162	11791-11800	NNS	denotes	Materials
T3163	11801-11804	CC	denotes	and
T3164	11805-11812	NNS	denotes	methods
T3165	11812-11813	-RRB-	denotes	)
T3166	11813-11814	.	denotes	.
T3167	11814-12105	sentence	denotes	Of the 1,264 olfactory receptor cDNAs, 1,176 matched a total of 419 olfactory receptor genes; the remaining cDNAs either matched an olfactory receptor below our 96% nucleotide identity threshold or had ambiguous matches encompassing more than one olfactory receptor gene region (see below).
T3168	11815-11817	IN	denotes	Of
T3170	11818-11821	DT	denotes	the
T3172	11822-11827	CD	denotes	1,264
T3173	11828-11837	JJ	denotes	olfactory
T3174	11838-11846	NN	denotes	receptor
T3171	11847-11852	NNS	denotes	cDNAs
T3175	11852-11854	,	denotes	,
T3176	11854-11859	CD	denotes	1,176
T3169	11860-11867	VBD	denotes	matched
T3178	11868-11869	DT	denotes	a
T3179	11870-11875	NN	denotes	total
T3180	11876-11878	IN	denotes	of
T3181	11879-11882	CD	denotes	419
T3183	11883-11892	JJ	denotes	olfactory
T3184	11893-11901	NN	denotes	receptor
T3182	11902-11907	NNS	denotes	genes
T3185	11907-11908	:	denotes	;
T3186	11909-11912	DT	denotes	the
T3188	11913-11922	VBG	denotes	remaining
T3187	11923-11928	NNS	denotes	cDNAs
T3189	11929-11935	CC	denotes	either
T3177	11936-11943	VBD	denotes	matched
T3190	11944-11946	DT	denotes	an
T3192	11947-11956	JJ	denotes	olfactory
T3191	11957-11965	NN	denotes	receptor
T3193	11966-11971	IN	denotes	below
T3194	11972-11975	PRP$	denotes	our
T3196	11976-11978	CD	denotes	96
T3197	11978-11979	NN	denotes	%
T3199	11980-11990	NN	denotes	nucleotide
T3198	11991-11999	NN	denotes	identity
T3195	12000-12009	NN	denotes	threshold
T3200	12010-12012	CC	denotes	or
T3201	12013-12016	VBD	denotes	had
T3202	12017-12026	JJ	denotes	ambiguous
T3203	12027-12034	NNS	denotes	matches
T3204	12035-12047	VBG	denotes	encompassing
T3205	12048-12052	JJR	denotes	more
T3207	12053-12057	IN	denotes	than
T3206	12058-12061	CD	denotes	one
T3209	12062-12071	JJ	denotes	olfactory
T3210	12072-12080	NN	denotes	receptor
T3211	12081-12085	NN	denotes	gene
T3208	12086-12092	NN	denotes	region
T3212	12093-12094	-LRB-	denotes	(
T3213	12094-12097	VB	denotes	see
T3214	12098-12103	RB	denotes	below
T3215	12103-12104	-RRB-	denotes	)
T3216	12104-12105	.	denotes	.
T3520	12107-12108	DT	denotes	A
T3522	12109-12114	NN	denotes	class
T3523	12115-12116	CD	denotes	I
T3524	12117-12126	JJ	denotes	olfactory
T3525	12127-12135	NN	denotes	receptor
T3526	12136-12146	JJ	denotes	degenerate
T3521	12147-12153	NN	denotes	primer
T3527	12154-12162	VBZ	denotes	broadens
T3528	12163-12175	JJ	denotes	phylogenetic
T3529	12176-12188	NN	denotes	distribution
T3530	12189-12191	IN	denotes	of
T3531	12192-12201	VBN	denotes	confirmed
T3533	12202-12211	JJ	denotes	olfactory
T3534	12212-12220	NN	denotes	receptor
T3532	12221-12226	NNS	denotes	genes
T3535	12226-12490	sentence	denotes	Previous analyses of the mammalian olfactory receptor family define two major phylogenetic clades, referred to as class I and II olfactory receptors, and suggest that class I olfactory receptors are more similar to fish olfactory receptors than are class IIs [5].
T3536	12227-12235	JJ	denotes	Previous
T3537	12236-12244	NNS	denotes	analyses
T3539	12245-12247	IN	denotes	of
T3540	12248-12251	DT	denotes	the
T3542	12252-12261	JJ	denotes	mammalian
T3543	12262-12271	JJ	denotes	olfactory
T3544	12272-12280	NN	denotes	receptor
T3541	12281-12287	NN	denotes	family
T3538	12288-12294	VBZ	denotes	define
T3545	12295-12298	CD	denotes	two
T3547	12299-12304	JJ	denotes	major
T3548	12305-12317	JJ	denotes	phylogenetic
T3546	12318-12324	NNS	denotes	clades
T3549	12324-12326	,	denotes	,
T3550	12326-12334	VBN	denotes	referred
T3551	12335-12337	IN	denotes	to
T3552	12338-12340	IN	denotes	as
T3553	12341-12346	NN	denotes	class
T3554	12347-12348	CD	denotes	I
T3556	12349-12352	CC	denotes	and
T3557	12353-12355	CD	denotes	II
T3558	12356-12365	JJ	denotes	olfactory
T3555	12366-12375	NNS	denotes	receptors
T3559	12375-12377	,	denotes	,
T3560	12377-12380	CC	denotes	and
T3561	12381-12388	VBP	denotes	suggest
T3562	12389-12393	IN	denotes	that
T3564	12394-12399	NN	denotes	class
T3566	12400-12401	CD	denotes	I
T3567	12402-12411	JJ	denotes	olfactory
T3565	12412-12421	NNS	denotes	receptors
T3563	12422-12425	VBP	denotes	are
T3568	12426-12430	RBR	denotes	more
T3569	12431-12438	JJ	denotes	similar
T3570	12439-12441	IN	denotes	to
T3571	12442-12446	NN	denotes	fish
T3573	12447-12456	JJ	denotes	olfactory
T3572	12457-12466	NNS	denotes	receptors
T3574	12467-12471	IN	denotes	than
T3575	12472-12475	VBP	denotes	are
T3576	12476-12481	NN	denotes	class
T3577	12482-12485	NNS	denotes	IIs
T3578	12486-12487	-LRB-	denotes	[
T3579	12487-12488	CD	denotes	5
T3580	12488-12489	-RRB-	denotes	]
T3581	12489-12490	.	denotes	.
T3582	12490-12849	sentence	denotes	Figure 1 illustrates the phylogenetic diversity of our cDNA collection, showing that we have confirmed expression of at least one olfactory receptor gene in each major clade of the class II olfactory receptor genes, or 391 out of 983 (40%) of all intact class II olfactory receptor genes where full-length genomic sequence data are available (blue branches).
T3583	12491-12497	NN	denotes	Figure
T3585	12498-12499	CD	denotes	1
T3584	12500-12511	VBZ	denotes	illustrates
T3586	12512-12515	DT	denotes	the
T3588	12516-12528	JJ	denotes	phylogenetic
T3587	12529-12538	NN	denotes	diversity
T3589	12539-12541	IN	denotes	of
T3590	12542-12545	PRP$	denotes	our
T3592	12546-12550	NN	denotes	cDNA
T3591	12551-12561	NN	denotes	collection
T3593	12561-12563	,	denotes	,
T3594	12563-12570	VBG	denotes	showing
T3595	12571-12575	IN	denotes	that
T3597	12576-12578	PRP	denotes	we
T3598	12579-12583	VBP	denotes	have
T3596	12584-12593	VBN	denotes	confirmed
T3599	12594-12604	NN	denotes	expression
T3600	12605-12607	IN	denotes	of
T3601	12608-12610	RB	denotes	at
T3603	12611-12616	RBS	denotes	least
T3602	12617-12620	CD	denotes	one
T3605	12621-12630	JJ	denotes	olfactory
T3606	12631-12639	NN	denotes	receptor
T3604	12640-12644	NN	denotes	gene
T3607	12645-12647	IN	denotes	in
T3608	12648-12652	DT	denotes	each
T3610	12653-12658	JJ	denotes	major
T3609	12659-12664	NN	denotes	clade
T3611	12665-12667	IN	denotes	of
T3612	12668-12671	DT	denotes	the
T3614	12672-12677	NN	denotes	class
T3615	12678-12680	CD	denotes	II
T3616	12681-12690	JJ	denotes	olfactory
T3617	12691-12699	NN	denotes	receptor
T3613	12700-12705	NNS	denotes	genes
T3618	12705-12707	,	denotes	,
T3619	12707-12709	CC	denotes	or
T3620	12710-12713	CD	denotes	391
T3622	12714-12717	IN	denotes	out
T3623	12718-12720	IN	denotes	of
T3621	12721-12724	CD	denotes	983
T3624	12725-12726	-LRB-	denotes	(
T3626	12726-12728	CD	denotes	40
T3625	12728-12729	NN	denotes	%
T3627	12729-12730	-RRB-	denotes	)
T3628	12731-12733	IN	denotes	of
T3629	12734-12737	DT	denotes	all
T3631	12738-12744	JJ	denotes	intact
T3632	12745-12750	NN	denotes	class
T3633	12751-12753	CD	denotes	II
T3634	12754-12763	JJ	denotes	olfactory
T3635	12764-12772	NN	denotes	receptor
T3630	12773-12778	NNS	denotes	genes
T3636	12779-12784	WRB	denotes	where
T3638	12785-12789	JJ	denotes	full
T3640	12789-12790	HYPH	denotes	-
T3639	12790-12796	NN	denotes	length
T3642	12797-12804	JJ	denotes	genomic
T3643	12805-12813	NN	denotes	sequence
T3641	12814-12818	NNS	denotes	data
T3637	12819-12822	VBP	denotes	are
T3644	12823-12832	JJ	denotes	available
T3645	12833-12834	-LRB-	denotes	(
T3647	12834-12838	JJ	denotes	blue
T3646	12839-12847	NNS	denotes	branches
T3648	12847-12848	-RRB-	denotes	)
T3649	12848-12849	.	denotes	.
T3650	12849-12942	sentence	denotes	The screen thus appears relatively unbiased in its coverage of class II olfactory receptors.
T3651	12850-12853	DT	denotes	The
T3652	12854-12860	NN	denotes	screen
T3654	12861-12865	RB	denotes	thus
T3653	12866-12873	VBZ	denotes	appears
T3655	12874-12884	RB	denotes	relatively
T3656	12885-12893	JJ	denotes	unbiased
T3657	12894-12896	IN	denotes	in
T3658	12897-12900	PRP$	denotes	its
T3659	12901-12909	NN	denotes	coverage
T3660	12910-12912	IN	denotes	of
T3661	12913-12918	NN	denotes	class
T3663	12919-12921	CD	denotes	II
T3664	12922-12931	JJ	denotes	olfactory
T3662	12932-12941	NNS	denotes	receptors
T3665	12941-12942	.	denotes	.
T3666	12942-13057	sentence	denotes	However, our random screen provided cDNAs for only two out of 124 intact, full-length class I olfactory receptors.
T3667	12943-12950	RB	denotes	However
T3669	12950-12952	,	denotes	,
T3670	12952-12955	PRP$	denotes	our
T3672	12956-12962	JJ	denotes	random
T3671	12963-12969	NN	denotes	screen
T3668	12970-12978	VBD	denotes	provided
T3673	12979-12984	NNS	denotes	cDNAs
T3674	12985-12988	IN	denotes	for
T3675	12989-12993	RB	denotes	only
T3677	12994-12997	CD	denotes	two
T3678	12998-13001	IN	denotes	out
T3679	13002-13004	IN	denotes	of
T3676	13005-13008	CD	denotes	124
T3681	13009-13015	JJ	denotes	intact
T3682	13015-13017	,	denotes	,
T3683	13017-13021	JJ	denotes	full
T3685	13021-13022	HYPH	denotes	-
T3684	13022-13028	NN	denotes	length
T3686	13029-13034	NN	denotes	class
T3687	13035-13036	CD	denotes	I
T3688	13037-13046	JJ	denotes	olfactory
T3680	13047-13056	NNS	denotes	receptors
T3689	13056-13057	.	denotes	.
T3690	13057-13240	sentence	denotes	In an attempt to broaden the phylogenetic coverage of our hybridization screen, we used additional degenerate probes on the adult library and screened an embryonic library (Table 2).
T3691	13058-13060	IN	denotes	In
T3693	13061-13063	DT	denotes	an
T3694	13064-13071	NN	denotes	attempt
T3695	13072-13074	TO	denotes	to
T3696	13075-13082	VB	denotes	broaden
T3697	13083-13086	DT	denotes	the
T3699	13087-13099	JJ	denotes	phylogenetic
T3698	13100-13108	NN	denotes	coverage
T3700	13109-13111	IN	denotes	of
T3701	13112-13115	PRP$	denotes	our
T3703	13116-13129	NN	denotes	hybridization
T3702	13130-13136	NN	denotes	screen
T3704	13136-13138	,	denotes	,
T3705	13138-13140	PRP	denotes	we
T3692	13141-13145	VBD	denotes	used
T3706	13146-13156	JJ	denotes	additional
T3708	13157-13167	JJ	denotes	degenerate
T3707	13168-13174	NNS	denotes	probes
T3709	13175-13177	IN	denotes	on
T3710	13178-13181	DT	denotes	the
T3712	13182-13187	JJ	denotes	adult
T3711	13188-13195	NN	denotes	library
T3713	13196-13199	CC	denotes	and
T3714	13200-13208	VBD	denotes	screened
T3715	13209-13211	DT	denotes	an
T3717	13212-13221	JJ	denotes	embryonic
T3716	13222-13229	NN	denotes	library
T3718	13230-13231	-LRB-	denotes	(
T3719	13231-13236	NN	denotes	Table
T3720	13237-13238	CD	denotes	2
T3721	13238-13239	-RRB-	denotes	)
T3722	13239-13240	.	denotes	.
T3723	13240-13323	sentence	denotes	These experiments did not increase the diversity of clones identified (not shown).
T3724	13241-13246	DT	denotes	These
T3725	13247-13258	NNS	denotes	experiments
T3727	13259-13262	VBD	denotes	did
T3728	13263-13266	RB	denotes	not
T3726	13267-13275	VB	denotes	increase
T3729	13276-13279	DT	denotes	the
T3730	13280-13289	NN	denotes	diversity
T3731	13290-13292	IN	denotes	of
T3732	13293-13299	NNS	denotes	clones
T3733	13300-13310	VBN	denotes	identified
T3734	13311-13312	-LRB-	denotes	(
T3736	13312-13315	RB	denotes	not
T3735	13316-13321	VBN	denotes	shown
T3737	13321-13322	-RRB-	denotes	)
T3738	13322-13323	.	denotes	.
T3739	13323-13476	sentence	denotes	This severe class I underrepresentation could be due to experimental bias - a consequence of using degenerate primers to create our hybridization probe.
T3740	13324-13328	DT	denotes	This
T3742	13329-13335	JJ	denotes	severe
T3743	13336-13341	NN	denotes	class
T3744	13342-13343	CD	denotes	I
T3741	13344-13363	NN	denotes	underrepresentation
T3746	13364-13369	MD	denotes	could
T3745	13370-13372	VB	denotes	be
T3747	13373-13376	IN	denotes	due
T3748	13377-13379	IN	denotes	to
T3749	13380-13392	JJ	denotes	experimental
T3750	13393-13397	NN	denotes	bias
T3751	13398-13399	:	denotes	-
T3752	13400-13401	DT	denotes	a
T3753	13402-13413	NN	denotes	consequence
T3754	13414-13416	IN	denotes	of
T3755	13417-13422	VBG	denotes	using
T3756	13423-13433	JJ	denotes	degenerate
T3757	13434-13441	NNS	denotes	primers
T3758	13442-13444	TO	denotes	to
T3759	13445-13451	VB	denotes	create
T3760	13452-13455	PRP$	denotes	our
T3762	13456-13469	NN	denotes	hybridization
T3761	13470-13475	NN	denotes	probe
T3763	13475-13476	.	denotes	.
T3764	13476-13577	sentence	denotes	Alternatively, class I genes might be expressed at extremely low levels in the olfactory epithelium.
T3765	13477-13490	RB	denotes	Alternatively
T3767	13490-13492	,	denotes	,
T3768	13492-13497	NN	denotes	class
T3770	13498-13499	CD	denotes	I
T3769	13500-13505	NNS	denotes	genes
T3771	13506-13511	MD	denotes	might
T3772	13512-13514	VB	denotes	be
T3766	13515-13524	VBN	denotes	expressed
T3773	13525-13527	IN	denotes	at
T3774	13528-13537	RB	denotes	extremely
T3775	13538-13541	JJ	denotes	low
T3776	13542-13548	NNS	denotes	levels
T3777	13549-13551	IN	denotes	in
T3778	13552-13555	DT	denotes	the
T3780	13556-13565	JJ	denotes	olfactory
T3779	13566-13576	NN	denotes	epithelium
T3781	13576-13577	.	denotes	.
T3782	13577-13850	sentence	denotes	In order to determine whether class I olfactory receptors are expressed in the olfactory epithelium, we designed a reverse-strand degenerate primer to recognize a motif in transmembrane domain 7 (PP{V/M/A/T}{F/L/I/M}NP) enriched among class I olfactory receptor sequences.
T3783	13578-13580	IN	denotes	In
T3785	13581-13586	NN	denotes	order
T3786	13587-13589	TO	denotes	to
T3787	13590-13599	VB	denotes	determine
T3788	13600-13607	IN	denotes	whether
T3790	13608-13613	NN	denotes	class
T3792	13614-13615	CD	denotes	I
T3793	13616-13625	JJ	denotes	olfactory
T3791	13626-13635	NNS	denotes	receptors
T3794	13636-13639	VBP	denotes	are
T3789	13640-13649	VBN	denotes	expressed
T3795	13650-13652	IN	denotes	in
T3796	13653-13656	DT	denotes	the
T3798	13657-13666	JJ	denotes	olfactory
T3797	13667-13677	NN	denotes	epithelium
T3799	13677-13679	,	denotes	,
T3800	13679-13681	PRP	denotes	we
T3784	13682-13690	VBD	denotes	designed
T3801	13691-13692	DT	denotes	a
T3803	13693-13700	JJ	denotes	reverse
T3805	13700-13701	HYPH	denotes	-
T3804	13701-13707	NN	denotes	strand
T3806	13708-13718	JJ	denotes	degenerate
T3802	13719-13725	NN	denotes	primer
T3807	13726-13728	TO	denotes	to
T3808	13729-13738	VB	denotes	recognize
T3809	13739-13740	DT	denotes	a
T3810	13741-13746	NN	denotes	motif
T3811	13747-13749	IN	denotes	in
T3812	13750-13763	NN	denotes	transmembrane
T3813	13764-13770	NN	denotes	domain
T3814	13771-13772	CD	denotes	7
T3815	13773-13774	-LRB-	denotes	(
T3817	13774-13776	NN	denotes	PP
T3818	13776-13777	-LRB-	denotes	{
T3819	13777-13778	NN	denotes	V
T3821	13778-13779	HYPH	denotes	/
T3822	13779-13780	NN	denotes	M
T3823	13780-13781	HYPH	denotes	/
T3824	13781-13782	NN	denotes	A
T3825	13782-13783	HYPH	denotes	/
T3820	13783-13784	NN	denotes	T
T3826	13784-13785	RBR	denotes	}
T3827	13785-13786	-LRB-	denotes	{
T3828	13786-13787	NN	denotes	F
T3830	13787-13788	HYPH	denotes	/
T3831	13788-13789	NN	denotes	L
T3832	13789-13790	HYPH	denotes	/
T3833	13790-13791	NN	denotes	I
T3834	13791-13792	HYPH	denotes	/
T3829	13792-13793	NN	denotes	M
T3835	13793-13794	RBR	denotes	}
T3816	13794-13796	NN	denotes	NP
T3836	13796-13797	-RRB-	denotes	)
T3837	13798-13806	VBN	denotes	enriched
T3838	13807-13812	IN	denotes	among
T3839	13813-13818	NN	denotes	class
T3841	13819-13820	CD	denotes	I
T3842	13821-13830	JJ	denotes	olfactory
T3843	13831-13839	NN	denotes	receptor
T3840	13840-13849	NNS	denotes	sequences
T3844	13849-13850	.	denotes	.
T3845	13850-14077	sentence	denotes	Most of the motif is shared among all olfactory receptors, but the first proline residue (at the primer's 3' end) is found in 121 out of 124 (98%) intact class I genes compared to only 37 out of 983 (4%) intact class II genes.
T3846	13851-13855	JJS	denotes	Most
T3848	13856-13858	IN	denotes	of
T3849	13859-13862	DT	denotes	the
T3850	13863-13868	NN	denotes	motif
T3851	13869-13871	VBZ	denotes	is
T3847	13872-13878	VBN	denotes	shared
T3852	13879-13884	IN	denotes	among
T3853	13885-13888	DT	denotes	all
T3855	13889-13898	JJ	denotes	olfactory
T3854	13899-13908	NNS	denotes	receptors
T3856	13908-13910	,	denotes	,
T3857	13910-13913	CC	denotes	but
T3858	13914-13917	DT	denotes	the
T3860	13918-13923	JJ	denotes	first
T3861	13924-13931	NN	denotes	proline
T3859	13932-13939	NN	denotes	residue
T3863	13940-13941	-LRB-	denotes	(
T3864	13941-13943	IN	denotes	at
T3865	13944-13947	DT	denotes	the
T3866	13948-13954	NN	denotes	primer
T3868	13954-13956	POS	denotes	's
T3869	13957-13958	CD	denotes	3
T3870	13958-13959	SYM	denotes	'
T3867	13960-13963	NN	denotes	end
T3871	13963-13964	-RRB-	denotes	)
T3872	13965-13967	VBZ	denotes	is
T3862	13968-13973	VBN	denotes	found
T3873	13974-13976	IN	denotes	in
T3874	13977-13980	CD	denotes	121
T3876	13981-13984	IN	denotes	out
T3877	13985-13987	IN	denotes	of
T3875	13988-13991	CD	denotes	124
T3879	13992-13993	-LRB-	denotes	(
T3881	13993-13995	CD	denotes	98
T3880	13995-13996	NN	denotes	%
T3882	13996-13997	-RRB-	denotes	)
T3883	13998-14004	JJ	denotes	intact
T3884	14005-14010	NN	denotes	class
T3885	14011-14012	CD	denotes	I
T3878	14013-14018	NNS	denotes	genes
T3886	14019-14027	VBN	denotes	compared
T3887	14028-14030	IN	denotes	to
T3888	14031-14035	RB	denotes	only
T3890	14036-14038	CD	denotes	37
T3891	14039-14042	IN	denotes	out
T3892	14043-14045	IN	denotes	of
T3889	14046-14049	CD	denotes	983
T3894	14050-14051	-LRB-	denotes	(
T3896	14051-14052	CD	denotes	4
T3895	14052-14053	NN	denotes	%
T3897	14053-14054	-RRB-	denotes	)
T3898	14055-14061	JJ	denotes	intact
T3899	14062-14067	NN	denotes	class
T3900	14068-14070	CD	denotes	II
T3893	14071-14076	NNS	denotes	genes
T3901	14076-14077	.	denotes	.
T3902	14077-14473	sentence	denotes	When combined with another olfactory receptor degenerate primer, P26 [17], this primer preferentially amplifies class I olfactory receptors from mouse genomic DNA: of 33 sequenced, cloned PCR products, 17 represented seven different class I olfactory receptors, six represented three different class II olfactory receptors, and ten represented five different non-olfactory receptor contaminants.
T3903	14078-14082	WRB	denotes	When
T3904	14083-14091	VBN	denotes	combined
T3906	14092-14096	IN	denotes	with
T3907	14097-14104	DT	denotes	another
T3909	14105-14114	JJ	denotes	olfactory
T3910	14115-14123	NN	denotes	receptor
T3911	14124-14134	JJ	denotes	degenerate
T3908	14135-14141	NN	denotes	primer
T3912	14141-14143	,	denotes	,
T3913	14143-14146	NN	denotes	P26
T3914	14147-14148	-LRB-	denotes	[
T3915	14148-14150	CD	denotes	17
T3916	14150-14151	-RRB-	denotes	]
T3917	14151-14153	,	denotes	,
T3918	14153-14157	DT	denotes	this
T3919	14158-14164	NN	denotes	primer
T3920	14165-14179	RB	denotes	preferentially
T3905	14180-14189	VBZ	denotes	amplifies
T3922	14190-14195	NN	denotes	class
T3924	14196-14197	CD	denotes	I
T3925	14198-14207	JJ	denotes	olfactory
T3923	14208-14217	NNS	denotes	receptors
T3926	14218-14222	IN	denotes	from
T3927	14223-14228	NN	denotes	mouse
T3929	14229-14236	JJ	denotes	genomic
T3928	14237-14240	NN	denotes	DNA
T3930	14240-14242	:	denotes	:
T3931	14242-14244	IN	denotes	of
T3932	14245-14247	CD	denotes	33
T3934	14248-14257	JJ	denotes	sequenced
T3935	14257-14259	,	denotes	,
T3936	14259-14265	JJ	denotes	cloned
T3937	14266-14269	NN	denotes	PCR
T3933	14270-14278	NNS	denotes	products
T3938	14278-14280	,	denotes	,
T3939	14280-14282	CD	denotes	17
T3921	14283-14294	VBD	denotes	represented
T3940	14295-14300	CD	denotes	seven
T3942	14301-14310	JJ	denotes	different
T3943	14311-14316	NN	denotes	class
T3944	14317-14318	CD	denotes	I
T3945	14319-14328	JJ	denotes	olfactory
T3941	14329-14338	NNS	denotes	receptors
T3946	14338-14340	,	denotes	,
T3947	14340-14343	CD	denotes	six
T3948	14344-14355	VBD	denotes	represented
T3949	14356-14361	CD	denotes	three
T3951	14362-14371	JJ	denotes	different
T3952	14372-14377	NN	denotes	class
T3953	14378-14380	CD	denotes	II
T3954	14381-14390	JJ	denotes	olfactory
T3950	14391-14400	NNS	denotes	receptors
T3955	14400-14402	,	denotes	,
T3956	14402-14405	CC	denotes	and
T3957	14406-14409	CD	denotes	ten
T3958	14410-14421	VBD	denotes	represented
T3959	14422-14426	CD	denotes	five
T3961	14427-14436	JJ	denotes	different
T3962	14437-14450	JJ	denotes	non-olfactory
T3963	14451-14459	NN	denotes	receptor
T3960	14460-14472	NNS	denotes	contaminants
T3964	14472-14473	.	denotes	.
T3965	14473-14700	sentence	denotes	Degenerate PCR, cloning, and sequencing from reverse-transcribed olfactory epithelium RNA showed that at least seven class I olfactory receptors are expressed, as well as one additional class II gene (colored red in Figure 1).
T3966	14474-14484	JJ	denotes	Degenerate
T3967	14485-14488	NN	denotes	PCR
T3969	14488-14490	,	denotes	,
T3970	14490-14497	NN	denotes	cloning
T3971	14497-14499	,	denotes	,
T3972	14499-14502	CC	denotes	and
T3973	14503-14513	NN	denotes	sequencing
T3974	14514-14518	IN	denotes	from
T3975	14519-14526	JJ	denotes	reverse
T3977	14526-14527	HYPH	denotes	-
T3976	14527-14538	VBN	denotes	transcribed
T3979	14539-14548	JJ	denotes	olfactory
T3980	14549-14559	NN	denotes	epithelium
T3978	14560-14563	NN	denotes	RNA
T3968	14564-14570	VBD	denotes	showed
T3981	14571-14575	IN	denotes	that
T3983	14576-14578	RB	denotes	at
T3985	14579-14584	RBS	denotes	least
T3984	14585-14590	CD	denotes	seven
T3987	14591-14596	NN	denotes	class
T3988	14597-14598	CD	denotes	I
T3989	14599-14608	JJ	denotes	olfactory
T3986	14609-14618	NNS	denotes	receptors
T3990	14619-14622	VBP	denotes	are
T3982	14623-14632	VBN	denotes	expressed
T3991	14632-14634	,	denotes	,
T3992	14634-14636	RB	denotes	as
T3994	14637-14641	RB	denotes	well
T3993	14642-14644	IN	denotes	as
T3996	14645-14648	CD	denotes	one
T3997	14649-14659	JJ	denotes	additional
T3998	14660-14665	NN	denotes	class
T3999	14666-14668	CD	denotes	II
T3995	14669-14673	NN	denotes	gene
T4000	14674-14675	-LRB-	denotes	(
T4001	14675-14682	VBN	denotes	colored
T4002	14683-14686	JJ	denotes	red
T4003	14687-14689	IN	denotes	in
T4004	14690-14696	NN	denotes	Figure
T4005	14697-14698	CD	denotes	1
T4006	14698-14699	-RRB-	denotes	)
T4007	14699-14700	.	denotes	.
T4008	14700-14915	sentence	denotes	However, no products could be obtained from the adult or the fetal olfactory epithelium cDNA libraries using the class I primer, suggesting that the libraries contain very low levels of class I olfactory receptors.
T4009	14701-14708	RB	denotes	However
T4011	14708-14710	,	denotes	,
T4012	14710-14712	DT	denotes	no
T4013	14713-14721	NNS	denotes	products
T4014	14722-14727	MD	denotes	could
T4015	14728-14730	VB	denotes	be
T4010	14731-14739	VBN	denotes	obtained
T4016	14740-14744	IN	denotes	from
T4017	14745-14748	DT	denotes	the
T4018	14749-14754	JJ	denotes	adult
T4019	14755-14757	CC	denotes	or
T4020	14758-14761	DT	denotes	the
T4021	14762-14767	JJ	denotes	fetal
T4023	14768-14777	JJ	denotes	olfactory
T4024	14778-14788	NN	denotes	epithelium
T4025	14789-14793	NN	denotes	cDNA
T4022	14794-14803	NNS	denotes	libraries
T4026	14804-14809	VBG	denotes	using
T4027	14810-14813	DT	denotes	the
T4029	14814-14819	NN	denotes	class
T4030	14820-14821	CD	denotes	I
T4028	14822-14828	NN	denotes	primer
T4031	14828-14830	,	denotes	,
T4032	14830-14840	VBG	denotes	suggesting
T4033	14841-14845	IN	denotes	that
T4035	14846-14849	DT	denotes	the
T4036	14850-14859	NNS	denotes	libraries
T4034	14860-14867	VBP	denotes	contain
T4037	14868-14872	RB	denotes	very
T4038	14873-14876	JJ	denotes	low
T4039	14877-14883	NNS	denotes	levels
T4040	14884-14886	IN	denotes	of
T4041	14887-14892	NN	denotes	class
T4043	14893-14894	CD	denotes	I
T4044	14895-14904	JJ	denotes	olfactory
T4042	14905-14914	NNS	denotes	receptors
T4045	14914-14915	.	denotes	.
T4046	14915-15199	sentence	denotes	We also confirmed expression of nine additional olfactory receptors (three class I and six class II, colored red in Figure 1) from subclades that were poorly represented in our cDNA screen using gene-specific primer pairs to amplify cDNA library or reverse-transcribed RNA templates.
T4047	14916-14918	PRP	denotes	We
T4049	14919-14923	RB	denotes	also
T4048	14924-14933	VBD	denotes	confirmed
T4050	14934-14944	NN	denotes	expression
T4051	14945-14947	IN	denotes	of
T4052	14948-14952	CD	denotes	nine
T4054	14953-14963	JJ	denotes	additional
T4055	14964-14973	JJ	denotes	olfactory
T4053	14974-14983	NNS	denotes	receptors
T4056	14984-14985	-LRB-	denotes	(
T4057	14985-14990	CD	denotes	three
T4058	14991-14996	NN	denotes	class
T4059	14997-14998	CD	denotes	I
T4060	14999-15002	CC	denotes	and
T4061	15003-15006	CD	denotes	six
T4062	15007-15012	NN	denotes	class
T4063	15013-15015	CD	denotes	II
T4064	15015-15017	,	denotes	,
T4065	15017-15024	VBN	denotes	colored
T4066	15025-15028	JJ	denotes	red
T4067	15029-15031	IN	denotes	in
T4068	15032-15038	NN	denotes	Figure
T4069	15039-15040	CD	denotes	1
T4070	15040-15041	-RRB-	denotes	)
T4071	15042-15046	IN	denotes	from
T4072	15047-15056	NNS	denotes	subclades
T4073	15057-15061	WDT	denotes	that
T4075	15062-15066	VBD	denotes	were
T4076	15067-15073	RB	denotes	poorly
T4074	15074-15085	VBN	denotes	represented
T4077	15086-15088	IN	denotes	in
T4078	15089-15092	PRP$	denotes	our
T4080	15093-15097	NN	denotes	cDNA
T4079	15098-15104	NN	denotes	screen
T4081	15105-15110	VBG	denotes	using
T4082	15111-15115	NN	denotes	gene
T4084	15115-15116	HYPH	denotes	-
T4083	15116-15124	JJ	denotes	specific
T4086	15125-15131	NN	denotes	primer
T4085	15132-15137	NNS	denotes	pairs
T4087	15138-15140	TO	denotes	to
T4088	15141-15148	VB	denotes	amplify
T4089	15149-15153	NN	denotes	cDNA
T4090	15154-15161	NN	denotes	library
T4091	15162-15164	CC	denotes	or
T4092	15165-15172	JJ	denotes	reverse
T4094	15172-15173	HYPH	denotes	-
T4093	15173-15184	VBN	denotes	transcribed
T4096	15185-15188	NN	denotes	RNA
T4095	15189-15198	NNS	denotes	templates
T4097	15198-15199	.	denotes	.
T4098	15199-15338	sentence	denotes	For two of the class I genes we had shown to be expressed, we determined relative transcript levels using quantitative RT-PCR (see below).
T4099	15200-15203	IN	denotes	For
T4101	15204-15207	CD	denotes	two
T4102	15208-15210	IN	denotes	of
T4103	15211-15214	DT	denotes	the
T4105	15215-15220	NN	denotes	class
T4106	15221-15222	CD	denotes	I
T4104	15223-15228	NNS	denotes	genes
T4107	15229-15231	PRP	denotes	we
T4109	15232-15235	VBD	denotes	had
T4108	15236-15241	VBN	denotes	shown
T4110	15242-15244	TO	denotes	to
T4112	15245-15247	VB	denotes	be
T4111	15248-15257	VBN	denotes	expressed
T4113	15257-15259	,	denotes	,
T4114	15259-15261	PRP	denotes	we
T4100	15262-15272	VBD	denotes	determined
T4115	15273-15281	JJ	denotes	relative
T4117	15282-15292	NN	denotes	transcript
T4116	15293-15299	NNS	denotes	levels
T4118	15300-15305	VBG	denotes	using
T4119	15306-15318	JJ	denotes	quantitative
T4121	15319-15321	NN	denotes	RT
T4122	15321-15322	HYPH	denotes	-
T4120	15322-15325	NN	denotes	PCR
T4123	15326-15327	-LRB-	denotes	(
T4124	15327-15330	VB	denotes	see
T4125	15331-15336	RB	denotes	below
T4126	15336-15337	-RRB-	denotes	)
T4127	15337-15338	.	denotes	.
T4128	15338-15667	sentence	denotes	Expression levels were similar to those observed for genes that were represented in our cDNA collection, suggesting that class I olfactory receptors are not under-represented in the olfactory epithelium, and that the dearth of class I cDNAs in our screen is likely to be due to bias in the libraries and/or hybridization probes.
T4129	15339-15349	NN	denotes	Expression
T4130	15350-15356	NNS	denotes	levels
T4131	15357-15361	VBD	denotes	were
T4132	15362-15369	JJ	denotes	similar
T4133	15370-15372	IN	denotes	to
T4134	15373-15378	DT	denotes	those
T4135	15379-15387	VBN	denotes	observed
T4136	15388-15391	IN	denotes	for
T4137	15392-15397	NNS	denotes	genes
T4138	15398-15402	WDT	denotes	that
T4140	15403-15407	VBD	denotes	were
T4139	15408-15419	VBN	denotes	represented
T4141	15420-15422	IN	denotes	in
T4142	15423-15426	PRP$	denotes	our
T4144	15427-15431	NN	denotes	cDNA
T4143	15432-15442	NN	denotes	collection
T4145	15442-15444	,	denotes	,
T4146	15444-15454	VBG	denotes	suggesting
T4147	15455-15459	IN	denotes	that
T4149	15460-15465	NN	denotes	class
T4151	15466-15467	CD	denotes	I
T4152	15468-15477	JJ	denotes	olfactory
T4150	15478-15487	NNS	denotes	receptors
T4153	15488-15491	VBP	denotes	are
T4154	15492-15495	RB	denotes	not
T4155	15496-15501	RB	denotes	under
T4156	15501-15502	HYPH	denotes	-
T4148	15502-15513	VBN	denotes	represented
T4157	15514-15516	IN	denotes	in
T4158	15517-15520	DT	denotes	the
T4160	15521-15530	JJ	denotes	olfactory
T4159	15531-15541	NN	denotes	epithelium
T4161	15541-15543	,	denotes	,
T4162	15543-15546	CC	denotes	and
T4163	15547-15551	IN	denotes	that
T4165	15552-15555	DT	denotes	the
T4166	15556-15562	NN	denotes	dearth
T4167	15563-15565	IN	denotes	of
T4168	15566-15571	NN	denotes	class
T4170	15572-15573	CD	denotes	I
T4169	15574-15579	NNS	denotes	cDNAs
T4171	15580-15582	IN	denotes	in
T4172	15583-15586	PRP$	denotes	our
T4173	15587-15593	NN	denotes	screen
T4164	15594-15596	VBZ	denotes	is
T4174	15597-15603	JJ	denotes	likely
T4175	15604-15606	TO	denotes	to
T4176	15607-15609	VB	denotes	be
T4177	15610-15613	IN	denotes	due
T4178	15614-15616	IN	denotes	to
T4179	15617-15621	NN	denotes	bias
T4180	15622-15624	IN	denotes	in
T4181	15625-15628	DT	denotes	the
T4182	15629-15638	NNS	denotes	libraries
T4183	15639-15642	CC	denotes	and
T4184	15642-15643	HYPH	denotes	/
T4185	15643-15645	CC	denotes	or
T4186	15646-15659	NN	denotes	hybridization
T4187	15660-15666	NNS	denotes	probes
T4188	15666-15667	.	denotes	.
T4620	15669-15673	DT	denotes	Some
T4622	15674-15683	JJ	denotes	olfactory
T4623	15684-15692	NN	denotes	receptor
T4621	15693-15698	NNS	denotes	genes
T4625	15699-15702	VBP	denotes	are
T4624	15703-15712	VBN	denotes	expressed
T4626	15713-15715	IN	denotes	at
T4627	15716-15722	JJR	denotes	higher
T4628	15723-15729	NNS	denotes	levels
T4629	15730-15734	IN	denotes	than
T4630	15735-15741	NNS	denotes	others
T4631	15741-15845	sentence	denotes	Our cDNA screen suggests that some olfactory receptor genes are expressed at higher levels than others.
T4632	15742-15745	PRP$	denotes	Our
T4634	15746-15750	NN	denotes	cDNA
T4633	15751-15757	NN	denotes	screen
T4635	15758-15766	VBZ	denotes	suggests
T4636	15767-15771	IN	denotes	that
T4638	15772-15776	DT	denotes	some
T4640	15777-15786	JJ	denotes	olfactory
T4641	15787-15795	NN	denotes	receptor
T4639	15796-15801	NNS	denotes	genes
T4642	15802-15805	VBP	denotes	are
T4637	15806-15815	VBN	denotes	expressed
T4643	15816-15818	IN	denotes	at
T4644	15819-15825	JJR	denotes	higher
T4645	15826-15832	NNS	denotes	levels
T4646	15833-15837	IN	denotes	than
T4647	15838-15844	NNS	denotes	others
T4648	15844-15845	.	denotes	.
T4649	15845-16326	sentence	denotes	If all olfactory receptor genes were expressed at equal levels, and our screen and library were unbiased in their coverage of the class II olfactory receptors, the number of cDNAs detected per class II olfactory receptor should follow a Poisson distribution, calculated based on the assumption that all 983 intact class II olfactory receptors have an equal chance of being represented in the screen, but that class I olfactory receptors and pseudogenes cannot be found (Figure 2).
T4650	15846-15848	IN	denotes	If
T4652	15849-15852	DT	denotes	all
T4654	15853-15862	JJ	denotes	olfactory
T4655	15863-15871	NN	denotes	receptor
T4653	15872-15877	NNS	denotes	genes
T4656	15878-15882	VBD	denotes	were
T4651	15883-15892	VBN	denotes	expressed
T4658	15893-15895	IN	denotes	at
T4659	15896-15901	JJ	denotes	equal
T4660	15902-15908	NNS	denotes	levels
T4661	15908-15910	,	denotes	,
T4662	15910-15913	CC	denotes	and
T4663	15914-15917	PRP$	denotes	our
T4664	15918-15924	NN	denotes	screen
T4666	15925-15928	CC	denotes	and
T4667	15929-15936	NN	denotes	library
T4665	15937-15941	VBD	denotes	were
T4668	15942-15950	JJ	denotes	unbiased
T4669	15951-15953	IN	denotes	in
T4670	15954-15959	PRP$	denotes	their
T4671	15960-15968	NN	denotes	coverage
T4672	15969-15971	IN	denotes	of
T4673	15972-15975	DT	denotes	the
T4675	15976-15981	NN	denotes	class
T4676	15982-15984	CD	denotes	II
T4677	15985-15994	JJ	denotes	olfactory
T4674	15995-16004	NNS	denotes	receptors
T4678	16004-16006	,	denotes	,
T4679	16006-16009	DT	denotes	the
T4680	16010-16016	NN	denotes	number
T4681	16017-16019	IN	denotes	of
T4682	16020-16025	NNS	denotes	cDNAs
T4683	16026-16034	VBN	denotes	detected
T4684	16035-16038	IN	denotes	per
T4685	16039-16044	NN	denotes	class
T4687	16045-16047	CD	denotes	II
T4688	16048-16057	JJ	denotes	olfactory
T4686	16058-16066	NN	denotes	receptor
T4689	16067-16073	MD	denotes	should
T4657	16074-16080	VB	denotes	follow
T4690	16081-16082	DT	denotes	a
T4692	16083-16090	NNP	denotes	Poisson
T4691	16091-16103	NN	denotes	distribution
T4693	16103-16105	,	denotes	,
T4694	16105-16115	VBN	denotes	calculated
T4695	16116-16121	VBN	denotes	based
T4696	16122-16124	IN	denotes	on
T4697	16125-16128	DT	denotes	the
T4698	16129-16139	NN	denotes	assumption
T4699	16140-16144	IN	denotes	that
T4701	16145-16148	DT	denotes	all
T4703	16149-16152	CD	denotes	983
T4704	16153-16159	JJ	denotes	intact
T4705	16160-16165	NN	denotes	class
T4706	16166-16168	CD	denotes	II
T4707	16169-16178	JJ	denotes	olfactory
T4702	16179-16188	NNS	denotes	receptors
T4700	16189-16193	VBP	denotes	have
T4708	16194-16196	DT	denotes	an
T4710	16197-16202	JJ	denotes	equal
T4709	16203-16209	NN	denotes	chance
T4711	16210-16212	IN	denotes	of
T4712	16213-16218	VBG	denotes	being
T4713	16219-16230	VBN	denotes	represented
T4714	16231-16233	IN	denotes	in
T4715	16234-16237	DT	denotes	the
T4716	16238-16244	NN	denotes	screen
T4717	16244-16246	,	denotes	,
T4718	16246-16249	CC	denotes	but
T4719	16250-16254	IN	denotes	that
T4721	16255-16260	NN	denotes	class
T4723	16261-16262	CD	denotes	I
T4724	16263-16272	JJ	denotes	olfactory
T4722	16273-16282	NNS	denotes	receptors
T4725	16283-16286	CC	denotes	and
T4726	16287-16298	NNS	denotes	pseudogenes
T4727	16299-16302	MD	denotes	can
T4728	16302-16305	RB	denotes	not
T4729	16306-16308	VB	denotes	be
T4720	16309-16314	VBN	denotes	found
T4730	16315-16316	-LRB-	denotes	(
T4731	16316-16322	NN	denotes	Figure
T4732	16323-16324	CD	denotes	2
T4733	16324-16325	-RRB-	denotes	)
T4734	16325-16326	.	denotes	.
T4735	16326-16527	sentence	denotes	We calculate a low probability (approximately one in 28) that we would observe any gene with at least eight matching cDNAs in the set of 1,176 cDNAs we assigned to single olfactory receptor sequences.
T4736	16327-16329	PRP	denotes	We
T4737	16330-16339	VBP	denotes	calculate
T4738	16340-16341	DT	denotes	a
T4740	16342-16345	JJ	denotes	low
T4739	16346-16357	NN	denotes	probability
T4741	16358-16359	-LRB-	denotes	(
T4743	16359-16372	RB	denotes	approximately
T4742	16373-16376	CD	denotes	one
T4744	16377-16379	IN	denotes	in
T4745	16380-16382	CD	denotes	28
T4746	16382-16383	-RRB-	denotes	)
T4747	16384-16388	IN	denotes	that
T4749	16389-16391	PRP	denotes	we
T4750	16392-16397	MD	denotes	would
T4748	16398-16405	VB	denotes	observe
T4751	16406-16409	DT	denotes	any
T4752	16410-16414	NN	denotes	gene
T4753	16415-16419	IN	denotes	with
T4754	16420-16422	RB	denotes	at
T4756	16423-16428	RBS	denotes	least
T4755	16429-16434	CD	denotes	eight
T4758	16435-16443	VBG	denotes	matching
T4757	16444-16449	NNS	denotes	cDNAs
T4759	16450-16452	IN	denotes	in
T4760	16453-16456	DT	denotes	the
T4761	16457-16460	NN	denotes	set
T4762	16461-16463	IN	denotes	of
T4763	16464-16469	CD	denotes	1,176
T4764	16470-16475	NNS	denotes	cDNAs
T4765	16476-16478	PRP	denotes	we
T4766	16479-16487	VBD	denotes	assigned
T4767	16488-16490	IN	denotes	to
T4768	16491-16497	JJ	denotes	single
T4770	16498-16507	JJ	denotes	olfactory
T4771	16508-16516	NN	denotes	receptor
T4769	16517-16526	NNS	denotes	sequences
T4772	16526-16527	.	denotes	.
T4773	16527-16706	sentence	denotes	However, for 17 olfactory receptors, we found ten or more matching cDNAs, suggesting that they might be expressed at higher levels than other olfactory receptor genes (Figure 2).
T4774	16528-16535	RB	denotes	However
T4776	16535-16537	,	denotes	,
T4777	16537-16540	IN	denotes	for
T4778	16541-16543	CD	denotes	17
T4780	16544-16553	JJ	denotes	olfactory
T4779	16554-16563	NNS	denotes	receptors
T4781	16563-16565	,	denotes	,
T4782	16565-16567	PRP	denotes	we
T4775	16568-16573	VBD	denotes	found
T4783	16574-16577	CD	denotes	ten
T4785	16578-16580	CC	denotes	or
T4786	16581-16585	JJR	denotes	more
T4787	16586-16594	NN	denotes	matching
T4784	16595-16600	NNS	denotes	cDNAs
T4788	16600-16602	,	denotes	,
T4789	16602-16612	VBG	denotes	suggesting
T4790	16613-16617	IN	denotes	that
T4792	16618-16622	PRP	denotes	they
T4793	16623-16628	MD	denotes	might
T4794	16629-16631	VB	denotes	be
T4791	16632-16641	VBN	denotes	expressed
T4795	16642-16644	IN	denotes	at
T4796	16645-16651	JJR	denotes	higher
T4797	16652-16658	NNS	denotes	levels
T4798	16659-16663	IN	denotes	than
T4799	16664-16669	JJ	denotes	other
T4801	16670-16679	JJ	denotes	olfactory
T4802	16680-16688	NN	denotes	receptor
T4800	16689-16694	NNS	denotes	genes
T4803	16695-16696	-LRB-	denotes	(
T4804	16696-16702	NN	denotes	Figure
T4805	16703-16704	CD	denotes	2
T4806	16704-16705	-RRB-	denotes	)
T4807	16705-16706	.	denotes	.
T4808	16706-16909	sentence	denotes	The two genes for which we found most cDNAs (AY318726/MOR28 and AY318727/MOR10) are genomically adjacent and in the well-studied olfactory receptor cluster next to the T-cell receptor α/δ locus [18,31].
T4809	16707-16710	DT	denotes	The
T4811	16711-16714	CD	denotes	two
T4810	16715-16720	NNS	denotes	genes
T4813	16721-16724	IN	denotes	for
T4815	16725-16730	WDT	denotes	which
T4816	16731-16733	PRP	denotes	we
T4814	16734-16739	VBD	denotes	found
T4817	16740-16744	JJS	denotes	most
T4818	16745-16750	NNS	denotes	cDNAs
T4819	16751-16752	-LRB-	denotes	(
T4820	16752-16760	NN	denotes	AY318726
T4822	16760-16761	HYPH	denotes	/
T4821	16761-16766	NN	denotes	MOR28
T4823	16767-16770	CC	denotes	and
T4824	16771-16779	NN	denotes	AY318727
T4826	16779-16780	HYPH	denotes	/
T4825	16780-16785	NN	denotes	MOR10
T4827	16785-16786	-RRB-	denotes	)
T4812	16787-16790	VBP	denotes	are
T4828	16791-16802	RB	denotes	genomically
T4829	16803-16811	JJ	denotes	adjacent
T4830	16812-16815	CC	denotes	and
T4831	16816-16818	IN	denotes	in
T4832	16819-16822	DT	denotes	the
T4834	16823-16827	RB	denotes	well
T4836	16827-16828	HYPH	denotes	-
T4835	16828-16835	VBN	denotes	studied
T4837	16836-16845	JJ	denotes	olfactory
T4838	16846-16854	NN	denotes	receptor
T4833	16855-16862	NN	denotes	cluster
T4839	16863-16867	JJ	denotes	next
T4840	16868-16870	IN	denotes	to
T4841	16871-16874	DT	denotes	the
T4843	16875-16876	NN	denotes	T
T4845	16876-16877	HYPH	denotes	-
T4844	16877-16881	NN	denotes	cell
T4846	16882-16890	NN	denotes	receptor
T4847	16891-16892	NN	denotes	α
T4849	16892-16893	HYPH	denotes	/
T4848	16893-16894	NN	denotes	δ
T4842	16895-16900	NN	denotes	locus
T4850	16901-16902	-LRB-	denotes	[
T4852	16902-16904	CD	denotes	18
T4853	16904-16905	,	denotes	,
T4851	16905-16907	CD	denotes	31
T4854	16907-16908	-RRB-	denotes	]
T4855	16908-16909	.	denotes	.
T4856	16909-17031	sentence	denotes	Quantitative RT-PCR of six olfactory receptors confirms that expression levels do indeed vary considerably between genes.
T4857	16910-16922	JJ	denotes	Quantitative
T4859	16923-16925	NN	denotes	RT
T4860	16925-16926	HYPH	denotes	-
T4858	16926-16929	NN	denotes	PCR
T4862	16930-16932	IN	denotes	of
T4863	16933-16936	CD	denotes	six
T4865	16937-16946	JJ	denotes	olfactory
T4864	16947-16956	NNS	denotes	receptors
T4861	16957-16965	VBZ	denotes	confirms
T4866	16966-16970	IN	denotes	that
T4868	16971-16981	NN	denotes	expression
T4869	16982-16988	NNS	denotes	levels
T4870	16989-16991	VBP	denotes	do
T4871	16992-16998	RB	denotes	indeed
T4867	16999-17003	VB	denotes	vary
T4872	17004-17016	RB	denotes	considerably
T4873	17017-17024	IN	denotes	between
T4874	17025-17030	NNS	denotes	genes
T4875	17030-17031	.	denotes	.
T4876	17031-17229	sentence	denotes	We used quantitative (real-time) PCR to measure olfactory epithelium transcript levels of six olfactory receptor genes and the ribosomal S16 gene in three mice of the same inbred strain (Figure 3).
T4877	17032-17034	PRP	denotes	We
T4878	17035-17039	VBD	denotes	used
T4879	17040-17052	JJ	denotes	quantitative
T4881	17053-17054	-LRB-	denotes	(
T4882	17054-17058	JJ	denotes	real
T4884	17058-17059	HYPH	denotes	-
T4883	17059-17063	NN	denotes	time
T4885	17063-17064	-RRB-	denotes	)
T4880	17065-17068	NN	denotes	PCR
T4886	17069-17071	TO	denotes	to
T4887	17072-17079	VB	denotes	measure
T4888	17080-17089	JJ	denotes	olfactory
T4889	17090-17100	NN	denotes	epithelium
T4891	17101-17111	NN	denotes	transcript
T4890	17112-17118	NNS	denotes	levels
T4892	17119-17121	IN	denotes	of
T4893	17122-17125	CD	denotes	six
T4895	17126-17135	JJ	denotes	olfactory
T4896	17136-17144	NN	denotes	receptor
T4894	17145-17150	NNS	denotes	genes
T4897	17151-17154	CC	denotes	and
T4898	17155-17158	DT	denotes	the
T4900	17159-17168	JJ	denotes	ribosomal
T4901	17169-17172	NN	denotes	S16
T4899	17173-17177	NN	denotes	gene
T4902	17178-17180	IN	denotes	in
T4903	17181-17186	CD	denotes	three
T4904	17187-17191	NNS	denotes	mice
T4905	17192-17194	IN	denotes	of
T4906	17195-17198	DT	denotes	the
T4908	17199-17203	JJ	denotes	same
T4909	17204-17210	JJ	denotes	inbred
T4907	17211-17217	NN	denotes	strain
T4910	17218-17219	-LRB-	denotes	(
T4911	17219-17225	NN	denotes	Figure
T4912	17226-17227	CD	denotes	3
T4913	17227-17228	-RRB-	denotes	)
T4914	17228-17229	.	denotes	.
T4915	17229-17402	sentence	denotes	These genes include two olfactory receptors with more than 20 matching cDNAs, two with one or two matching cDNAs and two class I olfactory receptors with no matching cDNAs.
T4916	17230-17235	DT	denotes	These
T4917	17236-17241	NNS	denotes	genes
T4918	17242-17249	VBP	denotes	include
T4919	17250-17253	CD	denotes	two
T4921	17254-17263	JJ	denotes	olfactory
T4920	17264-17273	NNS	denotes	receptors
T4922	17274-17278	IN	denotes	with
T4923	17279-17283	JJR	denotes	more
T4925	17284-17288	IN	denotes	than
T4924	17289-17291	CD	denotes	20
T4927	17292-17300	JJ	denotes	matching
T4926	17301-17306	NNS	denotes	cDNAs
T4928	17306-17308	,	denotes	,
T4929	17308-17311	CD	denotes	two
T4930	17312-17316	IN	denotes	with
T4931	17317-17320	CD	denotes	one
T4933	17321-17323	CC	denotes	or
T4934	17324-17327	CD	denotes	two
T4935	17328-17336	JJ	denotes	matching
T4932	17337-17342	NNS	denotes	cDNAs
T4936	17343-17346	CC	denotes	and
T4937	17347-17350	CD	denotes	two
T4939	17351-17356	NN	denotes	class
T4940	17357-17358	CD	denotes	I
T4941	17359-17368	JJ	denotes	olfactory
T4938	17369-17378	NNS	denotes	receptors
T4942	17379-17383	IN	denotes	with
T4943	17384-17386	DT	denotes	no
T4945	17387-17395	JJ	denotes	matching
T4944	17396-17401	NNS	denotes	cDNAs
T4946	17401-17402	.	denotes	.
T4947	17402-17629	sentence	denotes	In these assays, we measure transcript level per genomic copy of the gene by comparing how well a gene-specific primer pair amplifies reverse-transcribed RNA, relative to a standard curve of amplification of mouse genomic DNA.
T4948	17403-17405	IN	denotes	In
T4950	17406-17411	DT	denotes	these
T4951	17412-17418	NNS	denotes	assays
T4952	17418-17420	,	denotes	,
T4953	17420-17422	PRP	denotes	we
T4949	17423-17430	VBP	denotes	measure
T4954	17431-17441	NN	denotes	transcript
T4955	17442-17447	NN	denotes	level
T4956	17448-17451	IN	denotes	per
T4957	17452-17459	JJ	denotes	genomic
T4958	17460-17464	NN	denotes	copy
T4959	17465-17467	IN	denotes	of
T4960	17468-17471	DT	denotes	the
T4961	17472-17476	NN	denotes	gene
T4962	17477-17479	IN	denotes	by
T4963	17480-17489	VBG	denotes	comparing
T4964	17490-17493	WRB	denotes	how
T4965	17494-17498	RB	denotes	well
T4967	17499-17500	DT	denotes	a
T4969	17501-17505	NN	denotes	gene
T4971	17505-17506	HYPH	denotes	-
T4970	17506-17514	JJ	denotes	specific
T4972	17515-17521	NN	denotes	primer
T4968	17522-17526	NN	denotes	pair
T4966	17527-17536	VBZ	denotes	amplifies
T4973	17537-17544	JJ	denotes	reverse
T4975	17544-17545	HYPH	denotes	-
T4974	17545-17556	VBN	denotes	transcribed
T4976	17557-17560	NN	denotes	RNA
T4977	17560-17562	,	denotes	,
T4978	17562-17570	JJ	denotes	relative
T4979	17571-17573	IN	denotes	to
T4980	17574-17575	DT	denotes	a
T4982	17576-17584	JJ	denotes	standard
T4981	17585-17590	NN	denotes	curve
T4983	17591-17593	IN	denotes	of
T4984	17594-17607	NN	denotes	amplification
T4985	17608-17610	IN	denotes	of
T4986	17611-17616	NN	denotes	mouse
T4988	17617-17624	JJ	denotes	genomic
T4987	17625-17628	NN	denotes	DNA
T4989	17628-17629	.	denotes	.
T4990	17629-17742	sentence	denotes	We find that expression levels can vary by almost 300-fold between genes (for example, genes A and D, Figure 3).
T4991	17630-17632	PRP	denotes	We
T4992	17633-17637	VBP	denotes	find
T4993	17638-17642	IN	denotes	that
T4995	17643-17653	NN	denotes	expression
T4996	17654-17660	NNS	denotes	levels
T4997	17661-17664	MD	denotes	can
T4994	17665-17669	VB	denotes	vary
T4998	17670-17672	IN	denotes	by
T4999	17673-17679	RB	denotes	almost
T5000	17680-17688	JJ	denotes	300-fold
T5001	17689-17696	IN	denotes	between
T5002	17697-17702	NNS	denotes	genes
T5003	17703-17704	-LRB-	denotes	(
T5005	17704-17707	IN	denotes	for
T5006	17708-17715	NN	denotes	example
T5007	17715-17717	,	denotes	,
T5008	17717-17722	NNS	denotes	genes
T5009	17723-17724	NN	denotes	A
T5010	17725-17728	CC	denotes	and
T5011	17729-17730	NN	denotes	D
T5012	17730-17732	,	denotes	,
T5004	17732-17738	NN	denotes	Figure
T5013	17739-17740	CD	denotes	3
T5014	17740-17741	-RRB-	denotes	)
T5015	17741-17742	.	denotes	.
T5016	17742-18039	sentence	denotes	However, cDNA numbers are not a good indicator of expression level, a discrepancy that is likely to be due to bias in the screen (we used degenerate primers to make the probes, which will favor some genes over others) and in the libraries (oligo-dT priming will favor genes with shorter 3' UTRs).
T5017	17743-17750	RB	denotes	However
T5019	17750-17752	,	denotes	,
T5020	17752-17756	NN	denotes	cDNA
T5021	17757-17764	NNS	denotes	numbers
T5018	17765-17768	VBP	denotes	are
T5022	17769-17772	RB	denotes	not
T5023	17773-17774	DT	denotes	a
T5025	17775-17779	JJ	denotes	good
T5024	17780-17789	NN	denotes	indicator
T5026	17790-17792	IN	denotes	of
T5027	17793-17803	NN	denotes	expression
T5028	17804-17809	NN	denotes	level
T5029	17809-17811	,	denotes	,
T5030	17811-17812	DT	denotes	a
T5031	17813-17824	NN	denotes	discrepancy
T5032	17825-17829	WDT	denotes	that
T5033	17830-17832	VBZ	denotes	is
T5034	17833-17839	JJ	denotes	likely
T5035	17840-17842	TO	denotes	to
T5036	17843-17845	VB	denotes	be
T5037	17846-17849	IN	denotes	due
T5038	17850-17852	IN	denotes	to
T5039	17853-17857	NN	denotes	bias
T5040	17858-17860	IN	denotes	in
T5041	17861-17864	DT	denotes	the
T5042	17865-17871	NN	denotes	screen
T5043	17872-17873	-LRB-	denotes	(
T5045	17873-17875	PRP	denotes	we
T5044	17876-17880	VBD	denotes	used
T5046	17881-17891	JJ	denotes	degenerate
T5047	17892-17899	NNS	denotes	primers
T5048	17900-17902	TO	denotes	to
T5049	17903-17907	VB	denotes	make
T5050	17908-17911	DT	denotes	the
T5051	17912-17918	NNS	denotes	probes
T5052	17918-17920	,	denotes	,
T5053	17920-17925	WDT	denotes	which
T5055	17926-17930	MD	denotes	will
T5054	17931-17936	VB	denotes	favor
T5056	17937-17941	DT	denotes	some
T5057	17942-17947	NNS	denotes	genes
T5058	17948-17952	IN	denotes	over
T5059	17953-17959	NNS	denotes	others
T5060	17959-17960	-RRB-	denotes	)
T5061	17961-17964	CC	denotes	and
T5062	17965-17967	IN	denotes	in
T5063	17968-17971	DT	denotes	the
T5064	17972-17981	NNS	denotes	libraries
T5065	17982-17983	-LRB-	denotes	(
T5067	17983-17988	NN	denotes	oligo
T5069	17988-17989	HYPH	denotes	-
T5068	17989-17991	NN	denotes	dT
T5070	17992-17999	NN	denotes	priming
T5071	18000-18004	MD	denotes	will
T5066	18005-18010	VB	denotes	favor
T5072	18011-18016	NNS	denotes	genes
T5073	18017-18021	IN	denotes	with
T5074	18022-18029	JJR	denotes	shorter
T5076	18030-18031	CD	denotes	3
T5077	18031-18032	SYM	denotes	'
T5075	18033-18037	NNS	denotes	UTRs
T5078	18037-18038	-RRB-	denotes	)
T5079	18038-18039	.	denotes	.
T5080	18039-18340	sentence	denotes	For example, we observe large expression differences in all three mice between two genes for which similar numbers of cDNAs were found (genes A and B, Figure 3), and conversely, similar expression levels between two genes with a ten-fold difference in number of cDNAs found (genes B and C, Figure 3).
T5081	18040-18043	IN	denotes	For
T5083	18044-18051	NN	denotes	example
T5084	18051-18053	,	denotes	,
T5085	18053-18055	PRP	denotes	we
T5082	18056-18063	VBP	denotes	observe
T5086	18064-18069	JJ	denotes	large
T5088	18070-18080	NN	denotes	expression
T5087	18081-18092	NNS	denotes	differences
T5089	18093-18095	IN	denotes	in
T5090	18096-18099	PDT	denotes	all
T5092	18100-18105	CD	denotes	three
T5091	18106-18110	NNS	denotes	mice
T5093	18111-18118	IN	denotes	between
T5094	18119-18122	CD	denotes	two
T5095	18123-18128	NNS	denotes	genes
T5096	18129-18132	IN	denotes	for
T5098	18133-18138	WDT	denotes	which
T5099	18139-18146	JJ	denotes	similar
T5100	18147-18154	NNS	denotes	numbers
T5101	18155-18157	IN	denotes	of
T5102	18158-18163	NNS	denotes	cDNAs
T5103	18164-18168	VBD	denotes	were
T5097	18169-18174	VBN	denotes	found
T5104	18175-18176	-LRB-	denotes	(
T5105	18176-18181	NNS	denotes	genes
T5106	18182-18183	NN	denotes	A
T5107	18184-18187	CC	denotes	and
T5108	18188-18189	NN	denotes	B
T5109	18189-18191	,	denotes	,
T5110	18191-18197	NN	denotes	Figure
T5111	18198-18199	CD	denotes	3
T5112	18199-18200	-RRB-	denotes	)
T5113	18200-18202	,	denotes	,
T5114	18202-18205	CC	denotes	and
T5115	18206-18216	RB	denotes	conversely
T5117	18216-18218	,	denotes	,
T5118	18218-18225	JJ	denotes	similar
T5119	18226-18236	NN	denotes	expression
T5116	18237-18243	NNS	denotes	levels
T5120	18244-18251	IN	denotes	between
T5121	18252-18255	CD	denotes	two
T5122	18256-18261	NNS	denotes	genes
T5123	18262-18266	IN	denotes	with
T5124	18267-18268	DT	denotes	a
T5126	18269-18277	JJ	denotes	ten-fold
T5125	18278-18288	NN	denotes	difference
T5127	18289-18291	IN	denotes	in
T5128	18292-18298	NN	denotes	number
T5129	18299-18301	IN	denotes	of
T5130	18302-18307	NNS	denotes	cDNAs
T5131	18308-18313	VBN	denotes	found
T5132	18314-18315	-LRB-	denotes	(
T5133	18315-18320	NNS	denotes	genes
T5134	18321-18322	NN	denotes	B
T5135	18323-18326	CC	denotes	and
T5136	18327-18328	NN	denotes	C
T5137	18328-18330	,	denotes	,
T5138	18330-18336	NN	denotes	Figure
T5139	18337-18338	CD	denotes	3
T5140	18338-18339	-RRB-	denotes	)
T5141	18339-18340	.	denotes	.
T5142	18340-18693	sentence	denotes	Expression levels are mostly consistent between different mice: we find similar expression-level differences between olfactory receptor genes in all three mice examined (that is, the rank order of the six genes is similar among the three mice), although there is variation in expression level of some genes between mice (for example, gene E, Figure 3).
T5143	18341-18351	NN	denotes	Expression
T5144	18352-18358	NNS	denotes	levels
T5145	18359-18362	VBP	denotes	are
T5147	18363-18369	RB	denotes	mostly
T5148	18370-18380	JJ	denotes	consistent
T5149	18381-18388	IN	denotes	between
T5150	18389-18398	JJ	denotes	different
T5151	18399-18403	NNS	denotes	mice
T5152	18403-18405	:	denotes	:
T5153	18405-18407	PRP	denotes	we
T5146	18408-18412	VBP	denotes	find
T5154	18413-18420	JJ	denotes	similar
T5156	18421-18431	NN	denotes	expression
T5158	18431-18432	HYPH	denotes	-
T5157	18432-18437	NN	denotes	level
T5155	18438-18449	NNS	denotes	differences
T5159	18450-18457	IN	denotes	between
T5160	18458-18467	JJ	denotes	olfactory
T5161	18468-18476	NN	denotes	receptor
T5162	18477-18482	NNS	denotes	genes
T5163	18483-18485	IN	denotes	in
T5164	18486-18489	DT	denotes	all
T5166	18490-18495	CD	denotes	three
T5165	18496-18500	NNS	denotes	mice
T5167	18501-18509	VBN	denotes	examined
T5168	18510-18511	-LRB-	denotes	(
T5170	18511-18515	RB	denotes	that
T5171	18516-18518	RB	denotes	is
T5172	18518-18520	,	denotes	,
T5173	18520-18523	DT	denotes	the
T5175	18524-18528	NN	denotes	rank
T5174	18529-18534	NN	denotes	order
T5176	18535-18537	IN	denotes	of
T5177	18538-18541	DT	denotes	the
T5179	18542-18545	CD	denotes	six
T5178	18546-18551	NNS	denotes	genes
T5169	18552-18554	VBZ	denotes	is
T5180	18555-18562	JJ	denotes	similar
T5181	18563-18568	IN	denotes	among
T5182	18569-18572	DT	denotes	the
T5184	18573-18578	CD	denotes	three
T5183	18579-18583	NNS	denotes	mice
T5185	18583-18584	-RRB-	denotes	)
T5186	18584-18586	,	denotes	,
T5187	18586-18594	IN	denotes	although
T5189	18595-18600	EX	denotes	there
T5188	18601-18603	VBZ	denotes	is
T5190	18604-18613	NN	denotes	variation
T5191	18614-18616	IN	denotes	in
T5192	18617-18627	NN	denotes	expression
T5193	18628-18633	NN	denotes	level
T5194	18634-18636	IN	denotes	of
T5195	18637-18641	DT	denotes	some
T5196	18642-18647	NNS	denotes	genes
T5197	18648-18655	IN	denotes	between
T5198	18656-18660	NNS	denotes	mice
T5199	18661-18662	-LRB-	denotes	(
T5201	18662-18665	IN	denotes	for
T5202	18666-18673	NN	denotes	example
T5203	18673-18675	,	denotes	,
T5204	18675-18679	NN	denotes	gene
T5205	18680-18681	NN	denotes	E
T5206	18681-18683	,	denotes	,
T5200	18683-18689	NN	denotes	Figure
T5207	18690-18691	CD	denotes	3
T5208	18691-18692	-RRB-	denotes	)
T5209	18692-18693	.	denotes	.
T5210	18693-18923	sentence	denotes	In situ hybridization (Figure 4) shows that increased numbers of expressing cells account for some, but not all, of the difference in transcript levels between two of the genes tested by real-time PCR (genes A and D in Figure 3).
T5211	18694-18696	FW	denotes	In
T5212	18697-18701	FW	denotes	situ
T5213	18702-18715	NN	denotes	hybridization
T5215	18716-18717	-LRB-	denotes	(
T5216	18717-18723	NN	denotes	Figure
T5217	18724-18725	CD	denotes	4
T5218	18725-18726	-RRB-	denotes	)
T5214	18727-18732	VBZ	denotes	shows
T5219	18733-18737	IN	denotes	that
T5221	18738-18747	VBN	denotes	increased
T5222	18748-18755	NNS	denotes	numbers
T5223	18756-18758	IN	denotes	of
T5224	18759-18769	VBG	denotes	expressing
T5225	18770-18775	NNS	denotes	cells
T5220	18776-18783	VBP	denotes	account
T5226	18784-18787	IN	denotes	for
T5227	18788-18792	DT	denotes	some
T5228	18792-18794	,	denotes	,
T5229	18794-18797	CC	denotes	but
T5230	18798-18801	RB	denotes	not
T5231	18802-18805	DT	denotes	all
T5232	18805-18807	,	denotes	,
T5233	18807-18809	IN	denotes	of
T5234	18810-18813	DT	denotes	the
T5235	18814-18824	NN	denotes	difference
T5236	18825-18827	IN	denotes	in
T5237	18828-18838	NN	denotes	transcript
T5238	18839-18845	NNS	denotes	levels
T5239	18846-18853	IN	denotes	between
T5240	18854-18857	CD	denotes	two
T5241	18858-18860	IN	denotes	of
T5242	18861-18864	DT	denotes	the
T5243	18865-18870	NNS	denotes	genes
T5244	18871-18877	VBN	denotes	tested
T5245	18878-18880	IN	denotes	by
T5246	18881-18885	JJ	denotes	real
T5248	18885-18886	HYPH	denotes	-
T5247	18886-18890	NN	denotes	time
T5249	18891-18894	NN	denotes	PCR
T5250	18895-18896	-LRB-	denotes	(
T5251	18896-18901	NNS	denotes	genes
T5252	18902-18903	NN	denotes	A
T5253	18904-18907	CC	denotes	and
T5254	18908-18909	NN	denotes	D
T5255	18910-18912	IN	denotes	in
T5256	18913-18919	NN	denotes	Figure
T5257	18920-18921	CD	denotes	3
T5258	18921-18922	-RRB-	denotes	)
T5259	18922-18923	.	denotes	.
T5260	18923-19068	sentence	denotes	We hybridized alternate coronal serial sections spanning an entire olfactory epithelium of a young mouse (P6) with probes for gene A and gene D.
T5261	18924-18926	PRP	denotes	We
T5262	18927-18937	VBD	denotes	hybridized
T5263	18938-18947	JJ	denotes	alternate
T5265	18948-18955	JJ	denotes	coronal
T5266	18956-18962	JJ	denotes	serial
T5264	18963-18971	NNS	denotes	sections
T5267	18972-18980	VBG	denotes	spanning
T5268	18981-18983	DT	denotes	an
T5270	18984-18990	JJ	denotes	entire
T5271	18991-19000	JJ	denotes	olfactory
T5269	19001-19011	NN	denotes	epithelium
T5272	19012-19014	IN	denotes	of
T5273	19015-19016	DT	denotes	a
T5275	19017-19022	JJ	denotes	young
T5274	19023-19028	NN	denotes	mouse
T5276	19029-19030	-LRB-	denotes	(
T5277	19030-19032	NN	denotes	P6
T5278	19032-19033	-RRB-	denotes	)
T5279	19034-19038	IN	denotes	with
T5280	19039-19045	NNS	denotes	probes
T5281	19046-19049	IN	denotes	for
T5282	19050-19054	NN	denotes	gene
T5283	19055-19056	NN	denotes	A
T5284	19057-19060	CC	denotes	and
T5285	19061-19065	NN	denotes	gene
T5286	19066-19067	NN	denotes	D
T5287	19067-19068	.	denotes	.
T5288	19068-19205	sentence	denotes	Southern blot and BLAST analyses show that both probes are likely to hybridize to their intended target genes and no others (not shown).
T5289	19069-19077	NNP	denotes	Southern
T5290	19078-19082	NN	denotes	blot
T5292	19083-19086	CC	denotes	and
T5293	19087-19092	NN	denotes	BLAST
T5291	19093-19101	NNS	denotes	analyses
T5294	19102-19106	VBP	denotes	show
T5295	19107-19111	IN	denotes	that
T5297	19112-19116	DT	denotes	both
T5298	19117-19123	NNS	denotes	probes
T5296	19124-19127	VBP	denotes	are
T5299	19128-19134	JJ	denotes	likely
T5300	19135-19137	TO	denotes	to
T5301	19138-19147	VB	denotes	hybridize
T5302	19148-19150	IN	denotes	to
T5303	19151-19156	PRP$	denotes	their
T5305	19157-19165	VBN	denotes	intended
T5306	19166-19172	NN	denotes	target
T5304	19173-19178	NNS	denotes	genes
T5307	19179-19182	CC	denotes	and
T5308	19183-19185	DT	denotes	no
T5309	19186-19192	NNS	denotes	others
T5310	19193-19194	-LRB-	denotes	(
T5312	19194-19197	RB	denotes	not
T5311	19198-19203	VBN	denotes	shown
T5313	19203-19204	-RRB-	denotes	)
T5314	19204-19205	.	denotes	.
T5315	19205-19320	sentence	denotes	Gene A is expressed in zone 4 of the epithelium according to the nomenclature of Sullivan et al. [32] (Figure 4a).
T5316	19206-19210	NN	denotes	Gene
T5317	19211-19212	NN	denotes	A
T5319	19213-19215	VBZ	denotes	is
T5318	19216-19225	VBN	denotes	expressed
T5320	19226-19228	IN	denotes	in
T5321	19229-19233	NN	denotes	zone
T5322	19234-19235	CD	denotes	4
T5323	19236-19238	IN	denotes	of
T5324	19239-19242	DT	denotes	the
T5325	19243-19253	NN	denotes	epithelium
T5326	19254-19263	VBG	denotes	according
T5327	19264-19266	IN	denotes	to
T5328	19267-19270	DT	denotes	the
T5329	19271-19283	NN	denotes	nomenclature
T5330	19284-19286	IN	denotes	of
T5331	19287-19295	NNP	denotes	Sullivan
T5332	19296-19298	FW	denotes	et
T5333	19299-19302	FW	denotes	al.
T5334	19303-19304	-LRB-	denotes	[
T5335	19304-19306	CD	denotes	32
T5336	19306-19307	-RRB-	denotes	]
T5337	19308-19309	-LRB-	denotes	(
T5338	19309-19315	NN	denotes	Figure
T5339	19316-19318	CD	denotes	4a
T5340	19318-19319	-RRB-	denotes	)
T5341	19319-19320	.	denotes	.
T5342	19320-19652	sentence	denotes	The expression pattern of gene D does not correspond to any of the four 'classical' olfactory epithelial zones [14,15,32]: positive cells are found in regions of endoturbinates II and III and ectoturbinate 3, resembling the expression pattern seen previously for the OR37 subfamily and ORZ6 olfactory receptors [33,34] (Figure 4b).
T5343	19321-19324	DT	denotes	The
T5345	19325-19335	NN	denotes	expression
T5344	19336-19343	NN	denotes	pattern
T5347	19344-19346	IN	denotes	of
T5348	19347-19351	NN	denotes	gene
T5349	19352-19353	NN	denotes	D
T5350	19354-19358	VBZ	denotes	does
T5351	19359-19362	RB	denotes	not
T5346	19363-19373	VB	denotes	correspond
T5353	19374-19376	IN	denotes	to
T5354	19377-19380	DT	denotes	any
T5355	19381-19383	IN	denotes	of
T5356	19384-19387	DT	denotes	the
T5358	19388-19392	CD	denotes	four
T5359	19393-19394	``	denotes	'
T5360	19394-19403	JJ	denotes	classical
T5361	19403-19404	''	denotes	'
T5362	19405-19414	JJ	denotes	olfactory
T5363	19415-19425	JJ	denotes	epithelial
T5357	19426-19431	NNS	denotes	zones
T5364	19432-19433	-LRB-	denotes	[
T5366	19433-19435	CD	denotes	14
T5367	19435-19436	,	denotes	,
T5368	19436-19438	CD	denotes	15
T5369	19438-19439	,	denotes	,
T5365	19439-19441	CD	denotes	32
T5370	19441-19442	-RRB-	denotes	]
T5371	19442-19444	:	denotes	:
T5372	19444-19452	JJ	denotes	positive
T5373	19453-19458	NNS	denotes	cells
T5374	19459-19462	VBP	denotes	are
T5352	19463-19468	VBN	denotes	found
T5375	19469-19471	IN	denotes	in
T5376	19472-19479	NNS	denotes	regions
T5377	19480-19482	IN	denotes	of
T5378	19483-19497	NNS	denotes	endoturbinates
T5379	19498-19500	CD	denotes	II
T5380	19501-19504	CC	denotes	and
T5381	19505-19508	CD	denotes	III
T5382	19509-19512	CC	denotes	and
T5383	19513-19526	NN	denotes	ectoturbinate
T5384	19527-19528	CD	denotes	3
T5385	19528-19530	,	denotes	,
T5386	19530-19540	VBG	denotes	resembling
T5387	19541-19544	DT	denotes	the
T5389	19545-19555	NN	denotes	expression
T5388	19556-19563	NN	denotes	pattern
T5390	19564-19568	VBN	denotes	seen
T5391	19569-19579	RB	denotes	previously
T5392	19580-19583	IN	denotes	for
T5393	19584-19587	DT	denotes	the
T5395	19588-19592	NN	denotes	OR37
T5394	19593-19602	NN	denotes	subfamily
T5396	19603-19606	CC	denotes	and
T5397	19607-19611	NN	denotes	ORZ6
T5399	19612-19621	JJ	denotes	olfactory
T5398	19622-19631	NNS	denotes	receptors
T5400	19632-19633	-LRB-	denotes	[
T5402	19633-19635	CD	denotes	33
T5403	19635-19636	,	denotes	,
T5401	19636-19638	CD	denotes	34
T5404	19638-19639	-RRB-	denotes	]
T5405	19640-19641	-LRB-	denotes	(
T5406	19641-19647	NN	denotes	Figure
T5407	19648-19650	CD	denotes	4b
T5408	19650-19651	-RRB-	denotes	)
T5409	19651-19652	.	denotes	.
T5410	19652-19880	sentence	denotes	Counting the total number of positive cells in alternate sections across the entire epithelium, we find that gene A is expressed in 2,905 cells, about 12 times more cells than gene D, which is expressed in a total of 249 cells.
T5411	19653-19661	VBG	denotes	Counting
T5413	19662-19665	DT	denotes	the
T5415	19666-19671	JJ	denotes	total
T5414	19672-19678	NN	denotes	number
T5416	19679-19681	IN	denotes	of
T5417	19682-19690	JJ	denotes	positive
T5418	19691-19696	NNS	denotes	cells
T5419	19697-19699	IN	denotes	in
T5420	19700-19709	JJ	denotes	alternate
T5421	19710-19718	NNS	denotes	sections
T5422	19719-19725	IN	denotes	across
T5423	19726-19729	DT	denotes	the
T5425	19730-19736	JJ	denotes	entire
T5424	19737-19747	NN	denotes	epithelium
T5426	19747-19749	,	denotes	,
T5427	19749-19751	PRP	denotes	we
T5412	19752-19756	VBP	denotes	find
T5428	19757-19761	IN	denotes	that
T5430	19762-19766	NN	denotes	gene
T5431	19767-19768	NN	denotes	A
T5432	19769-19771	VBZ	denotes	is
T5429	19772-19781	VBN	denotes	expressed
T5433	19782-19784	IN	denotes	in
T5434	19785-19790	CD	denotes	2,905
T5435	19791-19796	NNS	denotes	cells
T5436	19796-19798	,	denotes	,
T5437	19798-19803	RB	denotes	about
T5438	19804-19806	CD	denotes	12
T5440	19807-19812	NNS	denotes	times
T5441	19813-19817	JJR	denotes	more
T5439	19818-19823	NNS	denotes	cells
T5442	19824-19828	IN	denotes	than
T5443	19829-19833	NN	denotes	gene
T5444	19834-19835	NN	denotes	D
T5445	19835-19837	,	denotes	,
T5446	19837-19842	WDT	denotes	which
T5448	19843-19845	VBZ	denotes	is
T5447	19846-19855	VBN	denotes	expressed
T5449	19856-19858	IN	denotes	in
T5450	19859-19860	DT	denotes	a
T5451	19861-19866	NN	denotes	total
T5452	19867-19869	IN	denotes	of
T5453	19870-19873	CD	denotes	249
T5454	19874-19879	NNS	denotes	cells
T5455	19879-19880	.	denotes	.
T5456	19880-20177	sentence	denotes	This 12-fold difference in numbers of expressing cells does not account for the almost 300-fold difference in RNA levels observed by real-time PCR, implying that the transcript level per expressing cell for gene A is about 25 times higher than transcript level in each expressing cell for gene D.
T5457	19881-19885	DT	denotes	This
T5459	19886-19893	JJ	denotes	12-fold
T5458	19894-19904	NN	denotes	difference
T5461	19905-19907	IN	denotes	in
T5462	19908-19915	NNS	denotes	numbers
T5463	19916-19918	IN	denotes	of
T5464	19919-19929	VBG	denotes	expressing
T5465	19930-19935	NNS	denotes	cells
T5466	19936-19940	VBZ	denotes	does
T5467	19941-19944	RB	denotes	not
T5460	19945-19952	VB	denotes	account
T5468	19953-19956	IN	denotes	for
T5469	19957-19960	DT	denotes	the
T5471	19961-19967	RB	denotes	almost
T5472	19968-19976	JJ	denotes	300-fold
T5470	19977-19987	NN	denotes	difference
T5473	19988-19990	IN	denotes	in
T5474	19991-19994	NN	denotes	RNA
T5475	19995-20001	NNS	denotes	levels
T5476	20002-20010	VBN	denotes	observed
T5477	20011-20013	IN	denotes	by
T5478	20014-20018	JJ	denotes	real
T5480	20018-20019	HYPH	denotes	-
T5479	20019-20023	NN	denotes	time
T5481	20024-20027	NN	denotes	PCR
T5482	20027-20029	,	denotes	,
T5483	20029-20037	VBG	denotes	implying
T5484	20038-20042	IN	denotes	that
T5486	20043-20046	DT	denotes	the
T5488	20047-20057	NN	denotes	transcript
T5487	20058-20063	NN	denotes	level
T5489	20064-20067	IN	denotes	per
T5490	20068-20078	VBG	denotes	expressing
T5491	20079-20083	NN	denotes	cell
T5492	20084-20087	IN	denotes	for
T5493	20088-20092	NN	denotes	gene
T5494	20093-20094	NN	denotes	A
T5485	20095-20097	VBZ	denotes	is
T5495	20098-20103	IN	denotes	about
T5496	20104-20106	CD	denotes	25
T5498	20107-20112	NNS	denotes	times
T5497	20113-20119	JJR	denotes	higher
T5499	20120-20124	IN	denotes	than
T5500	20125-20135	NN	denotes	transcript
T5501	20136-20141	NN	denotes	level
T5502	20142-20144	IN	denotes	in
T5503	20145-20149	DT	denotes	each
T5505	20150-20160	VBG	denotes	expressing
T5504	20161-20165	NN	denotes	cell
T5506	20166-20169	IN	denotes	for
T5507	20170-20174	NN	denotes	gene
T5508	20175-20176	NN	denotes	D
T5509	20176-20177	.	denotes	.
T5510	20177-20380	sentence	denotes	We note that hybridization intensities per positive neuron appear stronger for gene A than gene D after comparable exposure times, in accordance with the idea that transcript levels are higher per cell.
T5511	20178-20180	PRP	denotes	We
T5512	20181-20185	VBP	denotes	note
T5513	20186-20190	IN	denotes	that
T5515	20191-20204	NN	denotes	hybridization
T5516	20205-20216	NNS	denotes	intensities
T5517	20217-20220	IN	denotes	per
T5518	20221-20229	JJ	denotes	positive
T5519	20230-20236	NN	denotes	neuron
T5514	20237-20243	VBP	denotes	appear
T5520	20244-20252	JJR	denotes	stronger
T5521	20253-20256	IN	denotes	for
T5522	20257-20261	NN	denotes	gene
T5523	20262-20263	NN	denotes	A
T5524	20264-20268	IN	denotes	than
T5525	20269-20273	NN	denotes	gene
T5526	20274-20275	NN	denotes	D
T5527	20276-20281	IN	denotes	after
T5528	20282-20292	JJ	denotes	comparable
T5530	20293-20301	NN	denotes	exposure
T5529	20302-20307	NNS	denotes	times
T5531	20307-20309	,	denotes	,
T5532	20309-20311	IN	denotes	in
T5533	20312-20322	NN	denotes	accordance
T5534	20323-20327	IN	denotes	with
T5535	20328-20331	DT	denotes	the
T5536	20332-20336	NN	denotes	idea
T5537	20337-20341	IN	denotes	that
T5539	20342-20352	NN	denotes	transcript
T5540	20353-20359	NNS	denotes	levels
T5538	20360-20363	VBP	denotes	are
T5541	20364-20370	JJR	denotes	higher
T5542	20371-20374	IN	denotes	per
T5543	20375-20379	NN	denotes	cell
T5544	20379-20380	.	denotes	.
T5545	20380-20578	sentence	denotes	Thus, we suggest that expression in more cells and in higher levels per cell together account for the almost 300-fold higher olfactory epithelial RNA levels of gene A relative to gene D (Figure 3).
T5546	20381-20385	RB	denotes	Thus
T5548	20385-20387	,	denotes	,
T5549	20387-20389	PRP	denotes	we
T5547	20390-20397	VBP	denotes	suggest
T5550	20398-20402	IN	denotes	that
T5552	20403-20413	NN	denotes	expression
T5553	20414-20416	IN	denotes	in
T5554	20417-20421	JJR	denotes	more
T5555	20422-20427	NNS	denotes	cells
T5556	20428-20431	CC	denotes	and
T5557	20432-20434	IN	denotes	in
T5558	20435-20441	JJR	denotes	higher
T5559	20442-20448	NNS	denotes	levels
T5560	20449-20452	IN	denotes	per
T5561	20453-20457	NN	denotes	cell
T5562	20458-20466	RB	denotes	together
T5551	20467-20474	VBP	denotes	account
T5563	20475-20478	IN	denotes	for
T5564	20479-20482	DT	denotes	the
T5566	20483-20489	RB	denotes	almost
T5567	20490-20498	JJ	denotes	300-fold
T5568	20499-20505	JJR	denotes	higher
T5569	20506-20515	JJ	denotes	olfactory
T5571	20516-20526	JJ	denotes	epithelial
T5570	20527-20530	NN	denotes	RNA
T5565	20531-20537	NNS	denotes	levels
T5572	20538-20540	IN	denotes	of
T5573	20541-20545	NN	denotes	gene
T5574	20546-20547	NN	denotes	A
T5575	20548-20556	JJ	denotes	relative
T5576	20557-20559	IN	denotes	to
T5577	20560-20564	NN	denotes	gene
T5578	20565-20566	NN	denotes	D
T5579	20567-20568	-LRB-	denotes	(
T5580	20568-20574	NN	denotes	Figure
T5581	20575-20576	CD	denotes	3
T5582	20576-20577	-RRB-	denotes	)
T5583	20577-20578	.	denotes	.
T5714	20580-20584	RBS	denotes	Most
T5716	20585-20594	JJ	denotes	olfactory
T5717	20595-20603	NN	denotes	receptor
T5715	20604-20609	NNS	denotes	genes
T5718	20610-20614	VBP	denotes	have
T5719	20615-20622	JJ	denotes	several
T5721	20623-20638	JJ	denotes	transcriptional
T5720	20639-20647	NNS	denotes	isoforms
T5722	20647-20793	sentence	denotes	Our cDNA collection reveals that at least two thirds of the olfactory receptors sampled show alternative splicing of their 5' untranslated exons.
T5723	20648-20651	PRP$	denotes	Our
T5725	20652-20656	NN	denotes	cDNA
T5724	20657-20667	NN	denotes	collection
T5726	20668-20675	VBZ	denotes	reveals
T5727	20676-20680	IN	denotes	that
T5729	20681-20683	RB	denotes	at
T5731	20684-20689	RBS	denotes	least
T5730	20690-20693	CD	denotes	two
T5732	20694-20700	NNS	denotes	thirds
T5733	20701-20703	IN	denotes	of
T5734	20704-20707	DT	denotes	the
T5736	20708-20717	JJ	denotes	olfactory
T5735	20718-20727	NNS	denotes	receptors
T5737	20728-20735	VBN	denotes	sampled
T5728	20736-20740	VBP	denotes	show
T5738	20741-20752	JJ	denotes	alternative
T5739	20753-20761	NN	denotes	splicing
T5740	20762-20764	IN	denotes	of
T5741	20765-20770	PRP$	denotes	their
T5743	20771-20772	CD	denotes	5
T5744	20772-20773	SYM	denotes	'
T5745	20774-20786	JJ	denotes	untranslated
T5742	20787-20792	NNS	denotes	exons
T5746	20792-20793	.	denotes	.
T5747	20793-21226	sentence	denotes	Using a custom script to process sim4 alignments of cDNA and genomic sequences, we find two to eight different splice forms for 85 (45%) of the 191 genes for which we have had some opportunity to observe alternate splicing (that is, a minimum of two cDNAs, at least one of which is spliced), and 55 (67%) of the 82 genes for which we have four or more cDNAs (and thus a higher chance of observing any alternate splicing) (Figure 5).
T5748	20794-20799	VBG	denotes	Using
T5750	20800-20801	DT	denotes	a
T5752	20802-20808	NN	denotes	custom
T5751	20809-20815	NN	denotes	script
T5753	20816-20818	TO	denotes	to
T5754	20819-20826	VB	denotes	process
T5755	20827-20831	NN	denotes	sim4
T5756	20832-20842	NNS	denotes	alignments
T5757	20843-20845	IN	denotes	of
T5758	20846-20850	NN	denotes	cDNA
T5760	20851-20854	CC	denotes	and
T5761	20855-20862	JJ	denotes	genomic
T5759	20863-20872	NNS	denotes	sequences
T5762	20872-20874	,	denotes	,
T5763	20874-20876	PRP	denotes	we
T5749	20877-20881	VBP	denotes	find
T5764	20882-20885	CD	denotes	two
T5766	20886-20888	IN	denotes	to
T5765	20889-20894	CD	denotes	eight
T5768	20895-20904	JJ	denotes	different
T5769	20905-20911	NN	denotes	splice
T5767	20912-20917	NNS	denotes	forms
T5770	20918-20921	IN	denotes	for
T5771	20922-20924	CD	denotes	85
T5772	20925-20926	-LRB-	denotes	(
T5773	20926-20928	CD	denotes	45
T5774	20928-20929	NN	denotes	%
T5775	20929-20930	-RRB-	denotes	)
T5776	20931-20933	IN	denotes	of
T5777	20934-20937	DT	denotes	the
T5779	20938-20941	CD	denotes	191
T5778	20942-20947	NNS	denotes	genes
T5780	20948-20951	IN	denotes	for
T5782	20952-20957	WDT	denotes	which
T5783	20958-20960	PRP	denotes	we
T5784	20961-20965	VBP	denotes	have
T5781	20966-20969	VBN	denotes	had
T5785	20970-20974	DT	denotes	some
T5786	20975-20986	NN	denotes	opportunity
T5787	20987-20989	TO	denotes	to
T5788	20990-20997	VB	denotes	observe
T5789	20998-21007	JJ	denotes	alternate
T5790	21008-21016	NN	denotes	splicing
T5791	21017-21018	-LRB-	denotes	(
T5793	21018-21022	RB	denotes	that
T5794	21023-21025	RB	denotes	is
T5795	21025-21027	,	denotes	,
T5796	21027-21028	DT	denotes	a
T5792	21029-21036	NN	denotes	minimum
T5797	21037-21039	IN	denotes	of
T5798	21040-21043	CD	denotes	two
T5799	21044-21049	NNS	denotes	cDNAs
T5800	21049-21051	,	denotes	,
T5801	21051-21053	RB	denotes	at
T5802	21054-21059	RBS	denotes	least
T5803	21060-21063	CD	denotes	one
T5805	21064-21066	IN	denotes	of
T5806	21067-21072	WDT	denotes	which
T5807	21073-21075	VBZ	denotes	is
T5804	21076-21083	VBN	denotes	spliced
T5808	21083-21084	-RRB-	denotes	)
T5809	21084-21086	,	denotes	,
T5810	21086-21089	CC	denotes	and
T5811	21090-21092	CD	denotes	55
T5812	21093-21094	-LRB-	denotes	(
T5814	21094-21096	CD	denotes	67
T5813	21096-21097	NN	denotes	%
T5815	21097-21098	-RRB-	denotes	)
T5816	21099-21101	IN	denotes	of
T5817	21102-21105	DT	denotes	the
T5819	21106-21108	CD	denotes	82
T5818	21109-21114	NNS	denotes	genes
T5820	21115-21118	IN	denotes	for
T5822	21119-21124	WDT	denotes	which
T5823	21125-21127	PRP	denotes	we
T5821	21128-21132	VBP	denotes	have
T5824	21133-21137	CD	denotes	four
T5826	21138-21140	CC	denotes	or
T5827	21141-21145	JJR	denotes	more
T5825	21146-21151	NNS	denotes	cDNAs
T5828	21152-21153	-LRB-	denotes	(
T5829	21153-21156	CC	denotes	and
T5830	21157-21161	RB	denotes	thus
T5832	21162-21163	DT	denotes	a
T5833	21164-21170	JJR	denotes	higher
T5831	21171-21177	NN	denotes	chance
T5834	21178-21180	IN	denotes	of
T5835	21181-21190	VBG	denotes	observing
T5836	21191-21194	DT	denotes	any
T5838	21195-21204	JJ	denotes	alternate
T5837	21205-21213	NN	denotes	splicing
T5839	21213-21214	-RRB-	denotes	)
T5840	21215-21216	-LRB-	denotes	(
T5841	21216-21222	NN	denotes	Figure
T5842	21223-21224	CD	denotes	5
T5843	21224-21225	-RRB-	denotes	)
T5844	21225-21226	.	denotes	.
T5845	21226-21370	sentence	denotes	These alternative splice events are almost all restricted to the 5' UTR and include exon skipping and alternate splice-donor and -acceptor use.
T5846	21227-21232	DT	denotes	These
T5848	21233-21244	JJ	denotes	alternative
T5849	21245-21251	NN	denotes	splice
T5847	21252-21258	NNS	denotes	events
T5851	21259-21262	VBP	denotes	are
T5852	21263-21269	RB	denotes	almost
T5853	21270-21273	RB	denotes	all
T5850	21274-21284	VBN	denotes	restricted
T5854	21285-21287	IN	denotes	to
T5855	21288-21291	DT	denotes	the
T5857	21292-21293	CD	denotes	5
T5858	21293-21294	SYM	denotes	'
T5856	21295-21298	NN	denotes	UTR
T5859	21299-21302	CC	denotes	and
T5860	21303-21310	VBP	denotes	include
T5861	21311-21315	NN	denotes	exon
T5862	21316-21324	NN	denotes	skipping
T5863	21325-21328	CC	denotes	and
T5864	21329-21338	JJ	denotes	alternate
T5865	21339-21345	NN	denotes	splice
T5867	21345-21346	HYPH	denotes	-
T5868	21346-21351	NN	denotes	donor
T5869	21352-21355	CC	denotes	and
T5870	21356-21357	HYPH	denotes	-
T5871	21357-21365	NN	denotes	acceptor
T5866	21366-21369	NN	denotes	use
T5872	21369-21370	.	denotes	.
T5873	21370-21532	sentence	denotes	At least half of the olfactory receptors represented in our cDNA collection utilize more than one polyadenylation site, resulting in alternative 3' UTR isoforms.
T5874	21371-21373	RB	denotes	At
T5875	21374-21379	RBS	denotes	least
T5876	21380-21384	NN	denotes	half
T5878	21385-21387	IN	denotes	of
T5879	21388-21391	DT	denotes	the
T5881	21392-21401	JJ	denotes	olfactory
T5880	21402-21411	NNS	denotes	receptors
T5882	21412-21423	VBN	denotes	represented
T5883	21424-21426	IN	denotes	in
T5884	21427-21430	PRP$	denotes	our
T5886	21431-21435	NN	denotes	cDNA
T5885	21436-21446	NN	denotes	collection
T5877	21447-21454	VBP	denotes	utilize
T5887	21455-21459	JJR	denotes	more
T5889	21460-21464	IN	denotes	than
T5888	21465-21468	CD	denotes	one
T5891	21469-21484	NN	denotes	polyadenylation
T5890	21485-21489	NN	denotes	site
T5892	21489-21491	,	denotes	,
T5893	21491-21500	VBG	denotes	resulting
T5894	21501-21503	IN	denotes	in
T5895	21504-21515	JJ	denotes	alternative
T5897	21516-21517	CD	denotes	3
T5899	21517-21518	SYM	denotes	'
T5898	21519-21522	NN	denotes	UTR
T5896	21523-21531	NNS	denotes	isoforms
T5900	21531-21532	.	denotes	.
T5901	21532-21664	sentence	denotes	We have crudely estimated 3' UTR size for 1,169 cDNA clones by combining approximate insert size information with 5' sequence data.
T5902	21533-21535	PRP	denotes	We
T5904	21536-21540	VBP	denotes	have
T5905	21541-21548	RB	denotes	crudely
T5903	21549-21558	VBN	denotes	estimated
T5906	21559-21560	CD	denotes	3
T5908	21560-21561	SYM	denotes	'
T5907	21562-21565	NN	denotes	UTR
T5909	21566-21570	NN	denotes	size
T5910	21571-21574	IN	denotes	for
T5911	21575-21580	CD	denotes	1,169
T5913	21581-21585	NN	denotes	cDNA
T5912	21586-21592	NNS	denotes	clones
T5914	21593-21595	IN	denotes	by
T5915	21596-21605	VBG	denotes	combining
T5916	21606-21617	JJ	denotes	approximate
T5918	21618-21624	NN	denotes	insert
T5917	21625-21629	NN	denotes	size
T5919	21630-21641	NN	denotes	information
T5920	21642-21646	IN	denotes	with
T5921	21647-21648	CD	denotes	5
T5923	21648-21649	SYM	denotes	'
T5922	21650-21658	NN	denotes	sequence
T5924	21659-21663	NNS	denotes	data
T5925	21663-21664	.	denotes	.
T5926	21664-21805	sentence	denotes	More than one 3' UTR isoform is predicted for 43 of the 77 (56%) genes for which there are at least four cDNAs with 3' UTR size information.
T5927	21665-21669	JJR	denotes	More
T5929	21670-21674	IN	denotes	than
T5928	21675-21678	CD	denotes	one
T5931	21679-21680	CD	denotes	3
T5933	21680-21681	SYM	denotes	'
T5932	21682-21685	NN	denotes	UTR
T5930	21686-21693	NN	denotes	isoform
T5935	21694-21696	VBZ	denotes	is
T5934	21697-21706	VBN	denotes	predicted
T5936	21707-21710	IN	denotes	for
T5937	21711-21713	CD	denotes	43
T5939	21714-21716	IN	denotes	of
T5940	21717-21720	DT	denotes	the
T5938	21721-21723	CD	denotes	77
T5942	21724-21725	-LRB-	denotes	(
T5944	21725-21727	CD	denotes	56
T5943	21727-21728	NN	denotes	%
T5945	21728-21729	-RRB-	denotes	)
T5941	21730-21735	NNS	denotes	genes
T5946	21736-21739	IN	denotes	for
T5948	21740-21745	WDT	denotes	which
T5949	21746-21751	EX	denotes	there
T5947	21752-21755	VBP	denotes	are
T5950	21756-21758	RB	denotes	at
T5952	21759-21764	RBS	denotes	least
T5951	21765-21769	CD	denotes	four
T5953	21770-21775	NNS	denotes	cDNAs
T5954	21776-21780	IN	denotes	with
T5955	21781-21782	CD	denotes	3
T5957	21782-21783	SYM	denotes	'
T5956	21784-21787	NN	denotes	UTR
T5958	21788-21792	NN	denotes	size
T5959	21793-21804	NN	denotes	information
T5960	21804-21805	.	denotes	.
T5961	21805-21938	sentence	denotes	We confirmed the alternative polyadenylation isoforms of four out of five selected genes by sequencing the 3' end of 14 cDNA clones.
T5962	21806-21808	PRP	denotes	We
T5963	21809-21818	VBD	denotes	confirmed
T5964	21819-21822	DT	denotes	the
T5966	21823-21834	JJ	denotes	alternative
T5967	21835-21850	NN	denotes	polyadenylation
T5965	21851-21859	NNS	denotes	isoforms
T5968	21860-21862	IN	denotes	of
T5969	21863-21867	CD	denotes	four
T5971	21868-21871	IN	denotes	out
T5972	21872-21874	IN	denotes	of
T5970	21875-21879	CD	denotes	five
T5974	21880-21888	VBN	denotes	selected
T5973	21889-21894	NNS	denotes	genes
T5975	21895-21897	IN	denotes	by
T5976	21898-21908	VBG	denotes	sequencing
T5977	21909-21912	DT	denotes	the
T5979	21913-21914	CD	denotes	3
T5980	21914-21915	SYM	denotes	'
T5978	21916-21919	NN	denotes	end
T5981	21920-21922	IN	denotes	of
T5982	21923-21925	CD	denotes	14
T5984	21926-21930	NN	denotes	cDNA
T5983	21931-21937	NNS	denotes	clones
T5985	21937-21938	.	denotes	.
T5986	21938-22179	sentence	denotes	These 14 sequences also revealed one cDNA where the poly(A) tail was added 27 bp before the stop codon, and another where an intron was spliced out of the 3' UTR, contrary to the conventional stereotype of olfactory receptor gene structure.
T5987	21939-21944	DT	denotes	These
T5989	21945-21947	CD	denotes	14
T5988	21948-21957	NNS	denotes	sequences
T5991	21958-21962	RB	denotes	also
T5990	21963-21971	VBD	denotes	revealed
T5992	21972-21975	CD	denotes	one
T5993	21976-21980	NN	denotes	cDNA
T5994	21981-21986	WRB	denotes	where
T5996	21987-21990	DT	denotes	the
T5998	21991-21995	NN	denotes	poly
T5999	21995-21996	-LRB-	denotes	(
T6000	21996-21997	NN	denotes	A
T6001	21997-21998	-RRB-	denotes	)
T5997	21999-22003	NN	denotes	tail
T6002	22004-22007	VBD	denotes	was
T5995	22008-22013	VBN	denotes	added
T6003	22014-22016	CD	denotes	27
T6004	22017-22019	NN	denotes	bp
T6005	22020-22026	IN	denotes	before
T6006	22027-22030	DT	denotes	the
T6008	22031-22035	NN	denotes	stop
T6007	22036-22041	NN	denotes	codon
T6009	22041-22043	,	denotes	,
T6010	22043-22046	CC	denotes	and
T6011	22047-22054	DT	denotes	another
T6012	22055-22060	WRB	denotes	where
T6014	22061-22063	DT	denotes	an
T6015	22064-22070	NN	denotes	intron
T6016	22071-22074	VBD	denotes	was
T6013	22075-22082	VBN	denotes	spliced
T6017	22083-22086	IN	denotes	out
T6018	22087-22089	IN	denotes	of
T6019	22090-22093	DT	denotes	the
T6021	22094-22095	CD	denotes	3
T6022	22095-22096	SYM	denotes	'
T6020	22097-22100	NN	denotes	UTR
T6023	22100-22102	,	denotes	,
T6024	22102-22110	JJ	denotes	contrary
T6025	22111-22113	IN	denotes	to
T6026	22114-22117	DT	denotes	the
T6028	22118-22130	JJ	denotes	conventional
T6027	22131-22141	NN	denotes	stereotype
T6029	22142-22144	IN	denotes	of
T6030	22145-22154	JJ	denotes	olfactory
T6031	22155-22163	NN	denotes	receptor
T6032	22164-22168	NN	denotes	gene
T6033	22169-22178	NN	denotes	structure
T6034	22178-22179	.	denotes	.
T6417	22181-22182	DT	denotes	A
T6418	22183-22189	NN	denotes	subset
T6420	22190-22192	IN	denotes	of
T6421	22193-22202	JJ	denotes	olfactory
T6422	22203-22212	NNS	denotes	receptors
T6419	22213-22218	VBZ	denotes	shows
T6423	22219-22226	JJ	denotes	unusual
T6424	22227-22235	NN	denotes	splicing
T6425	22235-22431	sentence	denotes	We identified 62 cDNAs (5% of all olfactory receptor clones) from 38 intact olfactory receptors and one olfactory receptor pseudogene where a splice site within the protein-coding region is used.
T6426	22236-22238	PRP	denotes	We
T6427	22239-22249	VBD	denotes	identified
T6428	22250-22252	CD	denotes	62
T6429	22253-22258	NNS	denotes	cDNAs
T6430	22259-22260	-LRB-	denotes	(
T6432	22260-22261	CD	denotes	5
T6431	22261-22262	NN	denotes	%
T6433	22263-22265	IN	denotes	of
T6434	22266-22269	DT	denotes	all
T6436	22270-22279	JJ	denotes	olfactory
T6437	22280-22288	NN	denotes	receptor
T6435	22289-22295	NNS	denotes	clones
T6438	22295-22296	-RRB-	denotes	)
T6439	22297-22301	IN	denotes	from
T6440	22302-22304	CD	denotes	38
T6442	22305-22311	JJ	denotes	intact
T6443	22312-22321	JJ	denotes	olfactory
T6441	22322-22331	NNS	denotes	receptors
T6444	22332-22335	CC	denotes	and
T6445	22336-22339	CD	denotes	one
T6447	22340-22349	JJ	denotes	olfactory
T6448	22350-22358	NN	denotes	receptor
T6446	22359-22369	NN	denotes	pseudogene
T6449	22370-22375	WRB	denotes	where
T6451	22376-22377	DT	denotes	a
T6453	22378-22384	NN	denotes	splice
T6452	22385-22389	NN	denotes	site
T6454	22390-22396	IN	denotes	within
T6455	22397-22400	DT	denotes	the
T6457	22401-22408	NN	denotes	protein
T6459	22408-22409	HYPH	denotes	-
T6458	22409-22415	VBG	denotes	coding
T6456	22416-22422	NN	denotes	region
T6460	22423-22425	VBZ	denotes	is
T6450	22426-22430	VBN	denotes	used
T6461	22430-22431	.	denotes	.
T6462	22431-22638	sentence	denotes	For two genes (top two cDNAs, Figure 6), the predicted protein appears to be an intact olfactory receptor with three or ten amino acids, including the initiating methionine, contributed by an upstream exon.
T6463	22432-22435	IN	denotes	For
T6465	22436-22439	CD	denotes	two
T6466	22440-22445	NNS	denotes	genes
T6467	22446-22447	-LRB-	denotes	(
T6469	22447-22450	JJ	denotes	top
T6471	22451-22454	CD	denotes	two
T6470	22455-22460	NNS	denotes	cDNAs
T6472	22460-22462	,	denotes	,
T6468	22462-22468	NN	denotes	Figure
T6473	22469-22470	CD	denotes	6
T6474	22470-22471	-RRB-	denotes	)
T6475	22471-22473	,	denotes	,
T6476	22473-22476	DT	denotes	the
T6478	22477-22486	VBN	denotes	predicted
T6477	22487-22494	NN	denotes	protein
T6464	22495-22502	VBZ	denotes	appears
T6479	22503-22505	TO	denotes	to
T6480	22506-22508	VB	denotes	be
T6481	22509-22511	DT	denotes	an
T6483	22512-22518	JJ	denotes	intact
T6484	22519-22528	JJ	denotes	olfactory
T6482	22529-22537	NN	denotes	receptor
T6485	22538-22542	IN	denotes	with
T6486	22543-22548	CD	denotes	three
T6488	22549-22551	CC	denotes	or
T6489	22552-22555	CD	denotes	ten
T6490	22556-22561	NN	denotes	amino
T6487	22562-22567	NNS	denotes	acids
T6491	22567-22569	,	denotes	,
T6492	22569-22578	VBG	denotes	including
T6493	22579-22582	DT	denotes	the
T6495	22583-22593	VBG	denotes	initiating
T6494	22594-22604	NN	denotes	methionine
T6496	22604-22606	,	denotes	,
T6497	22606-22617	VBN	denotes	contributed
T6498	22618-22620	IN	denotes	by
T6499	22621-22623	DT	denotes	an
T6501	22624-22632	JJ	denotes	upstream
T6500	22633-22637	NN	denotes	exon
T6502	22637-22638	.	denotes	.
T6503	22638-22725	sentence	denotes	A similar gene structure was described previously for a human olfactory receptor [25].
T6504	22639-22640	DT	denotes	A
T6506	22641-22648	JJ	denotes	similar
T6507	22649-22653	NN	denotes	gene
T6505	22654-22663	NN	denotes	structure
T6509	22664-22667	VBD	denotes	was
T6508	22668-22677	VBN	denotes	described
T6510	22678-22688	RB	denotes	previously
T6511	22689-22692	IN	denotes	for
T6512	22693-22694	DT	denotes	a
T6514	22695-22700	JJ	denotes	human
T6515	22701-22710	JJ	denotes	olfactory
T6513	22711-22719	NN	denotes	receptor
T6516	22720-22721	-LRB-	denotes	[
T6517	22721-22723	CD	denotes	25
T6518	22723-22724	-RRB-	denotes	]
T6519	22724-22725	.	denotes	.
T6520	22725-22815	sentence	denotes	One of these two mouse genes has no start codon in its otherwise intact main coding exon.
T6521	22726-22729	CD	denotes	One
T6523	22730-22732	IN	denotes	of
T6524	22733-22738	DT	denotes	these
T6526	22739-22742	CD	denotes	two
T6527	22743-22748	NN	denotes	mouse
T6525	22749-22754	NNS	denotes	genes
T6522	22755-22758	VBZ	denotes	has
T6528	22759-22761	DT	denotes	no
T6530	22762-22767	NN	denotes	start
T6529	22768-22773	NN	denotes	codon
T6531	22774-22776	IN	denotes	in
T6532	22777-22780	PRP$	denotes	its
T6534	22781-22790	RB	denotes	otherwise
T6535	22791-22797	JJ	denotes	intact
T6536	22798-22802	JJ	denotes	main
T6537	22803-22809	VBG	denotes	coding
T6533	22810-22814	NN	denotes	exon
T6538	22814-22815	.	denotes	.
T6539	22815-22895	sentence	denotes	The unusual splicing thus rescues what would otherwise be a dysfunctional gene.
T6540	22816-22819	DT	denotes	The
T6542	22820-22827	JJ	denotes	unusual
T6541	22828-22836	NN	denotes	splicing
T6544	22837-22841	RB	denotes	thus
T6543	22842-22849	VBZ	denotes	rescues
T6545	22850-22854	WP	denotes	what
T6547	22855-22860	MD	denotes	would
T6548	22861-22870	RB	denotes	otherwise
T6546	22871-22873	VB	denotes	be
T6549	22874-22875	DT	denotes	a
T6551	22876-22889	JJ	denotes	dysfunctional
T6550	22890-22894	NN	denotes	gene
T6552	22894-22895	.	denotes	.
T6553	22895-23152	sentence	denotes	In most cases (60 out of 62 cDNAs), the unusual transcript appears to be an aberrant splice form - the transcript would probably not encode a functional protein because the splice introduces a frameshift or removes conserved functional residues (Figure 6).
T6554	22896-22898	IN	denotes	In
T6556	22899-22903	JJS	denotes	most
T6557	22904-22909	NNS	denotes	cases
T6558	22910-22911	-LRB-	denotes	(
T6559	22911-22913	CD	denotes	60
T6561	22914-22917	IN	denotes	out
T6562	22918-22920	IN	denotes	of
T6560	22921-22923	CD	denotes	62
T6563	22924-22929	NNS	denotes	cDNAs
T6564	22929-22930	-RRB-	denotes	)
T6565	22930-22932	,	denotes	,
T6566	22932-22935	DT	denotes	the
T6568	22936-22943	JJ	denotes	unusual
T6567	22944-22954	NN	denotes	transcript
T6569	22955-22962	VBZ	denotes	appears
T6570	22963-22965	TO	denotes	to
T6571	22966-22968	VB	denotes	be
T6572	22969-22971	DT	denotes	an
T6574	22972-22980	JJ	denotes	aberrant
T6575	22981-22987	NN	denotes	splice
T6573	22988-22992	NN	denotes	form
T6576	22993-22994	:	denotes	-
T6577	22995-22998	DT	denotes	the
T6578	22999-23009	NN	denotes	transcript
T6579	23010-23015	MD	denotes	would
T6580	23016-23024	RB	denotes	probably
T6581	23025-23028	RB	denotes	not
T6555	23029-23035	VB	denotes	encode
T6582	23036-23037	DT	denotes	a
T6584	23038-23048	JJ	denotes	functional
T6583	23049-23056	NN	denotes	protein
T6585	23057-23064	IN	denotes	because
T6587	23065-23068	DT	denotes	the
T6588	23069-23075	NN	denotes	splice
T6586	23076-23086	VBZ	denotes	introduces
T6589	23087-23088	DT	denotes	a
T6590	23089-23099	NN	denotes	frameshift
T6591	23100-23102	CC	denotes	or
T6592	23103-23110	VBZ	denotes	removes
T6593	23111-23120	VBN	denotes	conserved
T6595	23121-23131	JJ	denotes	functional
T6594	23132-23140	NNS	denotes	residues
T6596	23141-23142	-LRB-	denotes	(
T6597	23142-23148	NN	denotes	Figure
T6598	23149-23150	CD	denotes	6
T6599	23150-23151	-RRB-	denotes	)
T6600	23151-23152	.	denotes	.
T6601	23152-23283	sentence	denotes	For two clones (bottom two cDNAs, Figure 6), exon order in the cDNA clone is inconsistent with the corresponding genomic sequence.
T6602	23153-23156	IN	denotes	For
T6604	23157-23160	CD	denotes	two
T6605	23161-23167	NNS	denotes	clones
T6606	23168-23169	-LRB-	denotes	(
T6608	23169-23175	JJ	denotes	bottom
T6610	23176-23179	CD	denotes	two
T6609	23180-23185	NNS	denotes	cDNAs
T6611	23185-23187	,	denotes	,
T6607	23187-23193	NN	denotes	Figure
T6612	23194-23195	CD	denotes	6
T6613	23195-23196	-RRB-	denotes	)
T6614	23196-23198	,	denotes	,
T6615	23198-23202	NN	denotes	exon
T6616	23203-23208	NN	denotes	order
T6617	23209-23211	IN	denotes	in
T6618	23212-23215	DT	denotes	the
T6620	23216-23220	NN	denotes	cDNA
T6619	23221-23226	NN	denotes	clone
T6603	23227-23229	VBZ	denotes	is
T6621	23230-23242	JJ	denotes	inconsistent
T6622	23243-23247	IN	denotes	with
T6623	23248-23251	DT	denotes	the
T6625	23252-23265	JJ	denotes	corresponding
T6626	23266-23273	JJ	denotes	genomic
T6624	23274-23282	NN	denotes	sequence
T6627	23282-23283	.	denotes	.
T6628	23283-23444	sentence	denotes	It is difficult to imagine what kind of cloning artefact resulted in these severely scrambled cDNAs: we suggest that they derive from real but rare transcripts.
T6629	23284-23286	PRP	denotes	It
T6630	23287-23289	VBZ	denotes	is
T6632	23290-23299	JJ	denotes	difficult
T6633	23300-23302	TO	denotes	to
T6634	23303-23310	VB	denotes	imagine
T6635	23311-23315	WDT	denotes	what
T6636	23316-23320	NN	denotes	kind
T6638	23321-23323	IN	denotes	of
T6639	23324-23331	VBG	denotes	cloning
T6640	23332-23340	NN	denotes	artefact
T6637	23341-23349	VBD	denotes	resulted
T6641	23350-23352	IN	denotes	in
T6642	23353-23358	DT	denotes	these
T6644	23359-23367	RB	denotes	severely
T6645	23368-23377	VBN	denotes	scrambled
T6643	23378-23383	NNS	denotes	cDNAs
T6646	23383-23385	:	denotes	:
T6647	23385-23387	PRP	denotes	we
T6631	23388-23395	VBP	denotes	suggest
T6648	23396-23400	IN	denotes	that
T6650	23401-23405	PRP	denotes	they
T6649	23406-23412	VBP	denotes	derive
T6651	23413-23417	IN	denotes	from
T6652	23418-23422	JJ	denotes	real
T6654	23423-23426	CC	denotes	but
T6655	23427-23431	JJ	denotes	rare
T6653	23432-23443	NNS	denotes	transcripts
T6656	23443-23444	.	denotes	.
T6657	23444-23635	sentence	denotes	However, their low frequency in our cDNA collection suggests that splicing contrary to genomic organization does not contribute significantly to the olfactory receptor transcript repertoire.
T6658	23445-23452	RB	denotes	However
T6660	23452-23454	,	denotes	,
T6661	23454-23459	PRP$	denotes	their
T6663	23460-23463	JJ	denotes	low
T6662	23464-23473	NN	denotes	frequency
T6664	23474-23476	IN	denotes	in
T6665	23477-23480	PRP$	denotes	our
T6667	23481-23485	NN	denotes	cDNA
T6666	23486-23496	NN	denotes	collection
T6659	23497-23505	VBZ	denotes	suggests
T6668	23506-23510	IN	denotes	that
T6670	23511-23519	NN	denotes	splicing
T6671	23520-23528	JJ	denotes	contrary
T6672	23529-23531	IN	denotes	to
T6673	23532-23539	JJ	denotes	genomic
T6674	23540-23552	NN	denotes	organization
T6675	23553-23557	VBZ	denotes	does
T6676	23558-23561	RB	denotes	not
T6669	23562-23572	VB	denotes	contribute
T6677	23573-23586	RB	denotes	significantly
T6678	23587-23589	IN	denotes	to
T6679	23590-23593	DT	denotes	the
T6681	23594-23603	JJ	denotes	olfactory
T6682	23604-23612	NN	denotes	receptor
T6683	23613-23623	NN	denotes	transcript
T6680	23624-23634	NN	denotes	repertoire
T6684	23634-23635	.	denotes	.
T6685	23635-23811	sentence	denotes	For 21 of the 26 genes for which unusually spliced cDNAs were found, we also observe an alternative ('normal') isoform that does not use splice sites within the coding region.
T6686	23636-23639	IN	denotes	For
T6688	23640-23642	CD	denotes	21
T6690	23643-23645	IN	denotes	of
T6691	23646-23649	DT	denotes	the
T6689	23650-23652	CD	denotes	26
T6692	23653-23658	NNS	denotes	genes
T6693	23659-23662	IN	denotes	for
T6695	23663-23668	WDT	denotes	which
T6696	23669-23678	RB	denotes	unusually
T6697	23679-23686	VBN	denotes	spliced
T6698	23687-23692	NNS	denotes	cDNAs
T6699	23693-23697	VBD	denotes	were
T6694	23698-23703	VBN	denotes	found
T6700	23703-23705	,	denotes	,
T6701	23705-23707	PRP	denotes	we
T6702	23708-23712	RB	denotes	also
T6687	23713-23720	VBP	denotes	observe
T6703	23721-23723	DT	denotes	an
T6705	23724-23735	JJ	denotes	alternative
T6706	23736-23737	-LRB-	denotes	(
T6707	23737-23738	``	denotes	'
T6708	23738-23744	JJ	denotes	normal
T6709	23744-23745	''	denotes	'
T6710	23745-23746	-RRB-	denotes	)
T6704	23747-23754	NN	denotes	isoform
T6711	23755-23759	WDT	denotes	that
T6713	23760-23764	VBZ	denotes	does
T6714	23765-23768	RB	denotes	not
T6712	23769-23772	VB	denotes	use
T6715	23773-23779	NN	denotes	splice
T6716	23780-23785	NNS	denotes	sites
T6717	23786-23792	IN	denotes	within
T6718	23793-23796	DT	denotes	the
T6720	23797-23803	VBG	denotes	coding
T6719	23804-23810	NN	denotes	region
T6721	23810-23811	.	denotes	.
T6722	23811-23965	sentence	denotes	(For the remaining 13 of the 3' genes showing odd splicing, we have identified only one cDNA so have not determined whether normal isoforms are present.)
T6723	23812-23813	-LRB-	denotes	(
T6725	23813-23816	IN	denotes	For
T6726	23817-23820	DT	denotes	the
T6728	23821-23830	VBG	denotes	remaining
T6727	23831-23833	CD	denotes	13
T6729	23834-23836	IN	denotes	of
T6730	23837-23840	DT	denotes	the
T6732	23841-23842	CD	denotes	3
T6733	23842-23843	SYM	denotes	'
T6731	23844-23849	NNS	denotes	genes
T6734	23850-23857	VBG	denotes	showing
T6735	23858-23861	JJ	denotes	odd
T6736	23862-23870	NN	denotes	splicing
T6737	23870-23872	,	denotes	,
T6738	23872-23874	PRP	denotes	we
T6739	23875-23879	VBP	denotes	have
T6724	23880-23890	VBN	denotes	identified
T6740	23891-23895	RB	denotes	only
T6742	23896-23899	CD	denotes	one
T6741	23900-23904	NN	denotes	cDNA
T6743	23905-23907	CC	denotes	so
T6744	23908-23912	VBP	denotes	have
T6746	23913-23916	RB	denotes	not
T6745	23917-23927	VBN	denotes	determined
T6747	23928-23935	IN	denotes	whether
T6749	23936-23942	JJ	denotes	normal
T6750	23943-23951	NNS	denotes	isoforms
T6748	23952-23955	VBP	denotes	are
T6751	23956-23963	JJ	denotes	present
T6752	23963-23964	.	denotes	.
T6753	23964-23965	-RRB-	denotes	)
T6754	23965-24391	sentence	denotes	We were intrigued both by previous reports of splicing of human olfactory receptors near the major histocompatibility complex (MHC) cluster, where several genes splice over long distances to a common upstream exon [26,27] and by the idea that olfactory receptor transcriptional control could be achieved by DNA recombination mechanisms, perhaps with the result that transcripts would contain some sequence from another locus.
T6755	23966-23968	PRP	denotes	We
T6757	23969-23973	VBD	denotes	were
T6756	23974-23983	VBN	denotes	intrigued
T6758	23984-23988	CC	denotes	both
T6759	23989-23991	IN	denotes	by
T6760	23992-24000	JJ	denotes	previous
T6761	24001-24008	NNS	denotes	reports
T6762	24009-24011	IN	denotes	of
T6763	24012-24020	NN	denotes	splicing
T6764	24021-24023	IN	denotes	of
T6765	24024-24029	JJ	denotes	human
T6767	24030-24039	JJ	denotes	olfactory
T6766	24040-24049	NNS	denotes	receptors
T6768	24050-24054	IN	denotes	near
T6769	24055-24058	DT	denotes	the
T6771	24059-24064	JJ	denotes	major
T6772	24065-24083	NN	denotes	histocompatibility
T6770	24084-24091	NN	denotes	complex
T6774	24092-24093	-LRB-	denotes	(
T6775	24093-24096	NN	denotes	MHC
T6776	24096-24097	-RRB-	denotes	)
T6773	24098-24105	NN	denotes	cluster
T6777	24105-24107	,	denotes	,
T6778	24107-24112	WRB	denotes	where
T6780	24113-24120	JJ	denotes	several
T6781	24121-24126	NNS	denotes	genes
T6779	24127-24133	VBP	denotes	splice
T6782	24134-24138	IN	denotes	over
T6783	24139-24143	JJ	denotes	long
T6784	24144-24153	NNS	denotes	distances
T6785	24154-24156	IN	denotes	to
T6786	24157-24158	DT	denotes	a
T6788	24159-24165	JJ	denotes	common
T6789	24166-24174	JJ	denotes	upstream
T6787	24175-24179	NN	denotes	exon
T6790	24180-24181	-LRB-	denotes	[
T6792	24181-24183	CD	denotes	26
T6793	24183-24184	,	denotes	,
T6791	24184-24186	CD	denotes	27
T6794	24186-24187	-RRB-	denotes	]
T6795	24188-24191	CC	denotes	and
T6796	24192-24194	IN	denotes	by
T6797	24195-24198	DT	denotes	the
T6798	24199-24203	NN	denotes	idea
T6799	24204-24208	IN	denotes	that
T6801	24209-24218	JJ	denotes	olfactory
T6802	24219-24227	NN	denotes	receptor
T6804	24228-24243	JJ	denotes	transcriptional
T6803	24244-24251	NN	denotes	control
T6805	24252-24257	MD	denotes	could
T6806	24258-24260	VB	denotes	be
T6800	24261-24269	VBN	denotes	achieved
T6807	24270-24272	IN	denotes	by
T6808	24273-24276	NN	denotes	DNA
T6810	24277-24290	NN	denotes	recombination
T6809	24291-24301	NNS	denotes	mechanisms
T6811	24301-24303	,	denotes	,
T6812	24303-24310	RB	denotes	perhaps
T6813	24311-24315	IN	denotes	with
T6814	24316-24319	DT	denotes	the
T6815	24320-24326	NN	denotes	result
T6816	24327-24331	IN	denotes	that
T6818	24332-24343	NNS	denotes	transcripts
T6819	24344-24349	MD	denotes	would
T6817	24350-24357	VB	denotes	contain
T6820	24358-24362	DT	denotes	some
T6821	24363-24371	NN	denotes	sequence
T6822	24372-24376	IN	denotes	from
T6823	24377-24384	DT	denotes	another
T6824	24385-24390	NN	denotes	locus
T6825	24390-24391	.	denotes	.
T6826	24391-24624	sentence	denotes	We therefore verified that the entire sequence of each olfactory receptor EST matches the corresponding gene's genomic 'territory' (defined for this purpose as from 1 kb after the preceding gene to 1 kb after the gene's stop codon).
T6827	24392-24394	PRP	denotes	We
T6829	24395-24404	RB	denotes	therefore
T6828	24405-24413	VBD	denotes	verified
T6830	24414-24418	IN	denotes	that
T6832	24419-24422	DT	denotes	the
T6834	24423-24429	JJ	denotes	entire
T6833	24430-24438	NN	denotes	sequence
T6835	24439-24441	IN	denotes	of
T6836	24442-24446	DT	denotes	each
T6838	24447-24456	JJ	denotes	olfactory
T6839	24457-24465	NN	denotes	receptor
T6837	24466-24469	NN	denotes	EST
T6831	24470-24477	VBZ	denotes	matches
T6840	24478-24481	DT	denotes	the
T6842	24482-24495	VBG	denotes	corresponding
T6841	24496-24500	NN	denotes	gene
T6844	24500-24502	POS	denotes	's
T6845	24503-24510	JJ	denotes	genomic
T6846	24511-24512	``	denotes	'
T6843	24512-24521	NN	denotes	territory
T6847	24521-24522	''	denotes	'
T6848	24523-24524	-LRB-	denotes	(
T6849	24524-24531	VBN	denotes	defined
T6850	24532-24535	IN	denotes	for
T6851	24536-24540	DT	denotes	this
T6852	24541-24548	NN	denotes	purpose
T6853	24549-24551	IN	denotes	as
T6854	24552-24556	IN	denotes	from
T6855	24557-24558	CD	denotes	1
T6856	24559-24561	NN	denotes	kb
T6857	24562-24567	IN	denotes	after
T6858	24568-24571	DT	denotes	the
T6860	24572-24581	VBG	denotes	preceding
T6859	24582-24586	NN	denotes	gene
T6861	24587-24589	IN	denotes	to
T6862	24590-24591	CD	denotes	1
T6863	24592-24594	NN	denotes	kb
T6864	24595-24600	IN	denotes	after
T6865	24601-24604	DT	denotes	the
T6866	24605-24609	NN	denotes	gene
T6868	24609-24611	POS	denotes	's
T6869	24612-24616	NN	denotes	stop
T6867	24617-24622	NN	denotes	codon
T6870	24622-24623	-RRB-	denotes	)
T6871	24623-24624	.	denotes	.
T6872	24624-24769	sentence	denotes	We found no cDNAs where introns encompassed other olfactory receptor genes, as reported for olfactory receptors in the human MHC region [26,27].
T6873	24625-24627	PRP	denotes	We
T6874	24628-24633	VBD	denotes	found
T6875	24634-24636	DT	denotes	no
T6876	24637-24642	NNS	denotes	cDNAs
T6877	24643-24648	WRB	denotes	where
T6879	24649-24656	NNS	denotes	introns
T6878	24657-24668	VBD	denotes	encompassed
T6880	24669-24674	JJ	denotes	other
T6882	24675-24684	JJ	denotes	olfactory
T6883	24685-24693	NN	denotes	receptor
T6881	24694-24699	NNS	denotes	genes
T6884	24699-24701	,	denotes	,
T6885	24701-24703	IN	denotes	as
T6886	24704-24712	VBN	denotes	reported
T6887	24713-24716	IN	denotes	for
T6888	24717-24726	JJ	denotes	olfactory
T6889	24727-24736	NNS	denotes	receptors
T6890	24737-24739	IN	denotes	in
T6891	24740-24743	DT	denotes	the
T6893	24744-24749	JJ	denotes	human
T6894	24750-24753	NN	denotes	MHC
T6892	24754-24760	NN	denotes	region
T6895	24761-24762	-LRB-	denotes	[
T6897	24762-24764	CD	denotes	26
T6898	24764-24765	,	denotes	,
T6896	24765-24767	CD	denotes	27
T6899	24767-24768	-RRB-	denotes	]
T6900	24768-24769	.	denotes	.
T6901	24769-24900	sentence	denotes	Six cDNAs do extend further than a single gene's 'territory' and appear not to be artifacts of the sequencing or analysis process.
T6902	24770-24773	CD	denotes	Six
T6903	24774-24779	NNS	denotes	cDNAs
T6905	24780-24782	VBP	denotes	do
T6904	24783-24789	VB	denotes	extend
T6906	24790-24797	RBR	denotes	further
T6907	24798-24802	IN	denotes	than
T6908	24803-24804	DT	denotes	a
T6910	24805-24811	JJ	denotes	single
T6909	24812-24816	NN	denotes	gene
T6912	24816-24818	POS	denotes	's
T6913	24819-24820	``	denotes	'
T6911	24820-24829	NN	denotes	territory
T6914	24829-24830	''	denotes	'
T6915	24831-24834	CC	denotes	and
T6916	24835-24841	VBP	denotes	appear
T6917	24842-24845	RB	denotes	not
T6919	24846-24848	TO	denotes	to
T6918	24849-24851	VB	denotes	be
T6920	24852-24861	NNS	denotes	artifacts
T6921	24862-24864	IN	denotes	of
T6922	24865-24868	DT	denotes	the
T6924	24869-24879	NN	denotes	sequencing
T6925	24880-24882	CC	denotes	or
T6926	24883-24891	NN	denotes	analysis
T6923	24892-24899	NN	denotes	process
T6927	24899-24900	.	denotes	.
T6928	24900-25048	sentence	denotes	In each of these cases, the clones use splice sites within the 3' UTR and thus extend further than the arbitrary 1 kb downstream of the stop codon.
T6929	24901-24903	IN	denotes	In
T6931	24904-24908	DT	denotes	each
T6932	24909-24911	IN	denotes	of
T6933	24912-24917	DT	denotes	these
T6934	24918-24923	NNS	denotes	cases
T6935	24923-24925	,	denotes	,
T6936	24925-24928	DT	denotes	the
T6937	24929-24935	NNS	denotes	clones
T6930	24936-24939	VBP	denotes	use
T6938	24940-24946	NN	denotes	splice
T6939	24947-24952	NNS	denotes	sites
T6940	24953-24959	IN	denotes	within
T6941	24960-24963	DT	denotes	the
T6943	24964-24965	CD	denotes	3
T6944	24965-24966	SYM	denotes	'
T6942	24967-24970	NN	denotes	UTR
T6945	24971-24974	CC	denotes	and
T6946	24975-24979	RB	denotes	thus
T6947	24980-24986	VBP	denotes	extend
T6948	24987-24994	RBR	denotes	further
T6950	24995-24999	IN	denotes	than
T6951	25000-25003	DT	denotes	the
T6953	25004-25013	JJ	denotes	arbitrary
T6954	25014-25015	CD	denotes	1
T6952	25016-25018	NN	denotes	kb
T6949	25019-25029	JJ	denotes	downstream
T6955	25030-25032	IN	denotes	of
T6956	25033-25036	DT	denotes	the
T6958	25037-25041	NN	denotes	stop
T6957	25042-25047	NN	denotes	codon
T6959	25047-25048	.	denotes	.
T6960	25048-25178	sentence	denotes	Five of these six cDNAs also use splice-donor sites within the coding region and encode disrupted olfactory receptors (Figure 6).
T6961	25049-25053	CD	denotes	Five
T6963	25054-25056	IN	denotes	of
T6964	25057-25062	DT	denotes	these
T6966	25063-25066	CD	denotes	six
T6965	25067-25072	NNS	denotes	cDNAs
T6967	25073-25077	RB	denotes	also
T6962	25078-25081	VBP	denotes	use
T6968	25082-25088	NN	denotes	splice
T6970	25088-25089	HYPH	denotes	-
T6969	25089-25094	NN	denotes	donor
T6971	25095-25100	NNS	denotes	sites
T6972	25101-25107	IN	denotes	within
T6973	25108-25111	DT	denotes	the
T6975	25112-25118	VBG	denotes	coding
T6974	25119-25125	NN	denotes	region
T6976	25126-25129	CC	denotes	and
T6977	25130-25136	VBP	denotes	encode
T6978	25137-25146	VBN	denotes	disrupted
T6980	25147-25156	JJ	denotes	olfactory
T6979	25157-25166	NNS	denotes	receptors
T6981	25167-25168	-LRB-	denotes	(
T6982	25168-25174	NN	denotes	Figure
T6983	25175-25176	CD	denotes	6
T6984	25176-25177	-RRB-	denotes	)
T6985	25177-25178	.	denotes	.
T6986	25178-25277	sentence	denotes	In the sixth cDNA, a 2.6-kb intron is spliced out of the 3' UTR, leaving the coding region intact.
T6987	25179-25181	IN	denotes	In
T6989	25182-25185	DT	denotes	the
T6991	25186-25191	JJ	denotes	sixth
T6990	25192-25196	NN	denotes	cDNA
T6992	25196-25198	,	denotes	,
T6993	25198-25199	DT	denotes	a
T6995	25200-25203	CD	denotes	2.6
T6997	25203-25204	HYPH	denotes	-
T6996	25204-25206	NN	denotes	kb
T6994	25207-25213	NN	denotes	intron
T6998	25214-25216	VBZ	denotes	is
T6988	25217-25224	VBN	denotes	spliced
T6999	25225-25228	IN	denotes	out
T7000	25229-25231	IN	denotes	of
T7001	25232-25235	DT	denotes	the
T7003	25236-25237	CD	denotes	3
T7004	25237-25238	SYM	denotes	'
T7002	25239-25242	NN	denotes	UTR
T7005	25242-25244	,	denotes	,
T7006	25244-25251	VBG	denotes	leaving
T7007	25252-25255	DT	denotes	the
T7009	25256-25262	VBG	denotes	coding
T7008	25263-25269	NN	denotes	region
T7010	25270-25276	JJ	denotes	intact
T7011	25276-25277	.	denotes	.
T7012	25277-25512	sentence	denotes	If olfactory receptor transcriptional control is achieved by DNA recombination, the beginning of each transcript might derive from a donated promoter region, with the rest of the transcript coming from the native ORF-containing locus.
T7013	25278-25280	IN	denotes	If
T7015	25281-25290	JJ	denotes	olfactory
T7016	25291-25299	NN	denotes	receptor
T7018	25300-25315	JJ	denotes	transcriptional
T7017	25316-25323	NN	denotes	control
T7019	25324-25326	VBZ	denotes	is
T7014	25327-25335	VBN	denotes	achieved
T7021	25336-25338	IN	denotes	by
T7022	25339-25342	NN	denotes	DNA
T7023	25343-25356	NN	denotes	recombination
T7024	25356-25358	,	denotes	,
T7025	25358-25361	DT	denotes	the
T7026	25362-25371	NN	denotes	beginning
T7027	25372-25374	IN	denotes	of
T7028	25375-25379	DT	denotes	each
T7029	25380-25390	NN	denotes	transcript
T7030	25391-25396	MD	denotes	might
T7020	25397-25403	VB	denotes	derive
T7031	25404-25408	IN	denotes	from
T7032	25409-25410	DT	denotes	a
T7034	25411-25418	VBN	denotes	donated
T7035	25419-25427	NN	denotes	promoter
T7033	25428-25434	NN	denotes	region
T7036	25434-25436	,	denotes	,
T7037	25436-25440	IN	denotes	with
T7038	25441-25444	DT	denotes	the
T7039	25445-25449	NN	denotes	rest
T7041	25450-25452	IN	denotes	of
T7042	25453-25456	DT	denotes	the
T7043	25457-25467	NN	denotes	transcript
T7040	25468-25474	VBG	denotes	coming
T7044	25475-25479	IN	denotes	from
T7045	25480-25483	DT	denotes	the
T7047	25484-25490	JJ	denotes	native
T7048	25491-25494	NN	denotes	ORF
T7050	25494-25495	HYPH	denotes	-
T7049	25495-25505	VBG	denotes	containing
T7046	25506-25511	NN	denotes	locus
T7051	25511-25512	.	denotes	.
T7052	25512-25671	sentence	denotes	In order to examine the recombination hypothesis, we analyzed 115 cDNA clones for which sim4 failed to align 20 bp or more to the corresponding genomic locus.
T7053	25513-25515	IN	denotes	In
T7055	25516-25521	NN	denotes	order
T7056	25522-25524	TO	denotes	to
T7057	25525-25532	VB	denotes	examine
T7058	25533-25536	DT	denotes	the
T7060	25537-25550	NN	denotes	recombination
T7059	25551-25561	NN	denotes	hypothesis
T7061	25561-25563	,	denotes	,
T7062	25563-25565	PRP	denotes	we
T7054	25566-25574	VBD	denotes	analyzed
T7063	25575-25578	CD	denotes	115
T7065	25579-25583	NN	denotes	cDNA
T7064	25584-25590	NNS	denotes	clones
T7066	25591-25594	IN	denotes	for
T7068	25595-25600	WDT	denotes	which
T7069	25601-25605	NN	denotes	sim4
T7067	25606-25612	VBD	denotes	failed
T7070	25613-25615	TO	denotes	to
T7071	25616-25621	VB	denotes	align
T7072	25622-25624	CD	denotes	20
T7073	25625-25627	NNS	denotes	bp
T7074	25628-25630	CC	denotes	or
T7075	25631-25635	JJR	denotes	more
T7076	25636-25638	IN	denotes	to
T7077	25639-25642	DT	denotes	the
T7079	25643-25656	VBG	denotes	corresponding
T7080	25657-25664	JJ	denotes	genomic
T7078	25665-25670	NN	denotes	locus
T7081	25670-25671	.	denotes	.
T7082	25671-25840	sentence	denotes	In most cases, the missing sequence was explained by gaps in the genomic sequence or by matches that fell below our percent identity-based cutoff for reporting matches.
T7083	25672-25674	IN	denotes	In
T7085	25675-25679	JJS	denotes	most
T7086	25680-25685	NNS	denotes	cases
T7087	25685-25687	,	denotes	,
T7088	25687-25690	DT	denotes	the
T7090	25691-25698	JJ	denotes	missing
T7089	25699-25707	NN	denotes	sequence
T7091	25708-25711	VBD	denotes	was
T7084	25712-25721	VBN	denotes	explained
T7092	25722-25724	IN	denotes	by
T7093	25725-25729	NNS	denotes	gaps
T7094	25730-25732	IN	denotes	in
T7095	25733-25736	DT	denotes	the
T7097	25737-25744	JJ	denotes	genomic
T7096	25745-25753	NN	denotes	sequence
T7098	25754-25756	CC	denotes	or
T7099	25757-25759	IN	denotes	by
T7100	25760-25767	NNS	denotes	matches
T7101	25768-25772	WDT	denotes	that
T7102	25773-25777	VBD	denotes	fell
T7103	25778-25783	IN	denotes	below
T7104	25784-25787	PRP$	denotes	our
T7106	25788-25795	NN	denotes	percent
T7107	25796-25804	NN	denotes	identity
T7109	25804-25805	HYPH	denotes	-
T7108	25805-25810	VBN	denotes	based
T7105	25811-25817	NN	denotes	cutoff
T7110	25818-25821	IN	denotes	for
T7111	25822-25831	VBG	denotes	reporting
T7112	25832-25839	NNS	denotes	matches
T7113	25839-25840	.	denotes	.
T7114	25840-25977	sentence	denotes	For three cDNAs (from three different olfactory receptors), we found that the missing piece of sequence matched elsewhere in the genome.
T7115	25841-25844	IN	denotes	For
T7117	25845-25850	CD	denotes	three
T7118	25851-25856	NNS	denotes	cDNAs
T7119	25857-25858	-LRB-	denotes	(
T7120	25858-25862	IN	denotes	from
T7121	25863-25868	CD	denotes	three
T7123	25869-25878	JJ	denotes	different
T7124	25879-25888	JJ	denotes	olfactory
T7122	25889-25898	NNS	denotes	receptors
T7125	25898-25899	-RRB-	denotes	)
T7126	25899-25901	,	denotes	,
T7127	25901-25903	PRP	denotes	we
T7116	25904-25909	VBD	denotes	found
T7128	25910-25914	IN	denotes	that
T7130	25915-25918	DT	denotes	the
T7132	25919-25926	JJ	denotes	missing
T7131	25927-25932	NN	denotes	piece
T7133	25933-25935	IN	denotes	of
T7134	25936-25944	NN	denotes	sequence
T7129	25945-25952	VBN	denotes	matched
T7135	25953-25962	RB	denotes	elsewhere
T7136	25963-25965	IN	denotes	in
T7137	25966-25969	DT	denotes	the
T7138	25970-25976	NN	denotes	genome
T7139	25976-25977	.	denotes	.
T7140	25977-26057	sentence	denotes	Comparison with the public mouse genome assembly confirmed the distant matches.
T7141	25978-25988	NN	denotes	Comparison
T7143	25989-25993	IN	denotes	with
T7144	25994-25997	DT	denotes	the
T7146	25998-26004	JJ	denotes	public
T7147	26005-26010	NN	denotes	mouse
T7148	26011-26017	NN	denotes	genome
T7145	26018-26026	NN	denotes	assembly
T7142	26027-26036	VBD	denotes	confirmed
T7149	26037-26040	DT	denotes	the
T7151	26041-26048	JJ	denotes	distant
T7150	26049-26056	NNS	denotes	matches
T7152	26056-26057	.	denotes	.
T7153	26057-26262	sentence	denotes	With such a small number of cDNAs exhibiting a possible sign of DNA recombination (a sign that could also be interpreted as chimeric cDNA clones), we conclude that such rearrangement is unlikely to occur.
T7154	26058-26062	IN	denotes	With
T7156	26063-26067	PDT	denotes	such
T7158	26068-26069	DT	denotes	a
T7159	26070-26075	JJ	denotes	small
T7157	26076-26082	NN	denotes	number
T7161	26083-26085	IN	denotes	of
T7162	26086-26091	NNS	denotes	cDNAs
T7160	26092-26102	VBG	denotes	exhibiting
T7163	26103-26104	DT	denotes	a
T7165	26105-26113	JJ	denotes	possible
T7164	26114-26118	NN	denotes	sign
T7166	26119-26121	IN	denotes	of
T7167	26122-26125	NN	denotes	DNA
T7168	26126-26139	NN	denotes	recombination
T7169	26140-26141	-LRB-	denotes	(
T7170	26141-26142	DT	denotes	a
T7171	26143-26147	NN	denotes	sign
T7172	26148-26152	WDT	denotes	that
T7174	26153-26158	MD	denotes	could
T7175	26159-26163	RB	denotes	also
T7176	26164-26166	VB	denotes	be
T7173	26167-26178	VBN	denotes	interpreted
T7177	26179-26181	IN	denotes	as
T7178	26182-26190	JJ	denotes	chimeric
T7180	26191-26195	NN	denotes	cDNA
T7179	26196-26202	NNS	denotes	clones
T7181	26202-26203	-RRB-	denotes	)
T7182	26203-26205	,	denotes	,
T7183	26205-26207	PRP	denotes	we
T7155	26208-26216	VBP	denotes	conclude
T7184	26217-26221	IN	denotes	that
T7186	26222-26226	JJ	denotes	such
T7187	26227-26240	NN	denotes	rearrangement
T7185	26241-26243	VBZ	denotes	is
T7188	26244-26252	JJ	denotes	unlikely
T7189	26253-26255	TO	denotes	to
T7190	26256-26261	VB	denotes	occur
T7191	26261-26262	.	denotes	.
T7192	26262-26476	sentence	denotes	However, the possibility remains that DNA recombination is responsible for olfactory receptor transcriptional regulation, with the donated region contributing only promoter sequences but no part of the transcript.
T7193	26263-26270	RB	denotes	However
T7195	26270-26272	,	denotes	,
T7196	26272-26275	DT	denotes	the
T7197	26276-26287	NN	denotes	possibility
T7194	26288-26295	VBZ	denotes	remains
T7198	26296-26300	IN	denotes	that
T7200	26301-26304	NN	denotes	DNA
T7201	26305-26318	NN	denotes	recombination
T7199	26319-26321	VBZ	denotes	is
T7202	26322-26333	JJ	denotes	responsible
T7203	26334-26337	IN	denotes	for
T7204	26338-26347	JJ	denotes	olfactory
T7205	26348-26356	NN	denotes	receptor
T7207	26357-26372	JJ	denotes	transcriptional
T7206	26373-26383	NN	denotes	regulation
T7208	26383-26385	,	denotes	,
T7209	26385-26389	IN	denotes	with
T7210	26390-26393	DT	denotes	the
T7212	26394-26401	VBN	denotes	donated
T7211	26402-26408	NN	denotes	region
T7213	26409-26421	VBG	denotes	contributing
T7214	26422-26426	RB	denotes	only
T7216	26427-26435	NN	denotes	promoter
T7215	26436-26445	NNS	denotes	sequences
T7217	26446-26449	CC	denotes	but
T7218	26450-26452	DT	denotes	no
T7219	26453-26457	NN	denotes	part
T7220	26458-26460	IN	denotes	of
T7221	26461-26464	DT	denotes	the
T7222	26465-26475	NN	denotes	transcript
T7223	26475-26476	.	denotes	.
T7399	26478-26482	CC	denotes	Both
T7401	26483-26494	JJ	denotes	unclustered
T7402	26495-26504	JJ	denotes	olfactory
T7400	26505-26514	NNS	denotes	receptors
T7404	26515-26518	CC	denotes	and
T7405	26519-26528	JJ	denotes	olfactory
T7406	26529-26537	NN	denotes	receptor
T7407	26538-26549	NNS	denotes	pseudogenes
T7408	26550-26553	MD	denotes	can
T7409	26554-26556	VB	denotes	be
T7403	26557-26566	VBN	denotes	expressed
T7410	26566-26868	sentence	denotes	We were interested in whether olfactory receptors need to be part of a cluster in the genome in order to be transcribed, or if the clustered genomic organization of olfactory receptors is simply a consequence of the fact that local duplication is the major mechanism for expanding the gene family [1].
T7411	26567-26569	PRP	denotes	We
T7412	26570-26574	VBD	denotes	were
T7413	26575-26585	JJ	denotes	interested
T7414	26586-26588	IN	denotes	in
T7415	26589-26596	IN	denotes	whether
T7417	26597-26606	JJ	denotes	olfactory
T7418	26607-26616	NNS	denotes	receptors
T7416	26617-26621	VBP	denotes	need
T7419	26622-26624	TO	denotes	to
T7420	26625-26627	VB	denotes	be
T7421	26628-26632	NN	denotes	part
T7422	26633-26635	IN	denotes	of
T7423	26636-26637	DT	denotes	a
T7424	26638-26645	NN	denotes	cluster
T7425	26646-26648	IN	denotes	in
T7426	26649-26652	DT	denotes	the
T7427	26653-26659	NN	denotes	genome
T7428	26660-26662	IN	denotes	in
T7429	26663-26668	NN	denotes	order
T7430	26669-26671	TO	denotes	to
T7432	26672-26674	VB	denotes	be
T7431	26675-26686	VBN	denotes	transcribed
T7433	26686-26688	,	denotes	,
T7434	26688-26690	CC	denotes	or
T7435	26691-26693	IN	denotes	if
T7437	26694-26697	DT	denotes	the
T7439	26698-26707	JJ	denotes	clustered
T7440	26708-26715	JJ	denotes	genomic
T7438	26716-26728	NN	denotes	organization
T7441	26729-26731	IN	denotes	of
T7442	26732-26741	JJ	denotes	olfactory
T7443	26742-26751	NNS	denotes	receptors
T7436	26752-26754	VBZ	denotes	is
T7444	26755-26761	RB	denotes	simply
T7445	26762-26763	DT	denotes	a
T7446	26764-26775	NN	denotes	consequence
T7447	26776-26778	IN	denotes	of
T7448	26779-26782	DT	denotes	the
T7449	26783-26787	NN	denotes	fact
T7450	26788-26792	IN	denotes	that
T7452	26793-26798	JJ	denotes	local
T7453	26799-26810	NN	denotes	duplication
T7451	26811-26813	VBZ	denotes	is
T7454	26814-26817	DT	denotes	the
T7456	26818-26823	JJ	denotes	major
T7455	26824-26833	NN	denotes	mechanism
T7457	26834-26837	IN	denotes	for
T7458	26838-26847	VBG	denotes	expanding
T7459	26848-26851	DT	denotes	the
T7461	26852-26856	NN	denotes	gene
T7460	26857-26863	NN	denotes	family
T7462	26864-26865	-LRB-	denotes	[
T7463	26865-26866	CD	denotes	1
T7464	26866-26867	-RRB-	denotes	]
T7465	26867-26868	.	denotes	.
T7466	26868-27122	sentence	denotes	'Singleton' olfactory receptors (defined as full-length olfactory receptors without another olfactory receptor within 0.5 Mb) are more often pseudogenes than are olfactory receptors in clusters (8 out of 16 versus 271 out of 1,358; χ2 = 8.8, P < 0.005).
T7467	26869-26870	``	denotes	'
T7469	26870-26879	NN	denotes	Singleton
T7471	26879-26880	''	denotes	'
T7472	26881-26890	JJ	denotes	olfactory
T7470	26891-26900	NNS	denotes	receptors
T7473	26901-26902	-LRB-	denotes	(
T7474	26902-26909	VBN	denotes	defined
T7475	26910-26912	IN	denotes	as
T7476	26913-26917	JJ	denotes	full
T7478	26917-26918	HYPH	denotes	-
T7477	26918-26924	NN	denotes	length
T7480	26925-26934	JJ	denotes	olfactory
T7479	26935-26944	NNS	denotes	receptors
T7481	26945-26952	IN	denotes	without
T7482	26953-26960	DT	denotes	another
T7484	26961-26970	JJ	denotes	olfactory
T7483	26971-26979	NN	denotes	receptor
T7485	26980-26986	IN	denotes	within
T7486	26987-26990	CD	denotes	0.5
T7487	26991-26993	NNS	denotes	Mb
T7488	26993-26994	-RRB-	denotes	)
T7468	26995-26998	VBP	denotes	are
T7489	26999-27003	RBR	denotes	more
T7490	27004-27009	RB	denotes	often
T7491	27010-27021	NNS	denotes	pseudogenes
T7492	27022-27026	IN	denotes	than
T7493	27027-27030	VBP	denotes	are
T7494	27031-27040	JJ	denotes	olfactory
T7495	27041-27050	NNS	denotes	receptors
T7496	27051-27053	IN	denotes	in
T7497	27054-27062	NNS	denotes	clusters
T7498	27063-27064	-LRB-	denotes	(
T7500	27064-27065	CD	denotes	8
T7501	27066-27069	IN	denotes	out
T7502	27070-27072	IN	denotes	of
T7499	27073-27075	CD	denotes	16
T7503	27076-27082	CC	denotes	versus
T7504	27083-27086	CD	denotes	271
T7506	27087-27090	IN	denotes	out
T7507	27091-27093	IN	denotes	of
T7505	27094-27099	CD	denotes	1,358
T7508	27099-27100	:	denotes	;
T7509	27101-27103	NN	denotes	χ2
T7511	27104-27105	SYM	denotes	=
T7510	27106-27109	CD	denotes	8.8
T7512	27109-27111	,	denotes	,
T7513	27111-27112	NN	denotes	P
T7515	27113-27114	SYM	denotes	<
T7514	27115-27120	CD	denotes	0.005
T7516	27120-27121	-RRB-	denotes	)
T7517	27121-27122	.	denotes	.
T7518	27122-27373	sentence	denotes	Of the eight intact singleton olfactory receptors, two have matching cDNAs in our collection, a similar proportion as found for olfactory receptors in clusters, showing that clustering is not an absolute requirement for olfactory receptor expression.
T7519	27123-27125	IN	denotes	Of
T7521	27126-27129	DT	denotes	the
T7523	27130-27135	CD	denotes	eight
T7524	27136-27142	JJ	denotes	intact
T7525	27143-27152	NN	denotes	singleton
T7526	27153-27162	JJ	denotes	olfactory
T7522	27163-27172	NNS	denotes	receptors
T7527	27172-27174	,	denotes	,
T7528	27174-27177	CD	denotes	two
T7520	27178-27182	VBP	denotes	have
T7529	27183-27191	VBG	denotes	matching
T7530	27192-27197	NNS	denotes	cDNAs
T7531	27198-27200	IN	denotes	in
T7532	27201-27204	PRP$	denotes	our
T7533	27205-27215	NN	denotes	collection
T7534	27215-27217	,	denotes	,
T7535	27217-27218	DT	denotes	a
T7537	27219-27226	JJ	denotes	similar
T7536	27227-27237	NN	denotes	proportion
T7538	27238-27240	IN	denotes	as
T7539	27241-27246	VBN	denotes	found
T7540	27247-27250	IN	denotes	for
T7541	27251-27260	JJ	denotes	olfactory
T7542	27261-27270	NNS	denotes	receptors
T7543	27271-27273	IN	denotes	in
T7544	27274-27282	NNS	denotes	clusters
T7545	27282-27284	,	denotes	,
T7546	27284-27291	VBG	denotes	showing
T7547	27292-27296	IN	denotes	that
T7549	27297-27307	VBG	denotes	clustering
T7548	27308-27310	VBZ	denotes	is
T7550	27311-27314	RB	denotes	not
T7551	27315-27317	DT	denotes	an
T7553	27318-27326	JJ	denotes	absolute
T7552	27327-27338	NN	denotes	requirement
T7554	27339-27342	IN	denotes	for
T7555	27343-27352	JJ	denotes	olfactory
T7556	27353-27361	NN	denotes	receptor
T7557	27362-27372	NN	denotes	expression
T7558	27372-27373	.	denotes	.
T7559	27373-27571	sentence	denotes	However, it is possible these two expressed singleton genes are part of 'extended' olfactory receptor clusters - their nearest olfactory receptor neighbors are 1.7 Mb and 2.6 Mb away, respectively.
T7560	27374-27381	RB	denotes	However
T7562	27381-27383	,	denotes	,
T7563	27383-27385	PRP	denotes	it
T7564	27386-27388	VBZ	denotes	is
T7565	27389-27397	JJ	denotes	possible
T7566	27398-27403	DT	denotes	these
T7568	27404-27407	CD	denotes	two
T7569	27408-27417	VBN	denotes	expressed
T7570	27418-27427	NN	denotes	singleton
T7567	27428-27433	NNS	denotes	genes
T7571	27434-27437	VBP	denotes	are
T7572	27438-27442	NN	denotes	part
T7573	27443-27445	IN	denotes	of
T7574	27446-27447	``	denotes	'
T7575	27447-27455	VBN	denotes	extended
T7577	27455-27456	''	denotes	'
T7578	27457-27466	JJ	denotes	olfactory
T7579	27467-27475	NN	denotes	receptor
T7576	27476-27484	NNS	denotes	clusters
T7580	27485-27486	:	denotes	-
T7581	27487-27492	PRP$	denotes	their
T7583	27493-27500	JJS	denotes	nearest
T7584	27501-27510	JJ	denotes	olfactory
T7585	27511-27519	NN	denotes	receptor
T7582	27520-27529	NNS	denotes	neighbors
T7561	27530-27533	VBP	denotes	are
T7586	27534-27537	CD	denotes	1.7
T7587	27538-27540	NNS	denotes	Mb
T7589	27541-27544	CC	denotes	and
T7590	27545-27548	CD	denotes	2.6
T7591	27549-27551	NNS	denotes	Mb
T7588	27552-27556	RB	denotes	away
T7592	27556-27558	,	denotes	,
T7593	27558-27570	RB	denotes	respectively
T7594	27570-27571	.	denotes	.
T7595	27571-27705	sentence	denotes	We also find that some olfactory receptor pseudogenes are expressed, albeit with a lower probability than intact olfactory receptors.
T7596	27572-27574	PRP	denotes	We
T7598	27575-27579	RB	denotes	also
T7597	27580-27584	VBP	denotes	find
T7599	27585-27589	IN	denotes	that
T7601	27590-27594	DT	denotes	some
T7603	27595-27604	JJ	denotes	olfactory
T7604	27605-27613	NN	denotes	receptor
T7602	27614-27625	NNS	denotes	pseudogenes
T7605	27626-27629	VBP	denotes	are
T7600	27630-27639	VBN	denotes	expressed
T7606	27639-27641	,	denotes	,
T7607	27641-27647	IN	denotes	albeit
T7608	27648-27652	IN	denotes	with
T7609	27653-27654	DT	denotes	a
T7611	27655-27660	JJR	denotes	lower
T7610	27661-27672	NN	denotes	probability
T7612	27673-27677	IN	denotes	than
T7613	27678-27684	JJ	denotes	intact
T7615	27685-27694	JJ	denotes	olfactory
T7614	27695-27704	NNS	denotes	receptors
T7616	27704-27705	.	denotes	.
T7617	27705-27955	sentence	denotes	Considering the 1,392 olfactory receptor gene sequences for which reliable full-length data are available, 15 out of 285 (5%) apparent pseudogenes are represented in our cDNA collection, compared to 393 out of 1,107 (36%) intact olfactory receptors.
T7618	27706-27717	VBG	denotes	Considering
T7620	27718-27721	DT	denotes	the
T7622	27722-27727	CD	denotes	1,392
T7623	27728-27737	JJ	denotes	olfactory
T7624	27738-27746	NN	denotes	receptor
T7625	27747-27751	NN	denotes	gene
T7621	27752-27761	NNS	denotes	sequences
T7626	27762-27765	IN	denotes	for
T7628	27766-27771	WDT	denotes	which
T7629	27772-27780	JJ	denotes	reliable
T7631	27781-27785	JJ	denotes	full
T7633	27785-27786	HYPH	denotes	-
T7632	27786-27792	NN	denotes	length
T7630	27793-27797	NNS	denotes	data
T7627	27798-27801	VBP	denotes	are
T7634	27802-27811	JJ	denotes	available
T7635	27811-27813	,	denotes	,
T7636	27813-27815	CD	denotes	15
T7638	27816-27819	IN	denotes	out
T7639	27820-27822	IN	denotes	of
T7637	27823-27826	CD	denotes	285
T7641	27827-27828	-LRB-	denotes	(
T7643	27828-27829	CD	denotes	5
T7642	27829-27830	NN	denotes	%
T7644	27830-27831	-RRB-	denotes	)
T7645	27832-27840	JJ	denotes	apparent
T7640	27841-27852	NNS	denotes	pseudogenes
T7646	27853-27856	VBP	denotes	are
T7619	27857-27868	VBN	denotes	represented
T7647	27869-27871	IN	denotes	in
T7648	27872-27875	PRP$	denotes	our
T7650	27876-27880	NN	denotes	cDNA
T7649	27881-27891	NN	denotes	collection
T7651	27891-27893	,	denotes	,
T7652	27893-27901	VBN	denotes	compared
T7653	27902-27904	IN	denotes	to
T7654	27905-27908	CD	denotes	393
T7656	27909-27912	IN	denotes	out
T7657	27913-27915	IN	denotes	of
T7655	27916-27921	CD	denotes	1,107
T7659	27922-27923	-LRB-	denotes	(
T7661	27923-27925	CD	denotes	36
T7660	27925-27926	NN	denotes	%
T7662	27926-27927	-RRB-	denotes	)
T7663	27928-27934	JJ	denotes	intact
T7664	27935-27944	JJ	denotes	olfactory
T7658	27945-27954	NNS	denotes	receptors
T7665	27954-27955	.	denotes	.
T7666	27955-28060	sentence	denotes	However, three of these 15 'expressed pseudogenes' are intact genes in the public mouse genome sequence.
T7667	27956-27963	RB	denotes	However
T7669	27963-27965	,	denotes	,
T7670	27965-27970	CD	denotes	three
T7672	27971-27973	IN	denotes	of
T7673	27974-27979	DT	denotes	these
T7671	27980-27982	CD	denotes	15
T7675	27983-27984	``	denotes	'
T7676	27984-27993	JJ	denotes	expressed
T7674	27994-28005	NNS	denotes	pseudogenes
T7677	28005-28006	''	denotes	'
T7668	28007-28010	VBP	denotes	are
T7678	28011-28017	JJ	denotes	intact
T7679	28018-28023	NNS	denotes	genes
T7680	28024-28026	IN	denotes	in
T7681	28027-28030	DT	denotes	the
T7683	28031-28037	JJ	denotes	public
T7684	28038-28043	NN	denotes	mouse
T7685	28044-28050	NN	denotes	genome
T7682	28051-28059	NN	denotes	sequence
T7686	28059-28060	.	denotes	.
T7687	28060-28161	sentence	denotes	The defects in Celera's version of these genes may be due to sequencing errors or true polymorphism.
T7688	28061-28064	DT	denotes	The
T7689	28065-28072	NNS	denotes	defects
T7691	28073-28075	IN	denotes	in
T7692	28076-28082	NNP	denotes	Celera
T7694	28082-28084	POS	denotes	's
T7693	28085-28092	NN	denotes	version
T7695	28093-28095	IN	denotes	of
T7696	28096-28101	DT	denotes	these
T7697	28102-28107	NNS	denotes	genes
T7698	28108-28111	MD	denotes	may
T7690	28112-28114	VB	denotes	be
T7699	28115-28118	IN	denotes	due
T7700	28119-28121	IN	denotes	to
T7701	28122-28132	VBG	denotes	sequencing
T7702	28133-28139	NNS	denotes	errors
T7703	28140-28142	CC	denotes	or
T7704	28143-28147	JJ	denotes	true
T7705	28148-28160	NN	denotes	polymorphism
T7706	28160-28161	.	denotes	.
T7707	28161-28278	sentence	denotes	Publicly available mouse sequence confirms that 11 of the 12 remaining expressed pseudogenes are indeed pseudogenes.
T7708	28162-28170	RB	denotes	Publicly
T7709	28171-28180	JJ	denotes	available
T7711	28181-28186	NN	denotes	mouse
T7710	28187-28195	NN	denotes	sequence
T7712	28196-28204	VBZ	denotes	confirms
T7713	28205-28209	IN	denotes	that
T7715	28210-28212	CD	denotes	11
T7717	28213-28215	IN	denotes	of
T7718	28216-28219	DT	denotes	the
T7716	28220-28222	CD	denotes	12
T7720	28223-28232	VBG	denotes	remaining
T7721	28233-28242	JJ	denotes	expressed
T7719	28243-28254	NNS	denotes	pseudogenes
T7714	28255-28258	VBP	denotes	are
T7722	28259-28265	RB	denotes	indeed
T7723	28266-28277	NNS	denotes	pseudogenes
T7724	28277-28278	.	denotes	.
T7725	28278-28364	sentence	denotes	No public sequence matches the 12th 'expressed pseudogene' with 99% identity or more.
T7726	28279-28281	DT	denotes	No
T7728	28282-28288	JJ	denotes	public
T7727	28289-28297	NN	denotes	sequence
T7729	28298-28305	VBZ	denotes	matches
T7730	28306-28309	DT	denotes	the
T7732	28310-28314	JJ	denotes	12th
T7733	28315-28316	``	denotes	'
T7734	28316-28325	JJ	denotes	expressed
T7731	28326-28336	NN	denotes	pseudogene
T7735	28336-28337	''	denotes	'
T7736	28338-28342	IN	denotes	with
T7737	28343-28345	CD	denotes	99
T7738	28345-28346	NN	denotes	%
T7739	28347-28355	NN	denotes	identity
T7740	28356-28358	CC	denotes	or
T7741	28359-28363	JJR	denotes	more
T7742	28363-28364	.	denotes	.
T8686	28377-28379	PRP	denotes	We
T8688	28380-28384	VBP	denotes	have
T8687	28385-28395	VBN	denotes	identified
T8689	28396-28399	CC	denotes	and
T8690	28400-28409	VBN	denotes	sequenced
T8691	28410-28415	CD	denotes	1,264
T8693	28416-28423	JJ	denotes	odorant
T8694	28424-28432	NN	denotes	receptor
T8692	28433-28438	NNS	denotes	cDNAs
T8695	28439-28443	IN	denotes	from
T8696	28444-28447	CD	denotes	419
T8698	28448-28457	JJ	denotes	olfactory
T8699	28458-28466	NN	denotes	receptor
T8697	28467-28472	NNS	denotes	genes
T8700	28472-28474	,	denotes	,
T8701	28474-28484	VBG	denotes	confirming
T8702	28485-28490	PRP$	denotes	their
T8703	28491-28501	NN	denotes	expression
T8704	28502-28504	IN	denotes	in
T8705	28505-28508	DT	denotes	the
T8707	28509-28518	JJ	denotes	olfactory
T8706	28519-28529	NN	denotes	epithelium
T8708	28529-28530	.	denotes	.
T8709	28530-28802	sentence	denotes	We have thus validated the similarity-based prediction of over one-third of the intact olfactory receptor genes annotated in the mouse genome [1,2], thereby vastly increasing the proportion of the family for which experimental evidence of olfactory function is available.
T8710	28531-28533	PRP	denotes	We
T8712	28534-28538	VBP	denotes	have
T8713	28539-28543	RB	denotes	thus
T8711	28544-28553	VBN	denotes	validated
T8714	28554-28557	DT	denotes	the
T8716	28558-28568	NN	denotes	similarity
T8718	28568-28569	HYPH	denotes	-
T8717	28569-28574	VBN	denotes	based
T8715	28575-28585	NN	denotes	prediction
T8719	28586-28588	IN	denotes	of
T8720	28589-28593	IN	denotes	over
T8721	28594-28597	CD	denotes	one
T8722	28597-28598	HYPH	denotes	-
T8723	28598-28603	JJ	denotes	third
T8724	28604-28606	IN	denotes	of
T8725	28607-28610	DT	denotes	the
T8727	28611-28617	JJ	denotes	intact
T8728	28618-28627	JJ	denotes	olfactory
T8729	28628-28636	NN	denotes	receptor
T8726	28637-28642	NNS	denotes	genes
T8730	28643-28652	VBN	denotes	annotated
T8731	28653-28655	IN	denotes	in
T8732	28656-28659	DT	denotes	the
T8734	28660-28665	NN	denotes	mouse
T8733	28666-28672	NN	denotes	genome
T8735	28673-28674	-LRB-	denotes	[
T8737	28674-28675	CD	denotes	1
T8738	28675-28676	,	denotes	,
T8736	28676-28677	CD	denotes	2
T8739	28677-28678	-RRB-	denotes	]
T8740	28678-28680	,	denotes	,
T8741	28680-28687	RB	denotes	thereby
T8743	28688-28694	RB	denotes	vastly
T8742	28695-28705	VBG	denotes	increasing
T8744	28706-28709	DT	denotes	the
T8745	28710-28720	NN	denotes	proportion
T8746	28721-28723	IN	denotes	of
T8747	28724-28727	DT	denotes	the
T8748	28728-28734	NN	denotes	family
T8749	28735-28738	IN	denotes	for
T8751	28739-28744	WDT	denotes	which
T8752	28745-28757	JJ	denotes	experimental
T8753	28758-28766	NN	denotes	evidence
T8754	28767-28769	IN	denotes	of
T8755	28770-28779	JJ	denotes	olfactory
T8756	28780-28788	NN	denotes	function
T8750	28789-28791	VBZ	denotes	is
T8757	28792-28801	JJ	denotes	available
T8758	28801-28802	.	denotes	.
T8759	28802-29013	sentence	denotes	We have not found cDNAs for all olfactory receptor genes or an even phylogenetic distribution of cDNAs, probably because the libraries and/or our screen are biased toward certain olfactory receptor subfamilies.
T8760	28803-28805	PRP	denotes	We
T8762	28806-28810	VBP	denotes	have
T8763	28811-28814	RB	denotes	not
T8761	28815-28820	VBN	denotes	found
T8764	28821-28826	NNS	denotes	cDNAs
T8765	28827-28830	IN	denotes	for
T8766	28831-28834	DT	denotes	all
T8768	28835-28844	JJ	denotes	olfactory
T8769	28845-28853	NN	denotes	receptor
T8767	28854-28859	NNS	denotes	genes
T8770	28860-28862	CC	denotes	or
T8771	28863-28865	DT	denotes	an
T8773	28866-28870	RB	denotes	even
T8774	28871-28883	JJ	denotes	phylogenetic
T8772	28884-28896	NN	denotes	distribution
T8775	28897-28899	IN	denotes	of
T8776	28900-28905	NNS	denotes	cDNAs
T8777	28905-28907	,	denotes	,
T8778	28907-28915	RB	denotes	probably
T8780	28916-28923	IN	denotes	because
T8781	28924-28927	DT	denotes	the
T8782	28928-28937	NNS	denotes	libraries
T8783	28938-28941	CC	denotes	and
T8784	28941-28942	HYPH	denotes	/
T8785	28942-28944	CC	denotes	or
T8786	28945-28948	PRP$	denotes	our
T8787	28949-28955	NN	denotes	screen
T8779	28956-28959	VBP	denotes	are
T8788	28960-28966	JJ	denotes	biased
T8789	28967-28973	IN	denotes	toward
T8790	28974-28981	JJ	denotes	certain
T8792	28982-28991	JJ	denotes	olfactory
T8793	28992-29000	NN	denotes	receptor
T8791	29001-29012	NNS	denotes	subfamilies
T8794	29012-29013	.	denotes	.
T8795	29013-29359	sentence	denotes	Using RT-PCR with both degenerate and specific primers, we have confirmed olfactory expression of a number of additional olfactory receptors, bringing the total number of olfactory receptor genes verified in this study to 436, and ensuring that almost all phylogenetic clades have at least one representative with evidence of olfactory function.
T8796	29014-29019	VBG	denotes	Using
T8798	29020-29022	NN	denotes	RT
T8800	29022-29023	HYPH	denotes	-
T8799	29023-29026	NN	denotes	PCR
T8801	29027-29031	IN	denotes	with
T8802	29032-29036	CC	denotes	both
T8803	29037-29047	JJ	denotes	degenerate
T8805	29048-29051	CC	denotes	and
T8806	29052-29060	JJ	denotes	specific
T8804	29061-29068	NNS	denotes	primers
T8807	29068-29070	,	denotes	,
T8808	29070-29072	PRP	denotes	we
T8809	29073-29077	VBP	denotes	have
T8797	29078-29087	VBN	denotes	confirmed
T8810	29088-29097	JJ	denotes	olfactory
T8811	29098-29108	NN	denotes	expression
T8812	29109-29111	IN	denotes	of
T8813	29112-29113	DT	denotes	a
T8814	29114-29120	NN	denotes	number
T8815	29121-29123	IN	denotes	of
T8816	29124-29134	JJ	denotes	additional
T8818	29135-29144	JJ	denotes	olfactory
T8817	29145-29154	NNS	denotes	receptors
T8819	29154-29156	,	denotes	,
T8820	29156-29164	VBG	denotes	bringing
T8821	29165-29168	DT	denotes	the
T8823	29169-29174	JJ	denotes	total
T8822	29175-29181	NN	denotes	number
T8824	29182-29184	IN	denotes	of
T8825	29185-29194	JJ	denotes	olfactory
T8826	29195-29203	NN	denotes	receptor
T8827	29204-29209	NNS	denotes	genes
T8828	29210-29218	VBN	denotes	verified
T8829	29219-29221	IN	denotes	in
T8830	29222-29226	DT	denotes	this
T8831	29227-29232	NN	denotes	study
T8832	29233-29235	IN	denotes	to
T8833	29236-29239	CD	denotes	436
T8834	29239-29241	,	denotes	,
T8835	29241-29244	CC	denotes	and
T8836	29245-29253	VBG	denotes	ensuring
T8837	29254-29258	IN	denotes	that
T8839	29259-29265	RB	denotes	almost
T8841	29266-29269	DT	denotes	all
T8842	29270-29282	JJ	denotes	phylogenetic
T8840	29283-29289	NNS	denotes	clades
T8838	29290-29294	VBP	denotes	have
T8843	29295-29297	RB	denotes	at
T8845	29298-29303	RBS	denotes	least
T8844	29304-29307	CD	denotes	one
T8846	29308-29322	NN	denotes	representative
T8847	29323-29327	IN	denotes	with
T8848	29328-29336	NN	denotes	evidence
T8849	29337-29339	IN	denotes	of
T8850	29340-29349	JJ	denotes	olfactory
T8851	29350-29358	NN	denotes	function
T8852	29358-29359	.	denotes	.
T8853	29359-29492	sentence	denotes	Results of our cDNA library screen suggested that some olfactory receptors are expressed at significantly higher levels than others.
T8854	29360-29367	NNS	denotes	Results
T8856	29368-29370	IN	denotes	of
T8857	29371-29374	PRP$	denotes	our
T8859	29375-29379	NN	denotes	cDNA
T8860	29380-29387	NN	denotes	library
T8858	29388-29394	NN	denotes	screen
T8855	29395-29404	VBD	denotes	suggested
T8861	29405-29409	IN	denotes	that
T8863	29410-29414	DT	denotes	some
T8865	29415-29424	JJ	denotes	olfactory
T8864	29425-29434	NNS	denotes	receptors
T8866	29435-29438	VBP	denotes	are
T8862	29439-29448	VBN	denotes	expressed
T8867	29449-29451	IN	denotes	at
T8868	29452-29465	RB	denotes	significantly
T8869	29466-29472	JJR	denotes	higher
T8870	29473-29479	NNS	denotes	levels
T8871	29480-29484	IN	denotes	than
T8872	29485-29491	NNS	denotes	others
T8873	29491-29492	.	denotes	.
T8874	29492-29656	sentence	denotes	We used quantitative PCR to show that expression levels are indeed highly variable, with one olfactory receptor expressed at almost 300 times the level of another.
T8875	29493-29495	PRP	denotes	We
T8876	29496-29500	VBD	denotes	used
T8877	29501-29513	JJ	denotes	quantitative
T8878	29514-29517	NN	denotes	PCR
T8879	29518-29520	TO	denotes	to
T8880	29521-29525	VB	denotes	show
T8881	29526-29530	IN	denotes	that
T8883	29531-29541	NN	denotes	expression
T8884	29542-29548	NNS	denotes	levels
T8882	29549-29552	VBP	denotes	are
T8885	29553-29559	RB	denotes	indeed
T8886	29560-29566	RB	denotes	highly
T8887	29567-29575	JJ	denotes	variable
T8888	29575-29577	,	denotes	,
T8889	29577-29581	IN	denotes	with
T8891	29582-29585	CD	denotes	one
T8893	29586-29595	JJ	denotes	olfactory
T8892	29596-29604	NN	denotes	receptor
T8890	29605-29614	VBN	denotes	expressed
T8894	29615-29617	IN	denotes	at
T8895	29618-29624	RB	denotes	almost
T8896	29625-29628	CD	denotes	300
T8898	29629-29634	NNS	denotes	times
T8899	29635-29638	DT	denotes	the
T8897	29639-29644	NN	denotes	level
T8900	29645-29647	IN	denotes	of
T8901	29648-29655	DT	denotes	another
T8902	29655-29656	.	denotes	.
T8903	29656-29803	sentence	denotes	Higher expression levels could be due to increased transcript number per cell and/or a greater number of olfactory neurons 'choosing' those genes.
T8904	29657-29663	JJR	denotes	Higher
T8906	29664-29674	NN	denotes	expression
T8905	29675-29681	NNS	denotes	levels
T8908	29682-29687	MD	denotes	could
T8907	29688-29690	VB	denotes	be
T8909	29691-29694	IN	denotes	due
T8910	29695-29697	IN	denotes	to
T8911	29698-29707	VBN	denotes	increased
T8913	29708-29718	NN	denotes	transcript
T8912	29719-29725	NN	denotes	number
T8914	29726-29729	IN	denotes	per
T8915	29730-29734	NN	denotes	cell
T8916	29735-29738	CC	denotes	and
T8917	29738-29739	HYPH	denotes	/
T8918	29739-29741	CC	denotes	or
T8919	29742-29743	DT	denotes	a
T8921	29744-29751	JJR	denotes	greater
T8920	29752-29758	NN	denotes	number
T8923	29759-29761	IN	denotes	of
T8924	29762-29771	JJ	denotes	olfactory
T8925	29772-29779	NNS	denotes	neurons
T8926	29780-29781	``	denotes	'
T8922	29781-29789	VBG	denotes	choosing
T8927	29789-29790	''	denotes	'
T8928	29791-29796	DT	denotes	those
T8929	29797-29802	NNS	denotes	genes
T8930	29802-29803	.	denotes	.
T8931	29803-29899	sentence	denotes	For one pair of genes we tested, expression level differences appear to be due to both factors.
T8932	29804-29807	IN	denotes	For
T8934	29808-29811	CD	denotes	one
T8935	29812-29816	NN	denotes	pair
T8936	29817-29819	IN	denotes	of
T8937	29820-29825	NNS	denotes	genes
T8938	29826-29828	PRP	denotes	we
T8939	29829-29835	VBD	denotes	tested
T8940	29835-29837	,	denotes	,
T8941	29837-29847	NN	denotes	expression
T8942	29848-29853	NN	denotes	level
T8943	29854-29865	NNS	denotes	differences
T8933	29866-29872	VBP	denotes	appear
T8944	29873-29875	TO	denotes	to
T8945	29876-29878	VB	denotes	be
T8946	29879-29882	IN	denotes	due
T8947	29883-29885	IN	denotes	to
T8948	29886-29890	DT	denotes	both
T8949	29891-29898	NNS	denotes	factors
T8950	29898-29899	.	denotes	.
T8951	29899-30083	sentence	denotes	It would be interesting to collect data for additional genes to determine how the numbers of expressing cells and transcript levels per cell vary across the olfactory receptor family.
T8952	29900-29902	PRP	denotes	It
T8954	29903-29908	MD	denotes	would
T8953	29909-29911	VB	denotes	be
T8955	29912-29923	JJ	denotes	interesting
T8956	29924-29926	TO	denotes	to
T8957	29927-29934	VB	denotes	collect
T8958	29935-29939	NNS	denotes	data
T8959	29940-29943	IN	denotes	for
T8960	29944-29954	JJ	denotes	additional
T8961	29955-29960	NNS	denotes	genes
T8962	29961-29963	TO	denotes	to
T8963	29964-29973	VB	denotes	determine
T8964	29974-29977	WRB	denotes	how
T8966	29978-29981	DT	denotes	the
T8967	29982-29989	NNS	denotes	numbers
T8968	29990-29992	IN	denotes	of
T8969	29993-30003	VBG	denotes	expressing
T8970	30004-30009	NNS	denotes	cells
T8971	30010-30013	CC	denotes	and
T8972	30014-30024	NN	denotes	transcript
T8973	30025-30031	NNS	denotes	levels
T8974	30032-30035	IN	denotes	per
T8975	30036-30040	NN	denotes	cell
T8965	30041-30045	VBP	denotes	vary
T8976	30046-30052	IN	denotes	across
T8977	30053-30056	DT	denotes	the
T8979	30057-30066	JJ	denotes	olfactory
T8980	30067-30075	NN	denotes	receptor
T8978	30076-30082	NN	denotes	family
T8981	30082-30083	.	denotes	.
T8982	30083-30375	sentence	denotes	Data from a number of previous studies also show that different olfactory receptor genes, or even copies of the same olfactory receptor transgene in different genomic locations are expressed in different numbers of cells [14,18,35], but do not address the issue of transcript level per cell.
T8983	30084-30088	NNS	denotes	Data
T8985	30089-30093	IN	denotes	from
T8986	30094-30095	DT	denotes	a
T8987	30096-30102	NN	denotes	number
T8988	30103-30105	IN	denotes	of
T8989	30106-30114	JJ	denotes	previous
T8990	30115-30122	NNS	denotes	studies
T8991	30123-30127	RB	denotes	also
T8984	30128-30132	VBP	denotes	show
T8992	30133-30137	IN	denotes	that
T8994	30138-30147	JJ	denotes	different
T8996	30148-30157	JJ	denotes	olfactory
T8997	30158-30166	NN	denotes	receptor
T8995	30167-30172	NNS	denotes	genes
T8998	30172-30174	,	denotes	,
T8999	30174-30176	CC	denotes	or
T9000	30177-30181	RB	denotes	even
T9001	30182-30188	NNS	denotes	copies
T9002	30189-30191	IN	denotes	of
T9003	30192-30195	DT	denotes	the
T9005	30196-30200	JJ	denotes	same
T9006	30201-30210	JJ	denotes	olfactory
T9007	30211-30219	NN	denotes	receptor
T9004	30220-30229	NN	denotes	transgene
T9008	30230-30232	IN	denotes	in
T9009	30233-30242	JJ	denotes	different
T9011	30243-30250	JJ	denotes	genomic
T9010	30251-30260	NNS	denotes	locations
T9012	30261-30264	VBP	denotes	are
T8993	30265-30274	VBN	denotes	expressed
T9013	30275-30277	IN	denotes	in
T9014	30278-30287	JJ	denotes	different
T9015	30288-30295	NNS	denotes	numbers
T9016	30296-30298	IN	denotes	of
T9017	30299-30304	NNS	denotes	cells
T9018	30305-30306	-LRB-	denotes	[
T9020	30306-30308	CD	denotes	14
T9021	30308-30309	,	denotes	,
T9022	30309-30311	CD	denotes	18
T9023	30311-30312	,	denotes	,
T9019	30312-30314	CD	denotes	35
T9024	30314-30315	-RRB-	denotes	]
T9025	30315-30317	,	denotes	,
T9026	30317-30320	CC	denotes	but
T9027	30321-30323	VBP	denotes	do
T9029	30324-30327	RB	denotes	not
T9028	30328-30335	VB	denotes	address
T9030	30336-30339	DT	denotes	the
T9031	30340-30345	NN	denotes	issue
T9032	30346-30348	IN	denotes	of
T9033	30349-30359	NN	denotes	transcript
T9034	30360-30365	NN	denotes	level
T9035	30366-30369	IN	denotes	per
T9036	30370-30374	NN	denotes	cell
T9037	30374-30375	.	denotes	.
T9038	30375-30565	sentence	denotes	The fact that some genes are chosen more frequently, and when chosen may be expressed at higher levels per cell, is intriguing given each olfactory neuron's single-allele expression regime.
T9039	30376-30379	DT	denotes	The
T9040	30380-30384	NN	denotes	fact
T9042	30385-30389	IN	denotes	that
T9044	30390-30394	DT	denotes	some
T9045	30395-30400	NNS	denotes	genes
T9046	30401-30404	VBP	denotes	are
T9043	30405-30411	VBN	denotes	chosen
T9047	30412-30416	RBR	denotes	more
T9048	30417-30427	RB	denotes	frequently
T9049	30427-30429	,	denotes	,
T9050	30429-30432	CC	denotes	and
T9051	30433-30437	WRB	denotes	when
T9052	30438-30444	VBN	denotes	chosen
T9054	30445-30448	MD	denotes	may
T9055	30449-30451	VB	denotes	be
T9053	30452-30461	VBN	denotes	expressed
T9056	30462-30464	IN	denotes	at
T9057	30465-30471	JJR	denotes	higher
T9058	30472-30478	NNS	denotes	levels
T9059	30479-30482	IN	denotes	per
T9060	30483-30487	NN	denotes	cell
T9061	30487-30489	,	denotes	,
T9041	30489-30491	VBZ	denotes	is
T9062	30492-30502	JJ	denotes	intriguing
T9063	30503-30508	VBN	denotes	given
T9064	30509-30513	DT	denotes	each
T9066	30514-30523	JJ	denotes	olfactory
T9065	30524-30530	NN	denotes	neuron
T9068	30530-30532	POS	denotes	's
T9069	30533-30539	JJ	denotes	single
T9071	30539-30540	HYPH	denotes	-
T9070	30540-30546	NN	denotes	allele
T9072	30547-30557	NN	denotes	expression
T9067	30558-30564	NN	denotes	regime
T9073	30564-30565	.	denotes	.
T9074	30565-30635	sentence	denotes	The observation of unequal expression leads to a number of questions.
T9075	30566-30569	DT	denotes	The
T9076	30570-30581	NN	denotes	observation
T9078	30582-30584	IN	denotes	of
T9079	30585-30592	JJ	denotes	unequal
T9080	30593-30603	NN	denotes	expression
T9077	30604-30609	VBZ	denotes	leads
T9081	30610-30612	IN	denotes	to
T9082	30613-30614	DT	denotes	a
T9083	30615-30621	NN	denotes	number
T9084	30622-30624	IN	denotes	of
T9085	30625-30634	NNS	denotes	questions
T9086	30634-30635	.	denotes	.
T9087	30635-30887	sentence	denotes	It is known that each olfactory receptor is expressed in one of four zones of the olfactory epithelium [14,15]; do some zones choose from a smaller olfactory receptor sub-repertoire and thus express each olfactory receptor in a larger number of cells?
T9088	30636-30638	PRP	denotes	It
T9090	30639-30641	VBZ	denotes	is
T9089	30642-30647	VBN	denotes	known
T9092	30648-30652	IN	denotes	that
T9094	30653-30657	DT	denotes	each
T9096	30658-30667	JJ	denotes	olfactory
T9095	30668-30676	NN	denotes	receptor
T9097	30677-30679	VBZ	denotes	is
T9093	30680-30689	VBN	denotes	expressed
T9098	30690-30692	IN	denotes	in
T9099	30693-30696	CD	denotes	one
T9100	30697-30699	IN	denotes	of
T9101	30700-30704	CD	denotes	four
T9102	30705-30710	NNS	denotes	zones
T9103	30711-30713	IN	denotes	of
T9104	30714-30717	DT	denotes	the
T9106	30718-30727	JJ	denotes	olfactory
T9105	30728-30738	NN	denotes	epithelium
T9107	30739-30740	-LRB-	denotes	[
T9109	30740-30742	CD	denotes	14
T9110	30742-30743	,	denotes	,
T9108	30743-30745	CD	denotes	15
T9111	30745-30746	-RRB-	denotes	]
T9112	30746-30747	:	denotes	;
T9113	30748-30750	VB	denotes	do
T9114	30751-30755	DT	denotes	some
T9115	30756-30761	NNS	denotes	zones
T9091	30762-30768	VBP	denotes	choose
T9116	30769-30773	IN	denotes	from
T9117	30774-30775	DT	denotes	a
T9119	30776-30783	JJR	denotes	smaller
T9120	30784-30793	JJ	denotes	olfactory
T9121	30794-30802	NN	denotes	receptor
T9118	30803-30817	NN	denotes	sub-repertoire
T9122	30818-30821	CC	denotes	and
T9123	30822-30826	RB	denotes	thus
T9124	30827-30834	VB	denotes	express
T9125	30835-30839	DT	denotes	each
T9127	30840-30849	JJ	denotes	olfactory
T9126	30850-30858	NN	denotes	receptor
T9128	30859-30861	IN	denotes	in
T9129	30862-30863	DT	denotes	a
T9131	30864-30870	JJR	denotes	larger
T9130	30871-30877	NN	denotes	number
T9132	30878-30880	IN	denotes	of
T9133	30881-30886	NNS	denotes	cells
T9134	30886-30887	.	denotes	?
T9135	30887-31056	sentence	denotes	We note that several apparently highly expressed olfactory receptors (gene A, this study, and MOR10 and MOR28 [36]) are expressed in zone 4 of the olfactory epithelium.
T9136	30888-30890	PRP	denotes	We
T9137	30891-30895	VBP	denotes	note
T9138	30896-30900	IN	denotes	that
T9140	30901-30908	JJ	denotes	several
T9142	30909-30919	RB	denotes	apparently
T9144	30920-30926	RB	denotes	highly
T9143	30927-30936	VBN	denotes	expressed
T9145	30937-30946	JJ	denotes	olfactory
T9141	30947-30956	NNS	denotes	receptors
T9146	30957-30958	-LRB-	denotes	(
T9147	30958-30962	NN	denotes	gene
T9148	30963-30964	NN	denotes	A
T9149	30964-30966	,	denotes	,
T9151	30966-30970	DT	denotes	this
T9150	30971-30976	NN	denotes	study
T9152	30976-30978	,	denotes	,
T9153	30978-30981	CC	denotes	and
T9154	30982-30987	NN	denotes	MOR10
T9155	30988-30991	CC	denotes	and
T9156	30992-30997	NN	denotes	MOR28
T9157	30998-30999	-LRB-	denotes	[
T9158	30999-31001	CD	denotes	36
T9159	31001-31002	-RRB-	denotes	]
T9160	31002-31003	-RRB-	denotes	)
T9161	31004-31007	VBP	denotes	are
T9139	31008-31017	VBN	denotes	expressed
T9162	31018-31020	IN	denotes	in
T9163	31021-31025	NN	denotes	zone
T9164	31026-31027	CD	denotes	4
T9165	31028-31030	IN	denotes	of
T9166	31031-31034	DT	denotes	the
T9168	31035-31044	JJ	denotes	olfactory
T9167	31045-31055	NN	denotes	epithelium
T9169	31055-31056	.	denotes	.
T9170	31056-31219	sentence	denotes	Does activity-dependent neuronal competition [37] contribute to increased representation of the olfactory receptors that respond to common environmental odorants?
T9171	31057-31061	VBZ	denotes	Does
T9173	31062-31070	NN	denotes	activity
T9175	31070-31071	HYPH	denotes	-
T9174	31071-31080	JJ	denotes	dependent
T9177	31081-31089	JJ	denotes	neuronal
T9176	31090-31101	NN	denotes	competition
T9178	31102-31103	-LRB-	denotes	[
T9179	31103-31105	CD	denotes	37
T9180	31105-31106	-RRB-	denotes	]
T9172	31107-31117	VB	denotes	contribute
T9181	31118-31120	IN	denotes	to
T9182	31121-31130	VBN	denotes	increased
T9183	31131-31145	NN	denotes	representation
T9184	31146-31148	IN	denotes	of
T9185	31149-31152	DT	denotes	the
T9187	31153-31162	JJ	denotes	olfactory
T9186	31163-31172	NNS	denotes	receptors
T9188	31173-31177	WDT	denotes	that
T9189	31178-31185	VBP	denotes	respond
T9190	31186-31188	IN	denotes	to
T9191	31189-31195	JJ	denotes	common
T9193	31196-31209	JJ	denotes	environmental
T9192	31210-31218	NNS	denotes	odorants
T9194	31218-31219	.	denotes	?
T9195	31219-31288	sentence	denotes	Do the favored olfactory receptors have stronger promoter sequences?
T9196	31220-31222	VBP	denotes	Do
T9198	31223-31226	DT	denotes	the
T9200	31227-31234	JJ	denotes	favored
T9201	31235-31244	JJ	denotes	olfactory
T9199	31245-31254	NNS	denotes	receptors
T9197	31255-31259	VB	denotes	have
T9202	31260-31268	JJR	denotes	stronger
T9204	31269-31277	NN	denotes	promoter
T9203	31278-31287	NNS	denotes	sequences
T9205	31287-31288	.	denotes	?
T9206	31288-31404	sentence	denotes	Are some olfactory receptor mRNAs more stable than others, leading to higher transcript levels per expressing cell?
T9207	31289-31292	VBP	denotes	Are
T9208	31293-31297	DT	denotes	some
T9210	31298-31307	JJ	denotes	olfactory
T9211	31308-31316	NN	denotes	receptor
T9209	31317-31322	NNS	denotes	mRNAs
T9212	31323-31327	RBR	denotes	more
T9213	31328-31334	JJ	denotes	stable
T9214	31335-31339	IN	denotes	than
T9215	31340-31346	NNS	denotes	others
T9216	31346-31348	,	denotes	,
T9217	31348-31355	VBG	denotes	leading
T9218	31356-31358	IN	denotes	to
T9219	31359-31365	JJR	denotes	higher
T9221	31366-31376	NN	denotes	transcript
T9220	31377-31383	NNS	denotes	levels
T9222	31384-31387	IN	denotes	per
T9223	31388-31398	VBG	denotes	expressing
T9224	31399-31403	NN	denotes	cell
T9225	31403-31404	.	denotes	?
T9226	31404-31514	sentence	denotes	Are the favored olfactory receptors in more open chromatin conformation or more accessible genomic locations?
T9227	31405-31408	VBP	denotes	Are
T9228	31409-31412	DT	denotes	the
T9230	31413-31420	JJ	denotes	favored
T9231	31421-31430	JJ	denotes	olfactory
T9229	31431-31440	NNS	denotes	receptors
T9232	31441-31443	IN	denotes	in
T9233	31444-31448	RBR	denotes	more
T9234	31449-31453	JJ	denotes	open
T9236	31454-31463	NN	denotes	chromatin
T9235	31464-31476	NN	denotes	conformation
T9237	31477-31479	CC	denotes	or
T9238	31480-31484	RBR	denotes	more
T9239	31485-31495	JJ	denotes	accessible
T9241	31496-31503	JJ	denotes	genomic
T9240	31504-31513	NNS	denotes	locations
T9242	31513-31514	.	denotes	?
T9243	31514-31918	sentence	denotes	Transcription of apparent 'singleton' olfactory receptor genes (0.5 Mb or more from the nearest other olfactory receptor gene) suggests that there is no absolute requirement for genomic clustering for an olfactory receptor to be transcribed, consistent with observations that small olfactory receptor transgenes can be expressed correctly when integrated outside native olfactory receptor clusters [35].
T9244	31515-31528	NN	denotes	Transcription
T9246	31529-31531	IN	denotes	of
T9247	31532-31540	JJ	denotes	apparent
T9249	31541-31542	``	denotes	'
T9250	31542-31551	NN	denotes	singleton
T9251	31551-31552	''	denotes	'
T9252	31553-31562	JJ	denotes	olfactory
T9253	31563-31571	NN	denotes	receptor
T9248	31572-31577	NNS	denotes	genes
T9254	31578-31579	-LRB-	denotes	(
T9256	31579-31582	CD	denotes	0.5
T9255	31583-31585	NNS	denotes	Mb
T9257	31586-31588	CC	denotes	or
T9258	31589-31593	JJR	denotes	more
T9259	31594-31598	IN	denotes	from
T9260	31599-31602	DT	denotes	the
T9262	31603-31610	JJS	denotes	nearest
T9263	31611-31616	JJ	denotes	other
T9264	31617-31626	JJ	denotes	olfactory
T9265	31627-31635	NN	denotes	receptor
T9261	31636-31640	NN	denotes	gene
T9266	31640-31641	-RRB-	denotes	)
T9245	31642-31650	VBZ	denotes	suggests
T9267	31651-31655	IN	denotes	that
T9269	31656-31661	EX	denotes	there
T9268	31662-31664	VBZ	denotes	is
T9270	31665-31667	DT	denotes	no
T9272	31668-31676	JJ	denotes	absolute
T9271	31677-31688	NN	denotes	requirement
T9273	31689-31692	IN	denotes	for
T9274	31693-31700	JJ	denotes	genomic
T9275	31701-31711	NN	denotes	clustering
T9276	31712-31715	IN	denotes	for
T9278	31716-31718	DT	denotes	an
T9280	31719-31728	JJ	denotes	olfactory
T9279	31729-31737	NN	denotes	receptor
T9281	31738-31740	TO	denotes	to
T9282	31741-31743	VB	denotes	be
T9277	31744-31755	VBN	denotes	transcribed
T9283	31755-31757	,	denotes	,
T9284	31757-31767	JJ	denotes	consistent
T9285	31768-31772	IN	denotes	with
T9286	31773-31785	NNS	denotes	observations
T9287	31786-31790	IN	denotes	that
T9289	31791-31796	JJ	denotes	small
T9291	31797-31806	JJ	denotes	olfactory
T9292	31807-31815	NN	denotes	receptor
T9290	31816-31826	NNS	denotes	transgenes
T9293	31827-31830	MD	denotes	can
T9294	31831-31833	VB	denotes	be
T9288	31834-31843	VBN	denotes	expressed
T9295	31844-31853	RB	denotes	correctly
T9296	31854-31858	WRB	denotes	when
T9297	31859-31869	VBN	denotes	integrated
T9298	31870-31877	IN	denotes	outside
T9299	31878-31884	JJ	denotes	native
T9301	31885-31894	JJ	denotes	olfactory
T9302	31895-31903	NN	denotes	receptor
T9300	31904-31912	NNS	denotes	clusters
T9303	31913-31914	-LRB-	denotes	[
T9304	31914-31916	CD	denotes	35
T9305	31916-31917	-RRB-	denotes	]
T9306	31917-31918	.	denotes	.
T9307	31918-32187	sentence	denotes	However, the high pseudogene count among singleton olfactory receptor genes (50%, versus 20% for clustered olfactory receptor genes) suggests that not all genomic locations are favorable for olfactory receptor gene survival, perhaps due to transcriptional constraints.
T9308	31919-31926	RB	denotes	However
T9310	31926-31928	,	denotes	,
T9311	31928-31931	DT	denotes	the
T9313	31932-31936	JJ	denotes	high
T9314	31937-31947	NN	denotes	pseudogene
T9312	31948-31953	NN	denotes	count
T9315	31954-31959	IN	denotes	among
T9316	31960-31969	NN	denotes	singleton
T9318	31970-31979	JJ	denotes	olfactory
T9319	31980-31988	NN	denotes	receptor
T9317	31989-31994	NNS	denotes	genes
T9320	31995-31996	-LRB-	denotes	(
T9321	31996-31998	CD	denotes	50
T9322	31998-31999	NN	denotes	%
T9323	31999-32001	,	denotes	,
T9324	32001-32007	CC	denotes	versus
T9325	32008-32010	CD	denotes	20
T9326	32010-32011	NN	denotes	%
T9327	32012-32015	IN	denotes	for
T9328	32016-32025	VBN	denotes	clustered
T9330	32026-32035	JJ	denotes	olfactory
T9331	32036-32044	NN	denotes	receptor
T9329	32045-32050	NNS	denotes	genes
T9332	32050-32051	-RRB-	denotes	)
T9309	32052-32060	VBZ	denotes	suggests
T9333	32061-32065	IN	denotes	that
T9335	32066-32069	RB	denotes	not
T9337	32070-32073	DT	denotes	all
T9338	32074-32081	JJ	denotes	genomic
T9336	32082-32091	NNS	denotes	locations
T9334	32092-32095	VBP	denotes	are
T9339	32096-32105	JJ	denotes	favorable
T9340	32106-32109	IN	denotes	for
T9341	32110-32119	JJ	denotes	olfactory
T9342	32120-32128	NN	denotes	receptor
T9344	32129-32133	NN	denotes	gene
T9343	32134-32142	NN	denotes	survival
T9345	32142-32144	,	denotes	,
T9346	32144-32151	RB	denotes	perhaps
T9347	32152-32155	IN	denotes	due
T9348	32156-32158	IN	denotes	to
T9349	32159-32174	JJ	denotes	transcriptional
T9350	32175-32186	NNS	denotes	constraints
T9351	32186-32187	.	denotes	.
T9352	32187-32450	sentence	denotes	It is also possible that evolutionary factors may be responsible for reduced pseudogene content of clustered olfactory receptors - gene conversion between neighboring olfactory receptors could rescue inactivating mutations in clustered genes, but not singletons.
T9353	32188-32190	PRP	denotes	It
T9354	32191-32193	VBZ	denotes	is
T9356	32194-32198	RB	denotes	also
T9357	32199-32207	JJ	denotes	possible
T9358	32208-32212	IN	denotes	that
T9360	32213-32225	JJ	denotes	evolutionary
T9361	32226-32233	NNS	denotes	factors
T9362	32234-32237	MD	denotes	may
T9359	32238-32240	VB	denotes	be
T9363	32241-32252	JJ	denotes	responsible
T9364	32253-32256	IN	denotes	for
T9365	32257-32264	JJ	denotes	reduced
T9367	32265-32275	NN	denotes	pseudogene
T9366	32276-32283	NN	denotes	content
T9368	32284-32286	IN	denotes	of
T9369	32287-32296	JJ	denotes	clustered
T9371	32297-32306	JJ	denotes	olfactory
T9370	32307-32316	NNS	denotes	receptors
T9372	32317-32318	HYPH	denotes	-
T9373	32319-32323	NN	denotes	gene
T9374	32324-32334	NN	denotes	conversion
T9375	32335-32342	IN	denotes	between
T9376	32343-32354	VBG	denotes	neighboring
T9378	32355-32364	JJ	denotes	olfactory
T9377	32365-32374	NNS	denotes	receptors
T9379	32375-32380	MD	denotes	could
T9355	32381-32387	VB	denotes	rescue
T9380	32388-32400	VBG	denotes	inactivating
T9381	32401-32410	NNS	denotes	mutations
T9382	32411-32413	IN	denotes	in
T9383	32414-32423	JJ	denotes	clustered
T9384	32424-32429	NNS	denotes	genes
T9385	32429-32431	,	denotes	,
T9386	32431-32434	CC	denotes	but
T9387	32435-32438	RB	denotes	not
T9388	32439-32449	NNS	denotes	singletons
T9389	32449-32450	.	denotes	.
T9390	32450-32659	sentence	denotes	Before these questions about olfactory receptor gene choice can be answered, it will be important to measure expression levels of a larger number of genes, perhaps using an olfactory receptor gene microarray.
T9391	32451-32457	IN	denotes	Before
T9393	32458-32463	DT	denotes	these
T9394	32464-32473	NNS	denotes	questions
T9395	32474-32479	IN	denotes	about
T9396	32480-32489	JJ	denotes	olfactory
T9397	32490-32498	NN	denotes	receptor
T9398	32499-32503	NN	denotes	gene
T9399	32504-32510	NN	denotes	choice
T9400	32511-32514	MD	denotes	can
T9401	32515-32517	VB	denotes	be
T9392	32518-32526	VBN	denotes	answered
T9403	32526-32528	,	denotes	,
T9404	32528-32530	PRP	denotes	it
T9405	32531-32535	MD	denotes	will
T9402	32536-32538	VB	denotes	be
T9406	32539-32548	JJ	denotes	important
T9407	32549-32551	TO	denotes	to
T9408	32552-32559	VB	denotes	measure
T9409	32560-32570	NN	denotes	expression
T9410	32571-32577	NNS	denotes	levels
T9411	32578-32580	IN	denotes	of
T9412	32581-32582	DT	denotes	a
T9414	32583-32589	JJR	denotes	larger
T9413	32590-32596	NN	denotes	number
T9415	32597-32599	IN	denotes	of
T9416	32600-32605	NNS	denotes	genes
T9417	32605-32607	,	denotes	,
T9418	32607-32614	RB	denotes	perhaps
T9419	32615-32620	VBG	denotes	using
T9420	32621-32623	DT	denotes	an
T9422	32624-32633	JJ	denotes	olfactory
T9423	32634-32642	NN	denotes	receptor
T9424	32643-32647	NN	denotes	gene
T9421	32648-32658	NN	denotes	microarray
T9425	32658-32659	.	denotes	.
T9426	32659-32779	sentence	denotes	Our study provides at least partial data about the upstream transcript structures of over 300 olfactory receptor genes.
T9427	32660-32663	PRP$	denotes	Our
T9428	32664-32669	NN	denotes	study
T9429	32670-32678	VBZ	denotes	provides
T9430	32679-32681	RB	denotes	at
T9431	32682-32687	RBS	denotes	least
T9432	32688-32695	JJ	denotes	partial
T9433	32696-32700	NNS	denotes	data
T9434	32701-32706	IN	denotes	about
T9435	32707-32710	DT	denotes	the
T9437	32711-32719	JJ	denotes	upstream
T9438	32720-32730	NN	denotes	transcript
T9436	32731-32741	NNS	denotes	structures
T9439	32742-32744	IN	denotes	of
T9440	32745-32749	IN	denotes	over
T9441	32750-32753	CD	denotes	300
T9443	32754-32763	JJ	denotes	olfactory
T9444	32764-32772	NN	denotes	receptor
T9442	32773-32778	NNS	denotes	genes
T9445	32778-32779	.	denotes	.
T9446	32779-33005	sentence	denotes	These data provide tentative locations of a large set of promoter regions, allowing computational searches for shared sequence motifs that might be involved in the intriguing transcriptional regulation of olfactory receptors.
T9447	32780-32785	DT	denotes	These
T9448	32786-32790	NNS	denotes	data
T9449	32791-32798	VBP	denotes	provide
T9450	32799-32808	JJ	denotes	tentative
T9451	32809-32818	NNS	denotes	locations
T9452	32819-32821	IN	denotes	of
T9453	32822-32823	DT	denotes	a
T9455	32824-32829	JJ	denotes	large
T9454	32830-32833	NN	denotes	set
T9456	32834-32836	IN	denotes	of
T9457	32837-32845	NN	denotes	promoter
T9458	32846-32853	NNS	denotes	regions
T9459	32853-32855	,	denotes	,
T9460	32855-32863	VBG	denotes	allowing
T9461	32864-32877	JJ	denotes	computational
T9462	32878-32886	NNS	denotes	searches
T9463	32887-32890	IN	denotes	for
T9464	32891-32897	VBN	denotes	shared
T9466	32898-32906	NN	denotes	sequence
T9465	32907-32913	NNS	denotes	motifs
T9467	32914-32918	WDT	denotes	that
T9469	32919-32924	MD	denotes	might
T9470	32925-32927	VB	denotes	be
T9468	32928-32936	VBN	denotes	involved
T9471	32937-32939	IN	denotes	in
T9472	32940-32943	DT	denotes	the
T9474	32944-32954	JJ	denotes	intriguing
T9475	32955-32970	JJ	denotes	transcriptional
T9473	32971-32981	NN	denotes	regulation
T9476	32982-32984	IN	denotes	of
T9477	32985-32994	JJ	denotes	olfactory
T9478	32995-33004	NNS	denotes	receptors
T9479	33004-33005	.	denotes	.
T9480	33005-33123	sentence	denotes	However, given that not all cDNAs are full-length clones, some of these candidates will not be true promoter regions.
T9481	33006-33013	RB	denotes	However
T9483	33013-33015	,	denotes	,
T9484	33015-33020	VBN	denotes	given
T9485	33021-33025	IN	denotes	that
T9487	33026-33029	RB	denotes	not
T9489	33030-33033	DT	denotes	all
T9488	33034-33039	NNS	denotes	cDNAs
T9486	33040-33043	VBP	denotes	are
T9490	33044-33048	JJ	denotes	full
T9492	33048-33049	HYPH	denotes	-
T9491	33049-33055	NN	denotes	length
T9493	33056-33062	NNS	denotes	clones
T9494	33062-33064	,	denotes	,
T9495	33064-33068	DT	denotes	some
T9496	33069-33071	IN	denotes	of
T9497	33072-33077	DT	denotes	these
T9498	33078-33088	NNS	denotes	candidates
T9499	33089-33093	MD	denotes	will
T9500	33094-33097	RB	denotes	not
T9482	33098-33100	VB	denotes	be
T9501	33101-33105	JJ	denotes	true
T9503	33106-33114	NN	denotes	promoter
T9502	33115-33122	NNS	denotes	regions
T9504	33122-33123	.	denotes	.
T9505	33123-33302	sentence	denotes	The 5' UTR sequences we obtained will also aid in the design of experimental probes, for example, for in situ hybridizations or to immobilize on an olfactory receptor microarray.
T9506	33124-33127	DT	denotes	The
T9508	33128-33129	CD	denotes	5
T9510	33129-33130	SYM	denotes	'
T9509	33131-33134	NN	denotes	UTR
T9507	33135-33144	NNS	denotes	sequences
T9512	33145-33147	PRP	denotes	we
T9513	33148-33156	VBD	denotes	obtained
T9514	33157-33161	MD	denotes	will
T9515	33162-33166	RB	denotes	also
T9511	33167-33170	VB	denotes	aid
T9516	33171-33173	IN	denotes	in
T9517	33174-33177	DT	denotes	the
T9518	33178-33184	NN	denotes	design
T9519	33185-33187	IN	denotes	of
T9520	33188-33200	JJ	denotes	experimental
T9521	33201-33207	NNS	denotes	probes
T9522	33207-33209	,	denotes	,
T9523	33209-33212	IN	denotes	for
T9524	33213-33220	NN	denotes	example
T9525	33220-33222	,	denotes	,
T9526	33222-33225	IN	denotes	for
T9527	33226-33228	FW	denotes	in
T9528	33229-33233	FW	denotes	situ
T9529	33234-33248	NNS	denotes	hybridizations
T9530	33249-33251	CC	denotes	or
T9531	33252-33254	TO	denotes	to
T9532	33255-33265	VB	denotes	immobilize
T9533	33266-33268	IN	denotes	on
T9534	33269-33271	DT	denotes	an
T9536	33272-33281	JJ	denotes	olfactory
T9537	33282-33290	NN	denotes	receptor
T9535	33291-33301	NN	denotes	microarray
T9538	33301-33302	.	denotes	.
T9539	33302-33408	sentence	denotes	One of the challenges of such an array will be to design unique probes with which to represent each gene.
T9540	33303-33306	CD	denotes	One
T9542	33307-33309	IN	denotes	of
T9543	33310-33313	DT	denotes	the
T9544	33314-33324	NNS	denotes	challenges
T9545	33325-33327	IN	denotes	of
T9546	33328-33332	PDT	denotes	such
T9548	33333-33335	DT	denotes	an
T9547	33336-33341	NN	denotes	array
T9549	33342-33346	MD	denotes	will
T9541	33347-33349	VB	denotes	be
T9550	33350-33352	TO	denotes	to
T9551	33353-33359	VB	denotes	design
T9552	33360-33366	JJ	denotes	unique
T9553	33367-33373	NNS	denotes	probes
T9554	33374-33378	IN	denotes	with
T9556	33379-33384	WDT	denotes	which
T9557	33385-33387	TO	denotes	to
T9555	33388-33397	VB	denotes	represent
T9558	33398-33402	DT	denotes	each
T9559	33403-33407	NN	denotes	gene
T9560	33407-33408	.	denotes	.
T9561	33408-33509	sentence	denotes	Often, the coding region of olfactory receptors is highly similar between recently duplicated genes.
T9562	33409-33414	RB	denotes	Often
T9564	33414-33416	,	denotes	,
T9565	33416-33419	DT	denotes	the
T9567	33420-33426	VBG	denotes	coding
T9566	33427-33433	NN	denotes	region
T9568	33434-33436	IN	denotes	of
T9569	33437-33446	JJ	denotes	olfactory
T9570	33447-33456	NNS	denotes	receptors
T9563	33457-33459	VBZ	denotes	is
T9571	33460-33466	RB	denotes	highly
T9572	33467-33474	JJ	denotes	similar
T9573	33475-33482	IN	denotes	between
T9574	33483-33491	RB	denotes	recently
T9575	33492-33502	VBN	denotes	duplicated
T9576	33503-33508	NNS	denotes	genes
T9577	33508-33509	.	denotes	.
T9578	33509-33658	sentence	denotes	Many pairs of similar olfactory receptors show more sequence divergence in the UTRs than the protein-coding region (J.Y., unpublished observations).
T9579	33510-33514	JJ	denotes	Many
T9580	33515-33520	NNS	denotes	pairs
T9582	33521-33523	IN	denotes	of
T9583	33524-33531	JJ	denotes	similar
T9585	33532-33541	JJ	denotes	olfactory
T9584	33542-33551	NNS	denotes	receptors
T9581	33552-33556	VBP	denotes	show
T9586	33557-33561	JJR	denotes	more
T9588	33562-33570	NN	denotes	sequence
T9587	33571-33581	NN	denotes	divergence
T9589	33582-33584	IN	denotes	in
T9590	33585-33588	DT	denotes	the
T9591	33589-33593	NNS	denotes	UTRs
T9592	33594-33598	IN	denotes	than
T9593	33599-33602	DT	denotes	the
T9595	33603-33610	NN	denotes	protein
T9597	33610-33611	HYPH	denotes	-
T9596	33611-33617	VBG	denotes	coding
T9594	33618-33624	NN	denotes	region
T9598	33625-33626	-LRB-	denotes	(
T9599	33626-33630	NNP	denotes	J.Y.
T9600	33630-33632	,	denotes	,
T9601	33632-33643	JJ	denotes	unpublished
T9602	33644-33656	NNS	denotes	observations
T9603	33656-33657	-RRB-	denotes	)
T9604	33657-33658	.	denotes	.
T9605	33658-33818	sentence	denotes	The UTRs would therefore make a better choice of sequence from which to design unique oligonucleotides to distinguish closely related olfactory receptor genes.
T9606	33659-33662	DT	denotes	The
T9607	33663-33667	NNS	denotes	UTRs
T9609	33668-33673	MD	denotes	would
T9610	33674-33683	RB	denotes	therefore
T9608	33684-33688	VB	denotes	make
T9611	33689-33690	DT	denotes	a
T9613	33691-33697	JJR	denotes	better
T9612	33698-33704	NN	denotes	choice
T9614	33705-33707	IN	denotes	of
T9615	33708-33716	NN	denotes	sequence
T9616	33717-33721	IN	denotes	from
T9618	33722-33727	WDT	denotes	which
T9619	33728-33730	TO	denotes	to
T9617	33731-33737	VB	denotes	design
T9620	33738-33744	JJ	denotes	unique
T9621	33745-33761	NNS	denotes	oligonucleotides
T9622	33762-33764	TO	denotes	to
T9623	33765-33776	VB	denotes	distinguish
T9624	33777-33784	RB	denotes	closely
T9625	33785-33792	JJ	denotes	related
T9627	33793-33802	JJ	denotes	olfactory
T9628	33803-33811	NN	denotes	receptor
T9626	33812-33817	NNS	denotes	genes
T9629	33817-33818	.	denotes	.
T9630	33818-34009	sentence	denotes	Locations of these regions in genomic sequence are difficult to predict - our study provides 5' UTR sequences of 343 genes and the approximate 3' UTR length for 399 olfactory receptor genes.
T9631	33819-33828	NNS	denotes	Locations
T9633	33829-33831	IN	denotes	of
T9634	33832-33837	DT	denotes	these
T9635	33838-33845	NNS	denotes	regions
T9636	33846-33848	IN	denotes	in
T9637	33849-33856	JJ	denotes	genomic
T9638	33857-33865	NN	denotes	sequence
T9632	33866-33869	VBP	denotes	are
T9640	33870-33879	JJ	denotes	difficult
T9641	33880-33882	TO	denotes	to
T9642	33883-33890	VB	denotes	predict
T9643	33891-33892	:	denotes	-
T9644	33893-33896	PRP$	denotes	our
T9645	33897-33902	NN	denotes	study
T9639	33903-33911	VBZ	denotes	provides
T9646	33912-33913	CD	denotes	5
T9648	33913-33914	SYM	denotes	'
T9647	33915-33918	NN	denotes	UTR
T9649	33919-33928	NNS	denotes	sequences
T9650	33929-33931	IN	denotes	of
T9651	33932-33935	CD	denotes	343
T9652	33936-33941	NNS	denotes	genes
T9653	33942-33945	CC	denotes	and
T9654	33946-33949	DT	denotes	the
T9656	33950-33961	JJ	denotes	approximate
T9657	33962-33963	CD	denotes	3
T9659	33963-33964	SYM	denotes	'
T9658	33965-33968	NN	denotes	UTR
T9655	33969-33975	NN	denotes	length
T9660	33976-33979	IN	denotes	for
T9661	33980-33983	CD	denotes	399
T9663	33984-33993	JJ	denotes	olfactory
T9664	33994-34002	NN	denotes	receptor
T9662	34003-34008	NNS	denotes	genes
T9665	34008-34009	.	denotes	.
T9666	34009-34319	sentence	denotes	Probe design must also account for the multiple transcriptional isoforms observed for many olfactory receptors - depending on the question being asked, probes could be designed in shared sequence to determine the total level of all isoforms, or in unique exons to measure the level of each isoform separately.
T9667	34010-34015	NN	denotes	Probe
T9668	34016-34022	NN	denotes	design
T9670	34023-34027	MD	denotes	must
T9671	34028-34032	RB	denotes	also
T9669	34033-34040	VB	denotes	account
T9673	34041-34044	IN	denotes	for
T9674	34045-34048	DT	denotes	the
T9676	34049-34057	JJ	denotes	multiple
T9677	34058-34073	JJ	denotes	transcriptional
T9675	34074-34082	NNS	denotes	isoforms
T9678	34083-34091	VBN	denotes	observed
T9679	34092-34095	IN	denotes	for
T9680	34096-34100	JJ	denotes	many
T9682	34101-34110	JJ	denotes	olfactory
T9681	34111-34120	NNS	denotes	receptors
T9683	34121-34122	:	denotes	-
T9684	34123-34132	VBG	denotes	depending
T9685	34133-34135	IN	denotes	on
T9686	34136-34139	DT	denotes	the
T9687	34140-34148	NN	denotes	question
T9688	34149-34154	VBG	denotes	being
T9689	34155-34160	VBD	denotes	asked
T9690	34160-34162	,	denotes	,
T9691	34162-34168	NNS	denotes	probes
T9692	34169-34174	MD	denotes	could
T9693	34175-34177	VB	denotes	be
T9672	34178-34186	VBN	denotes	designed
T9694	34187-34189	IN	denotes	in
T9695	34190-34196	VBN	denotes	shared
T9696	34197-34205	NN	denotes	sequence
T9697	34206-34208	TO	denotes	to
T9698	34209-34218	VB	denotes	determine
T9699	34219-34222	DT	denotes	the
T9701	34223-34228	JJ	denotes	total
T9700	34229-34234	NN	denotes	level
T9702	34235-34237	IN	denotes	of
T9703	34238-34241	DT	denotes	all
T9704	34242-34250	NNS	denotes	isoforms
T9705	34250-34252	,	denotes	,
T9706	34252-34254	CC	denotes	or
T9707	34255-34257	IN	denotes	in
T9708	34258-34264	JJ	denotes	unique
T9709	34265-34270	NNS	denotes	exons
T9710	34271-34273	TO	denotes	to
T9711	34274-34281	VB	denotes	measure
T9712	34282-34285	DT	denotes	the
T9713	34286-34291	NN	denotes	level
T9714	34292-34294	IN	denotes	of
T9715	34295-34299	DT	denotes	each
T9716	34300-34307	NN	denotes	isoform
T9717	34308-34318	RB	denotes	separately
T9718	34318-34319	.	denotes	.
T9719	34319-34557	sentence	denotes	We find that the majority of the olfactory receptors, like most non-olfactory receptor genes [38,39], are transcribed as multiple isoforms, involving alternative splicing of 5' untranslated exons and alternate polyadenylation-site usage.
T9720	34320-34322	PRP	denotes	We
T9721	34323-34327	VBP	denotes	find
T9722	34328-34332	IN	denotes	that
T9724	34333-34336	DT	denotes	the
T9725	34337-34345	NN	denotes	majority
T9726	34346-34348	IN	denotes	of
T9727	34349-34352	DT	denotes	the
T9729	34353-34362	JJ	denotes	olfactory
T9728	34363-34372	NNS	denotes	receptors
T9730	34372-34374	,	denotes	,
T9731	34374-34378	IN	denotes	like
T9732	34379-34383	JJS	denotes	most
T9734	34384-34397	JJ	denotes	non-olfactory
T9735	34398-34406	NN	denotes	receptor
T9733	34407-34412	NNS	denotes	genes
T9736	34413-34414	-LRB-	denotes	[
T9738	34414-34416	CD	denotes	38
T9739	34416-34417	,	denotes	,
T9737	34417-34419	CD	denotes	39
T9740	34419-34420	-RRB-	denotes	]
T9741	34420-34422	,	denotes	,
T9742	34422-34425	VBP	denotes	are
T9723	34426-34437	VBN	denotes	transcribed
T9743	34438-34440	IN	denotes	as
T9744	34441-34449	JJ	denotes	multiple
T9745	34450-34458	NNS	denotes	isoforms
T9746	34458-34460	,	denotes	,
T9747	34460-34469	VBG	denotes	involving
T9748	34470-34481	JJ	denotes	alternative
T9749	34482-34490	NN	denotes	splicing
T9750	34491-34493	IN	denotes	of
T9751	34494-34495	CD	denotes	5
T9753	34495-34496	SYM	denotes	'
T9754	34497-34509	JJ	denotes	untranslated
T9752	34510-34515	NNS	denotes	exons
T9755	34516-34519	CC	denotes	and
T9756	34520-34529	JJ	denotes	alternate
T9758	34530-34545	NN	denotes	polyadenylation
T9760	34545-34546	HYPH	denotes	-
T9759	34546-34550	NN	denotes	site
T9757	34551-34556	NN	denotes	usage
T9761	34556-34557	.	denotes	.
T9762	34557-34732	sentence	denotes	The act of splicing itself may be important for efficient mRNA export from the nucleus [40] or to couple olfactory receptor coding regions with genomically distant promoters.
T9763	34558-34561	DT	denotes	The
T9764	34562-34565	NN	denotes	act
T9766	34566-34568	IN	denotes	of
T9767	34569-34577	VBG	denotes	splicing
T9768	34578-34584	PRP	denotes	itself
T9769	34585-34588	MD	denotes	may
T9765	34589-34591	VB	denotes	be
T9770	34592-34601	JJ	denotes	important
T9771	34602-34605	IN	denotes	for
T9773	34606-34615	JJ	denotes	efficient
T9775	34616-34620	NN	denotes	mRNA
T9774	34621-34627	NN	denotes	export
T9776	34628-34632	IN	denotes	from
T9777	34633-34636	DT	denotes	the
T9778	34637-34644	NN	denotes	nucleus
T9779	34645-34646	-LRB-	denotes	[
T9772	34646-34648	CD	denotes	40
T9780	34648-34649	-RRB-	denotes	]
T9781	34650-34652	CC	denotes	or
T9782	34653-34655	TO	denotes	to
T9783	34656-34662	VB	denotes	couple
T9784	34663-34672	JJ	denotes	olfactory
T9785	34673-34681	NN	denotes	receptor
T9787	34682-34688	NN	denotes	coding
T9786	34689-34696	NNS	denotes	regions
T9788	34697-34701	IN	denotes	with
T9789	34702-34713	RB	denotes	genomically
T9790	34714-34721	JJ	denotes	distant
T9791	34722-34731	NNS	denotes	promoters
T9792	34731-34732	.	denotes	.
T9793	34732-34905	sentence	denotes	The exact nature of the spliced transcript might be unimportant, such that several isoforms might be produced simply because multiple functional splice sites are available.
T9794	34733-34736	DT	denotes	The
T9796	34737-34742	JJ	denotes	exact
T9795	34743-34749	NN	denotes	nature
T9798	34750-34752	IN	denotes	of
T9799	34753-34756	DT	denotes	the
T9801	34757-34764	VBN	denotes	spliced
T9800	34765-34775	NN	denotes	transcript
T9802	34776-34781	MD	denotes	might
T9797	34782-34784	VB	denotes	be
T9803	34785-34796	JJ	denotes	unimportant
T9804	34796-34798	,	denotes	,
T9805	34798-34802	JJ	denotes	such
T9807	34803-34807	IN	denotes	that
T9808	34808-34815	JJ	denotes	several
T9809	34816-34824	NNS	denotes	isoforms
T9810	34825-34830	MD	denotes	might
T9811	34831-34833	VB	denotes	be
T9806	34834-34842	VBN	denotes	produced
T9812	34843-34849	RB	denotes	simply
T9814	34850-34857	IN	denotes	because
T9815	34858-34866	JJ	denotes	multiple
T9817	34867-34877	JJ	denotes	functional
T9818	34878-34884	NN	denotes	splice
T9816	34885-34890	NNS	denotes	sites
T9813	34891-34894	VBP	denotes	are
T9819	34895-34904	JJ	denotes	available
T9820	34904-34905	.	denotes	.
T9821	34905-35094	sentence	denotes	Alternatively, the multiplicity of transcriptional isoforms might have functional significance, as UTRs may contain signals controlling mRNA stability, localization or degradation [41,42].
T9822	34906-34919	RB	denotes	Alternatively
T9824	34919-34921	,	denotes	,
T9825	34921-34924	DT	denotes	the
T9826	34925-34937	NN	denotes	multiplicity
T9827	34938-34940	IN	denotes	of
T9828	34941-34956	JJ	denotes	transcriptional
T9829	34957-34965	NNS	denotes	isoforms
T9830	34966-34971	MD	denotes	might
T9823	34972-34976	VB	denotes	have
T9831	34977-34987	JJ	denotes	functional
T9832	34988-35000	NN	denotes	significance
T9833	35000-35002	,	denotes	,
T9834	35002-35004	IN	denotes	as
T9836	35005-35009	NNS	denotes	UTRs
T9837	35010-35013	MD	denotes	may
T9835	35014-35021	VB	denotes	contain
T9838	35022-35029	NNS	denotes	signals
T9839	35030-35041	VBG	denotes	controlling
T9840	35042-35046	NN	denotes	mRNA
T9841	35047-35056	NN	denotes	stability
T9842	35056-35058	,	denotes	,
T9843	35058-35070	NN	denotes	localization
T9844	35071-35073	CC	denotes	or
T9845	35074-35085	NN	denotes	degradation
T9846	35086-35087	-LRB-	denotes	[
T9848	35087-35089	CD	denotes	41
T9849	35089-35090	,	denotes	,
T9847	35090-35092	CD	denotes	42
T9850	35092-35093	-RRB-	denotes	]
T9851	35093-35094	.	denotes	.
T9852	35094-35226	sentence	denotes	Our study shows that about 5% of olfactory receptor transcripts do not fit the current notion of olfactory receptor gene structure.
T9853	35095-35098	PRP$	denotes	Our
T9854	35099-35104	NN	denotes	study
T9855	35105-35110	VBZ	denotes	shows
T9856	35111-35115	IN	denotes	that
T9858	35116-35121	IN	denotes	about
T9859	35122-35123	CD	denotes	5
T9860	35123-35124	NN	denotes	%
T9861	35125-35127	IN	denotes	of
T9862	35128-35137	JJ	denotes	olfactory
T9863	35138-35146	NN	denotes	receptor
T9864	35147-35158	NNS	denotes	transcripts
T9865	35159-35161	VBP	denotes	do
T9866	35162-35165	RB	denotes	not
T9857	35166-35169	VB	denotes	fit
T9867	35170-35173	DT	denotes	the
T9869	35174-35181	JJ	denotes	current
T9868	35182-35188	NN	denotes	notion
T9870	35189-35191	IN	denotes	of
T9871	35192-35201	JJ	denotes	olfactory
T9872	35202-35210	NN	denotes	receptor
T9873	35211-35215	NN	denotes	gene
T9874	35216-35225	NN	denotes	structure
T9875	35225-35226	.	denotes	.
T9876	35226-35296	sentence	denotes	Occasionally, an intron is spliced out of the 3' untranslated region.
T9877	35227-35239	RB	denotes	Occasionally
T9879	35239-35241	,	denotes	,
T9880	35241-35243	DT	denotes	an
T9881	35244-35250	NN	denotes	intron
T9882	35251-35253	VBZ	denotes	is
T9878	35254-35261	VBN	denotes	spliced
T9883	35262-35265	IN	denotes	out
T9884	35266-35268	IN	denotes	of
T9885	35269-35272	DT	denotes	the
T9887	35273-35274	CD	denotes	3
T9888	35274-35275	SYM	denotes	'
T9889	35276-35288	JJ	denotes	untranslated
T9886	35289-35295	NN	denotes	region
T9890	35295-35296	.	denotes	.
T9891	35296-35476	sentence	denotes	A number of cDNAs use splice sites within the olfactory receptor's ORF, meaning that their protein product is different from that predicted on the basis of genomic sequence alone.
T9892	35297-35298	DT	denotes	A
T9893	35299-35305	NN	denotes	number
T9895	35306-35308	IN	denotes	of
T9896	35309-35314	NNS	denotes	cDNAs
T9894	35315-35318	VBP	denotes	use
T9897	35319-35325	NN	denotes	splice
T9898	35326-35331	NNS	denotes	sites
T9899	35332-35338	IN	denotes	within
T9900	35339-35342	DT	denotes	the
T9902	35343-35352	JJ	denotes	olfactory
T9901	35353-35361	NN	denotes	receptor
T9904	35361-35363	POS	denotes	's
T9903	35364-35367	NN	denotes	ORF
T9905	35367-35369	,	denotes	,
T9906	35369-35376	VBG	denotes	meaning
T9907	35377-35381	IN	denotes	that
T9909	35382-35387	PRP$	denotes	their
T9911	35388-35395	NN	denotes	protein
T9910	35396-35403	NN	denotes	product
T9908	35404-35406	VBZ	denotes	is
T9912	35407-35416	JJ	denotes	different
T9913	35417-35421	IN	denotes	from
T9914	35422-35426	DT	denotes	that
T9915	35427-35436	VBN	denotes	predicted
T9916	35437-35439	IN	denotes	on
T9917	35440-35443	DT	denotes	the
T9918	35444-35449	NN	denotes	basis
T9919	35450-35452	IN	denotes	of
T9920	35453-35460	JJ	denotes	genomic
T9921	35461-35469	NN	denotes	sequence
T9922	35470-35475	RB	denotes	alone
T9923	35475-35476	.	denotes	.
T9924	35476-35729	sentence	denotes	In two such cases, the transcript would encode a functional olfactory receptor, with the initiating methionine and first few amino acids encoded by an upstream exon, as has been observed previously for a subtelomeric human olfactory receptor gene [25].
T9925	35477-35479	IN	denotes	In
T9927	35480-35483	CD	denotes	two
T9929	35484-35488	JJ	denotes	such
T9928	35489-35494	NNS	denotes	cases
T9930	35494-35496	,	denotes	,
T9931	35496-35499	DT	denotes	the
T9932	35500-35510	NN	denotes	transcript
T9933	35511-35516	MD	denotes	would
T9926	35517-35523	VB	denotes	encode
T9934	35524-35525	DT	denotes	a
T9936	35526-35536	JJ	denotes	functional
T9937	35537-35546	JJ	denotes	olfactory
T9935	35547-35555	NN	denotes	receptor
T9938	35555-35557	,	denotes	,
T9939	35557-35561	IN	denotes	with
T9941	35562-35565	DT	denotes	the
T9943	35566-35576	VBG	denotes	initiating
T9942	35577-35587	NN	denotes	methionine
T9944	35588-35591	CC	denotes	and
T9945	35592-35597	JJ	denotes	first
T9947	35598-35601	JJ	denotes	few
T9948	35602-35607	NN	denotes	amino
T9946	35608-35613	NNS	denotes	acids
T9940	35614-35621	VBN	denotes	encoded
T9949	35622-35624	IN	denotes	by
T9950	35625-35627	DT	denotes	an
T9952	35628-35636	JJ	denotes	upstream
T9951	35637-35641	NN	denotes	exon
T9953	35641-35643	,	denotes	,
T9954	35643-35645	IN	denotes	as
T9956	35646-35649	VBZ	denotes	has
T9957	35650-35654	VBN	denotes	been
T9955	35655-35663	VBN	denotes	observed
T9958	35664-35674	RB	denotes	previously
T9959	35675-35678	IN	denotes	for
T9960	35679-35680	DT	denotes	a
T9962	35681-35693	JJ	denotes	subtelomeric
T9963	35694-35699	JJ	denotes	human
T9964	35700-35709	JJ	denotes	olfactory
T9965	35710-35718	NN	denotes	receptor
T9961	35719-35723	NN	denotes	gene
T9966	35724-35725	-LRB-	denotes	[
T9967	35725-35727	CD	denotes	25
T9968	35727-35728	-RRB-	denotes	]
T9969	35728-35729	.	denotes	.
T9970	35729-35928	sentence	denotes	Such within-ORF splicing might increase protein-coding diversity, although, given the small number of genes involved, splicing is unlikely to significantly affect the functional receptor repertoire.
T9971	35730-35734	JJ	denotes	Such
T9973	35735-35741	IN	denotes	within
T9974	35741-35742	HYPH	denotes	-
T9975	35742-35745	NN	denotes	ORF
T9972	35746-35754	NN	denotes	splicing
T9977	35755-35760	MD	denotes	might
T9976	35761-35769	VB	denotes	increase
T9978	35770-35777	NN	denotes	protein
T9980	35777-35778	HYPH	denotes	-
T9979	35778-35784	VBG	denotes	coding
T9981	35785-35794	NN	denotes	diversity
T9982	35794-35796	,	denotes	,
T9983	35796-35804	IN	denotes	although
T9985	35804-35806	,	denotes	,
T9986	35806-35811	VBN	denotes	given
T9987	35812-35815	DT	denotes	the
T9989	35816-35821	JJ	denotes	small
T9988	35822-35828	NN	denotes	number
T9990	35829-35831	IN	denotes	of
T9991	35832-35837	NNS	denotes	genes
T9992	35838-35846	VBN	denotes	involved
T9993	35846-35848	,	denotes	,
T9994	35848-35856	NN	denotes	splicing
T9984	35857-35859	VBZ	denotes	is
T9995	35860-35868	JJ	denotes	unlikely
T9996	35869-35871	TO	denotes	to
T9998	35872-35885	RB	denotes	significantly
T9997	35886-35892	VB	denotes	affect
T9999	35893-35896	DT	denotes	the
T10001	35897-35907	JJ	denotes	functional
T10002	35908-35916	NN	denotes	receptor
T10000	35917-35927	NN	denotes	repertoire
T10003	35927-35928	.	denotes	.
T10004	35928-36147	sentence	denotes	Most of the atypical splice forms we observe appear to encode non-functional transcripts, containing frameshifts or lacking a start codon or other functional residues conserved throughout the olfactory receptor family.
T10005	35929-35933	JJS	denotes	Most
T10007	35934-35936	IN	denotes	of
T10008	35937-35940	DT	denotes	the
T10010	35941-35949	JJ	denotes	atypical
T10011	35950-35956	NN	denotes	splice
T10009	35957-35962	NNS	denotes	forms
T10012	35963-35965	PRP	denotes	we
T10013	35966-35973	VBP	denotes	observe
T10006	35974-35980	VBP	denotes	appear
T10014	35981-35983	TO	denotes	to
T10015	35984-35990	VB	denotes	encode
T10016	35991-36005	JJ	denotes	non-functional
T10017	36006-36017	NNS	denotes	transcripts
T10018	36017-36019	,	denotes	,
T10019	36019-36029	VBG	denotes	containing
T10020	36030-36041	NNS	denotes	frameshifts
T10021	36042-36044	CC	denotes	or
T10022	36045-36052	VBG	denotes	lacking
T10023	36053-36054	DT	denotes	a
T10025	36055-36060	NN	denotes	start
T10024	36061-36066	NN	denotes	codon
T10026	36067-36069	CC	denotes	or
T10027	36070-36075	JJ	denotes	other
T10029	36076-36086	JJ	denotes	functional
T10028	36087-36095	NNS	denotes	residues
T10030	36096-36105	VBN	denotes	conserved
T10031	36106-36116	IN	denotes	throughout
T10032	36117-36120	DT	denotes	the
T10034	36121-36130	JJ	denotes	olfactory
T10035	36131-36139	NN	denotes	receptor
T10033	36140-36146	NN	denotes	family
T10036	36146-36147	.	denotes	.
T10037	36147-36310	sentence	denotes	These nonfunctional transcripts are probably aberrant by-products of the splicing system [43] that have not yet been degraded by RNA surveillance systems [40,41].
T10038	36148-36153	DT	denotes	These
T10040	36154-36167	JJ	denotes	nonfunctional
T10039	36168-36179	NNS	denotes	transcripts
T10041	36180-36183	VBP	denotes	are
T10042	36184-36192	RB	denotes	probably
T10043	36193-36201	JJ	denotes	aberrant
T10045	36202-36204	NN	denotes	by
T10046	36204-36205	HYPH	denotes	-
T10044	36205-36213	NNS	denotes	products
T10047	36214-36216	IN	denotes	of
T10048	36217-36220	DT	denotes	the
T10050	36221-36229	NN	denotes	splicing
T10049	36230-36236	NN	denotes	system
T10051	36237-36238	-LRB-	denotes	[
T10052	36238-36240	CD	denotes	43
T10053	36240-36241	-RRB-	denotes	]
T10054	36242-36246	WDT	denotes	that
T10056	36247-36251	VBP	denotes	have
T10057	36252-36255	RB	denotes	not
T10058	36256-36259	RB	denotes	yet
T10059	36260-36264	VBN	denotes	been
T10055	36265-36273	VBN	denotes	degraded
T10060	36274-36276	IN	denotes	by
T10061	36277-36280	NN	denotes	RNA
T10063	36281-36293	NN	denotes	surveillance
T10062	36294-36301	NNS	denotes	systems
T10064	36302-36303	-LRB-	denotes	[
T10066	36303-36305	CD	denotes	40
T10067	36305-36306	,	denotes	,
T10065	36306-36308	CD	denotes	41
T10068	36308-36309	-RRB-	denotes	]
T10069	36309-36310	.	denotes	.
T10070	36310-36464	sentence	denotes	The neurons expressing these aberrant transcripts might also make normal transcripts for the same genes and thus produce a functional olfactory receptor.
T10071	36311-36314	DT	denotes	The
T10072	36315-36322	NNS	denotes	neurons
T10074	36323-36333	VBG	denotes	expressing
T10075	36334-36339	DT	denotes	these
T10077	36340-36348	JJ	denotes	aberrant
T10076	36349-36360	NNS	denotes	transcripts
T10078	36361-36366	MD	denotes	might
T10079	36367-36371	RB	denotes	also
T10073	36372-36376	VB	denotes	make
T10080	36377-36383	JJ	denotes	normal
T10081	36384-36395	NNS	denotes	transcripts
T10082	36396-36399	IN	denotes	for
T10083	36400-36403	DT	denotes	the
T10085	36404-36408	JJ	denotes	same
T10084	36409-36414	NNS	denotes	genes
T10086	36415-36418	CC	denotes	and
T10087	36419-36423	RB	denotes	thus
T10088	36424-36431	VB	denotes	produce
T10089	36432-36433	DT	denotes	a
T10091	36434-36444	JJ	denotes	functional
T10092	36445-36454	JJ	denotes	olfactory
T10090	36455-36463	NN	denotes	receptor
T10093	36463-36464	.	denotes	.
T10094	36464-36753	sentence	denotes	Alternatively, the unusual transcriptional regulation of olfactory receptors might ensure that only one splice isoform is expressed per cell (unlikely, but possible if an RNA-based feedback mechanism operates), thus condemning cells expressing these aberrant isoforms to be dysfunctional.
T10095	36465-36478	RB	denotes	Alternatively
T10097	36478-36480	,	denotes	,
T10098	36480-36483	DT	denotes	the
T10100	36484-36491	JJ	denotes	unusual
T10101	36492-36507	JJ	denotes	transcriptional
T10099	36508-36518	NN	denotes	regulation
T10102	36519-36521	IN	denotes	of
T10103	36522-36531	JJ	denotes	olfactory
T10104	36532-36541	NNS	denotes	receptors
T10105	36542-36547	MD	denotes	might
T10096	36548-36554	VB	denotes	ensure
T10106	36555-36559	IN	denotes	that
T10108	36560-36564	RB	denotes	only
T10110	36565-36568	CD	denotes	one
T10111	36569-36575	NN	denotes	splice
T10109	36576-36583	NN	denotes	isoform
T10112	36584-36586	VBZ	denotes	is
T10107	36587-36596	VBN	denotes	expressed
T10113	36597-36600	IN	denotes	per
T10114	36601-36605	NN	denotes	cell
T10115	36606-36607	-LRB-	denotes	(
T10116	36607-36615	JJ	denotes	unlikely
T10117	36615-36617	,	denotes	,
T10118	36617-36620	CC	denotes	but
T10119	36621-36629	JJ	denotes	possible
T10120	36630-36632	IN	denotes	if
T10122	36633-36635	DT	denotes	an
T10124	36636-36639	NN	denotes	RNA
T10126	36639-36640	HYPH	denotes	-
T10125	36640-36645	VBN	denotes	based
T10127	36646-36654	NN	denotes	feedback
T10123	36655-36664	NN	denotes	mechanism
T10121	36665-36673	VBZ	denotes	operates
T10128	36673-36674	-RRB-	denotes	)
T10129	36674-36676	,	denotes	,
T10130	36676-36680	RB	denotes	thus
T10131	36681-36691	VBG	denotes	condemning
T10132	36692-36697	NNS	denotes	cells
T10134	36698-36708	VBG	denotes	expressing
T10135	36709-36714	DT	denotes	these
T10137	36715-36723	JJ	denotes	aberrant
T10136	36724-36732	NNS	denotes	isoforms
T10138	36733-36735	TO	denotes	to
T10133	36736-36738	VB	denotes	be
T10139	36739-36752	JJ	denotes	dysfunctional
T10140	36752-36753	.	denotes	.
T10141	36753-36916	sentence	denotes	We also observe transcripts from a small number of olfactory receptor pseudogenes, as previously described for three human olfactory receptor pseudogenes [26,44].
T10142	36754-36756	PRP	denotes	We
T10144	36757-36761	RB	denotes	also
T10143	36762-36769	VBP	denotes	observe
T10145	36770-36781	NNS	denotes	transcripts
T10146	36782-36786	IN	denotes	from
T10147	36787-36788	DT	denotes	a
T10149	36789-36794	JJ	denotes	small
T10148	36795-36801	NN	denotes	number
T10150	36802-36804	IN	denotes	of
T10151	36805-36814	JJ	denotes	olfactory
T10152	36815-36823	NN	denotes	receptor
T10153	36824-36835	NNS	denotes	pseudogenes
T10154	36835-36837	,	denotes	,
T10155	36837-36839	IN	denotes	as
T10157	36840-36850	RB	denotes	previously
T10156	36851-36860	VBN	denotes	described
T10158	36861-36864	IN	denotes	for
T10159	36865-36870	CD	denotes	three
T10161	36871-36876	JJ	denotes	human
T10162	36877-36886	JJ	denotes	olfactory
T10163	36887-36895	NN	denotes	receptor
T10160	36896-36907	NNS	denotes	pseudogenes
T10164	36908-36909	-LRB-	denotes	[
T10166	36909-36911	CD	denotes	26
T10167	36911-36912	,	denotes	,
T10165	36912-36914	CD	denotes	44
T10168	36914-36915	-RRB-	denotes	]
T10169	36915-36916	.	denotes	.
T10170	36916-37190	sentence	denotes	Although many fewer pseudogenes than intact genes were represented in our cDNA collection, some neurons in the olfactory epithelium evidently express disrupted olfactory receptors and thus might be unable to respond to odorants or to correctly innervate the olfactory bulb.
T10171	36917-36925	IN	denotes	Although
T10173	36926-36930	JJ	denotes	many
T10174	36931-36936	JJR	denotes	fewer
T10175	36937-36948	NNS	denotes	pseudogenes
T10176	36949-36953	IN	denotes	than
T10177	36954-36960	JJ	denotes	intact
T10178	36961-36966	NNS	denotes	genes
T10179	36967-36971	VBD	denotes	were
T10172	36972-36983	VBN	denotes	represented
T10181	36984-36986	IN	denotes	in
T10182	36987-36990	PRP$	denotes	our
T10184	36991-36995	NN	denotes	cDNA
T10183	36996-37006	NN	denotes	collection
T10185	37006-37008	,	denotes	,
T10186	37008-37012	DT	denotes	some
T10187	37013-37020	NNS	denotes	neurons
T10188	37021-37023	IN	denotes	in
T10189	37024-37027	DT	denotes	the
T10191	37028-37037	JJ	denotes	olfactory
T10190	37038-37048	NN	denotes	epithelium
T10192	37049-37058	RB	denotes	evidently
T10180	37059-37066	VBP	denotes	express
T10193	37067-37076	JJ	denotes	disrupted
T10195	37077-37086	JJ	denotes	olfactory
T10194	37087-37096	NNS	denotes	receptors
T10196	37097-37100	CC	denotes	and
T10197	37101-37105	RB	denotes	thus
T10199	37106-37111	MD	denotes	might
T10198	37112-37114	VB	denotes	be
T10200	37115-37121	JJ	denotes	unable
T10201	37122-37124	TO	denotes	to
T10202	37125-37132	VB	denotes	respond
T10203	37133-37135	IN	denotes	to
T10204	37136-37144	NNS	denotes	odorants
T10205	37145-37147	CC	denotes	or
T10206	37148-37150	TO	denotes	to
T10208	37151-37160	RB	denotes	correctly
T10207	37161-37170	VB	denotes	innervate
T10209	37171-37174	DT	denotes	the
T10211	37175-37184	JJ	denotes	olfactory
T10210	37185-37189	NN	denotes	bulb
T10212	37189-37190	.	denotes	.
T10213	37190-37462	sentence	denotes	Wang, Axel and coworkers have shown that an artificial transgenic olfactory receptor gene containing two nonsense mutations can support development of an olfactory neuron, but that pseudogene-expressing neurons fail to converge on a glomerulus in the olfactory bulb [45].
T10214	37191-37195	NNP	denotes	Wang
T10216	37195-37197	,	denotes	,
T10217	37197-37201	NNP	denotes	Axel
T10218	37202-37205	CC	denotes	and
T10219	37206-37215	NNS	denotes	coworkers
T10220	37216-37220	VBP	denotes	have
T10215	37221-37226	VBN	denotes	shown
T10221	37227-37231	IN	denotes	that
T10223	37232-37234	DT	denotes	an
T10225	37235-37245	JJ	denotes	artificial
T10226	37246-37256	JJ	denotes	transgenic
T10227	37257-37266	JJ	denotes	olfactory
T10228	37267-37275	NN	denotes	receptor
T10224	37276-37280	NN	denotes	gene
T10229	37281-37291	VBG	denotes	containing
T10230	37292-37295	CD	denotes	two
T10232	37296-37304	JJ	denotes	nonsense
T10231	37305-37314	NNS	denotes	mutations
T10233	37315-37318	MD	denotes	can
T10222	37319-37326	VB	denotes	support
T10234	37327-37338	NN	denotes	development
T10235	37339-37341	IN	denotes	of
T10236	37342-37344	DT	denotes	an
T10238	37345-37354	JJ	denotes	olfactory
T10237	37355-37361	NN	denotes	neuron
T10239	37361-37363	,	denotes	,
T10240	37363-37366	CC	denotes	but
T10241	37367-37371	IN	denotes	that
T10243	37372-37382	NN	denotes	pseudogene
T10245	37382-37383	HYPH	denotes	-
T10244	37383-37393	VBG	denotes	expressing
T10246	37394-37401	NNS	denotes	neurons
T10242	37402-37406	VBP	denotes	fail
T10247	37407-37409	TO	denotes	to
T10248	37410-37418	VB	denotes	converge
T10249	37419-37421	IN	denotes	on
T10250	37422-37423	DT	denotes	a
T10251	37424-37434	NN	denotes	glomerulus
T10252	37435-37437	IN	denotes	in
T10253	37438-37441	DT	denotes	the
T10255	37442-37451	JJ	denotes	olfactory
T10254	37452-37456	NN	denotes	bulb
T10256	37457-37458	-LRB-	denotes	[
T10257	37458-37460	CD	denotes	45
T10258	37460-37461	-RRB-	denotes	]
T10259	37461-37462	.	denotes	.
T10260	37462-37797	sentence	denotes	By analogy with an olfactory receptor deletion mutant [45], it is likely that most pseudogene-expressing neurons die or switch to express a different olfactory receptor gene, leaving a small number of pseudogene-expressing neurons in adult mice, but at greatly reduced levels compared to neurons expressing intact olfactory receptors.
T10261	37463-37465	IN	denotes	By
T10263	37466-37473	NN	denotes	analogy
T10264	37474-37478	IN	denotes	with
T10265	37479-37481	DT	denotes	an
T10267	37482-37491	JJ	denotes	olfactory
T10268	37492-37500	NN	denotes	receptor
T10269	37501-37509	NN	denotes	deletion
T10266	37510-37516	NN	denotes	mutant
T10270	37517-37518	-LRB-	denotes	[
T10271	37518-37520	CD	denotes	45
T10272	37520-37521	-RRB-	denotes	]
T10273	37521-37523	,	denotes	,
T10274	37523-37525	PRP	denotes	it
T10262	37526-37528	VBZ	denotes	is
T10275	37529-37535	JJ	denotes	likely
T10276	37536-37540	IN	denotes	that
T10278	37541-37545	JJS	denotes	most
T10280	37546-37556	NN	denotes	pseudogene
T10282	37556-37557	HYPH	denotes	-
T10281	37557-37567	VBG	denotes	expressing
T10279	37568-37575	NNS	denotes	neurons
T10277	37576-37579	VBP	denotes	die
T10283	37580-37582	CC	denotes	or
T10284	37583-37589	VBP	denotes	switch
T10285	37590-37592	TO	denotes	to
T10286	37593-37600	VB	denotes	express
T10287	37601-37602	DT	denotes	a
T10289	37603-37612	JJ	denotes	different
T10290	37613-37622	JJ	denotes	olfactory
T10291	37623-37631	NN	denotes	receptor
T10288	37632-37636	NN	denotes	gene
T10292	37636-37638	,	denotes	,
T10293	37638-37645	VBG	denotes	leaving
T10294	37646-37647	DT	denotes	a
T10296	37648-37653	JJ	denotes	small
T10295	37654-37660	NN	denotes	number
T10297	37661-37663	IN	denotes	of
T10298	37664-37674	NN	denotes	pseudogene
T10300	37674-37675	HYPH	denotes	-
T10299	37675-37685	VBG	denotes	expressing
T10301	37686-37693	NNS	denotes	neurons
T10302	37694-37696	IN	denotes	in
T10303	37697-37702	JJ	denotes	adult
T10304	37703-37707	NNS	denotes	mice
T10305	37707-37709	,	denotes	,
T10306	37709-37712	CC	denotes	but
T10307	37713-37715	IN	denotes	at
T10308	37716-37723	RB	denotes	greatly
T10309	37724-37731	VBN	denotes	reduced
T10310	37732-37738	NNS	denotes	levels
T10311	37739-37747	VBN	denotes	compared
T10312	37748-37750	IN	denotes	to
T10313	37751-37758	NNS	denotes	neurons
T10314	37759-37769	VBG	denotes	expressing
T10315	37770-37776	JJ	denotes	intact
T10317	37777-37786	JJ	denotes	olfactory
T10316	37787-37796	NNS	denotes	receptors
T10318	37796-37797	.	denotes	.
T10408	37811-37814	PRP$	denotes	Our
T10409	37815-37820	NN	denotes	study
T10411	37821-37824	VBZ	denotes	has
T10410	37825-37833	VBN	denotes	provided
T10412	37834-37836	DT	denotes	an
T10414	37837-37846	JJ	denotes	olfactory
T10415	37847-37855	NN	denotes	receptor
T10416	37856-37860	NN	denotes	cDNA
T10413	37861-37869	NN	denotes	resource
T10417	37870-37882	VBG	denotes	representing
T10418	37883-37887	IN	denotes	over
T10419	37888-37891	CD	denotes	one
T10420	37891-37892	HYPH	denotes	-
T10421	37892-37897	JJ	denotes	third
T10422	37898-37900	IN	denotes	of
T10423	37901-37904	DT	denotes	the
T10425	37905-37914	JJ	denotes	olfactory
T10426	37915-37923	NN	denotes	receptor
T10427	37924-37928	NN	denotes	gene
T10424	37929-37935	NN	denotes	family
T10428	37935-37936	.	denotes	.
T10429	37936-38035	sentence	denotes	We have thus established over 400 annotated olfactory receptor genes as having olfactory function.
T10430	37937-37939	PRP	denotes	We
T10432	37940-37944	VBP	denotes	have
T10433	37945-37949	RB	denotes	thus
T10431	37950-37961	VBN	denotes	established
T10434	37962-37966	IN	denotes	over
T10435	37967-37970	CD	denotes	400
T10437	37971-37980	JJ	denotes	annotated
T10438	37981-37990	JJ	denotes	olfactory
T10439	37991-37999	NN	denotes	receptor
T10436	38000-38005	NNS	denotes	genes
T10440	38006-38008	IN	denotes	as
T10441	38009-38015	VBG	denotes	having
T10442	38016-38025	JJ	denotes	olfactory
T10443	38026-38034	NN	denotes	function
T10444	38034-38035	.	denotes	.
T10445	38035-38169	sentence	denotes	The sequences we generated demonstrate that the majority of the olfactory receptor gene family has multiple transcriptional isoforms.
T10446	38036-38039	DT	denotes	The
T10447	38040-38049	NNS	denotes	sequences
T10449	38050-38052	PRP	denotes	we
T10450	38053-38062	VBD	denotes	generated
T10448	38063-38074	VBP	denotes	demonstrate
T10451	38075-38079	IN	denotes	that
T10453	38080-38083	DT	denotes	the
T10454	38084-38092	NN	denotes	majority
T10455	38093-38095	IN	denotes	of
T10456	38096-38099	DT	denotes	the
T10458	38100-38109	JJ	denotes	olfactory
T10459	38110-38118	NN	denotes	receptor
T10460	38119-38123	NN	denotes	gene
T10457	38124-38130	NN	denotes	family
T10452	38131-38134	VBZ	denotes	has
T10461	38135-38143	JJ	denotes	multiple
T10463	38144-38159	JJ	denotes	transcriptional
T10462	38160-38168	NNS	denotes	isoforms
T10464	38168-38169	.	denotes	.
T10465	38169-38264	sentence	denotes	Most olfactory receptor transcripts encode functional receptor proteins, with rare exceptions.
T10466	38170-38174	RBS	denotes	Most
T10468	38175-38184	JJ	denotes	olfactory
T10469	38185-38193	NN	denotes	receptor
T10467	38194-38205	NNS	denotes	transcripts
T10470	38206-38212	VBP	denotes	encode
T10471	38213-38223	JJ	denotes	functional
T10473	38224-38232	NN	denotes	receptor
T10472	38233-38241	NN	denotes	proteins
T10474	38241-38243	,	denotes	,
T10475	38243-38247	IN	denotes	with
T10476	38248-38252	JJ	denotes	rare
T10477	38253-38263	NNS	denotes	exceptions
T10478	38263-38264	.	denotes	.
T10479	38264-38476	sentence	denotes	We show that individual olfactory receptor genes can have vastly different expression levels, an intriguing finding in light of the unusual one-neuron one-gene transcriptional regime of the olfactory epithelium.
T10480	38265-38267	PRP	denotes	We
T10481	38268-38272	VBP	denotes	show
T10482	38273-38277	IN	denotes	that
T10484	38278-38288	JJ	denotes	individual
T10486	38289-38298	JJ	denotes	olfactory
T10487	38299-38307	NN	denotes	receptor
T10485	38308-38313	NNS	denotes	genes
T10488	38314-38317	MD	denotes	can
T10483	38318-38322	VB	denotes	have
T10489	38323-38329	RB	denotes	vastly
T10490	38330-38339	JJ	denotes	different
T10492	38340-38350	NN	denotes	expression
T10491	38351-38357	NNS	denotes	levels
T10493	38357-38359	,	denotes	,
T10494	38359-38361	DT	denotes	an
T10496	38362-38372	JJ	denotes	intriguing
T10495	38373-38380	NN	denotes	finding
T10497	38381-38383	IN	denotes	in
T10498	38384-38389	NN	denotes	light
T10499	38390-38392	IN	denotes	of
T10500	38393-38396	DT	denotes	the
T10502	38397-38404	JJ	denotes	unusual
T10503	38405-38408	CD	denotes	one
T10505	38408-38409	HYPH	denotes	-
T10504	38409-38415	NN	denotes	neuron
T10506	38416-38419	CD	denotes	one
T10508	38419-38420	HYPH	denotes	-
T10507	38420-38424	NN	denotes	gene
T10509	38425-38440	JJ	denotes	transcriptional
T10501	38441-38447	NN	denotes	regime
T10510	38448-38450	IN	denotes	of
T10511	38451-38454	DT	denotes	the
T10513	38455-38464	JJ	denotes	olfactory
T10512	38465-38475	NN	denotes	epithelium
T10514	38475-38476	.	denotes	.
T10515	38476-38625	sentence	denotes	Our results and the sequences we provide will facilitate future global studies of the mechanisms and dynamics of olfactory receptor gene expression.
T10516	38477-38480	PRP$	denotes	Our
T10517	38481-38488	NNS	denotes	results
T10519	38489-38492	CC	denotes	and
T10520	38493-38496	DT	denotes	the
T10521	38497-38506	NNS	denotes	sequences
T10522	38507-38509	PRP	denotes	we
T10523	38510-38517	VBP	denotes	provide
T10524	38518-38522	MD	denotes	will
T10518	38523-38533	VB	denotes	facilitate
T10525	38534-38540	JJ	denotes	future
T10527	38541-38547	JJ	denotes	global
T10526	38548-38555	NNS	denotes	studies
T10528	38556-38558	IN	denotes	of
T10529	38559-38562	DT	denotes	the
T10530	38563-38573	NNS	denotes	mechanisms
T10531	38574-38577	CC	denotes	and
T10532	38578-38586	NNS	denotes	dynamics
T10533	38587-38589	IN	denotes	of
T10534	38590-38599	JJ	denotes	olfactory
T10535	38600-38608	NN	denotes	receptor
T10536	38609-38613	NN	denotes	gene
T10537	38614-38624	NN	denotes	expression
T10538	38624-38625	.	denotes	.
T10657	38650-38664	NN	denotes	Identification
T10658	38665-38667	IN	denotes	of
T10659	38668-38677	JJ	denotes	olfactory
T10660	38678-38686	NN	denotes	receptor
T10661	38687-38692	NNS	denotes	cDNAs
T10662	38692-39016	sentence	denotes	An adult mouse cDNA library made from the olfactory epithelium of a single animal was provided by Leslie Vosshall (Rockefeller University, New York, NY, USA), and an embryonic library (made from the olfactory epithelia of several E16.5-E18.5 embryos) was provided by Tyler Cutforth (Columbia University, New York, NY, USA).
T10663	38693-38695	DT	denotes	An
T10665	38696-38701	JJ	denotes	adult
T10666	38702-38707	NN	denotes	mouse
T10667	38708-38712	NN	denotes	cDNA
T10664	38713-38720	NN	denotes	library
T10669	38721-38725	VBN	denotes	made
T10670	38726-38730	IN	denotes	from
T10671	38731-38734	DT	denotes	the
T10673	38735-38744	JJ	denotes	olfactory
T10672	38745-38755	NN	denotes	epithelium
T10674	38756-38758	IN	denotes	of
T10675	38759-38760	DT	denotes	a
T10677	38761-38767	JJ	denotes	single
T10676	38768-38774	NN	denotes	animal
T10678	38775-38778	VBD	denotes	was
T10668	38779-38787	VBN	denotes	provided
T10679	38788-38790	IN	denotes	by
T10680	38791-38797	NNP	denotes	Leslie
T10681	38798-38806	NNP	denotes	Vosshall
T10682	38807-38808	-LRB-	denotes	(
T10684	38808-38819	NNP	denotes	Rockefeller
T10683	38820-38830	NNP	denotes	University
T10685	38830-38832	,	denotes	,
T10686	38832-38835	NNP	denotes	New
T10687	38836-38840	NNP	denotes	York
T10688	38840-38842	,	denotes	,
T10689	38842-38844	NNP	denotes	NY
T10690	38844-38846	,	denotes	,
T10691	38846-38849	NNP	denotes	USA
T10692	38849-38850	-RRB-	denotes	)
T10693	38850-38852	,	denotes	,
T10694	38852-38855	CC	denotes	and
T10695	38856-38858	DT	denotes	an
T10697	38859-38868	JJ	denotes	embryonic
T10696	38869-38876	NN	denotes	library
T10699	38877-38878	-LRB-	denotes	(
T10700	38878-38882	VBN	denotes	made
T10701	38883-38887	IN	denotes	from
T10702	38888-38891	DT	denotes	the
T10704	38892-38901	JJ	denotes	olfactory
T10703	38902-38911	NNS	denotes	epithelia
T10705	38912-38914	IN	denotes	of
T10706	38915-38922	JJ	denotes	several
T10708	38923-38928	NN	denotes	E16.5
T10710	38928-38929	HYPH	denotes	-
T10709	38929-38934	NN	denotes	E18.5
T10707	38935-38942	NNS	denotes	embryos
T10711	38942-38943	-RRB-	denotes	)
T10712	38944-38947	VBD	denotes	was
T10698	38948-38956	VBN	denotes	provided
T10713	38957-38959	IN	denotes	by
T10714	38960-38965	NNP	denotes	Tyler
T10715	38966-38974	NNP	denotes	Cutforth
T10716	38975-38976	-LRB-	denotes	(
T10718	38976-38984	NNP	denotes	Columbia
T10717	38985-38995	NNP	denotes	University
T10719	38995-38997	,	denotes	,
T10720	38997-39000	NNP	denotes	New
T10721	39001-39005	NNP	denotes	York
T10722	39005-39007	,	denotes	,
T10723	39007-39009	NNP	denotes	NY
T10724	39009-39011	,	denotes	,
T10725	39011-39014	NNP	denotes	USA
T10726	39014-39015	-RRB-	denotes	)
T10727	39015-39016	.	denotes	.
T10728	39016-39139	sentence	denotes	Both libraries were oligo-dT primed and directionally cloned into the lambdaZAP-XR vector (Stratagene, La Jolla, CA, USA).
T10729	39017-39021	DT	denotes	Both
T10730	39022-39031	NNS	denotes	libraries
T10732	39032-39036	VBD	denotes	were
T10733	39037-39042	NN	denotes	oligo
T10734	39042-39043	HYPH	denotes	-
T10735	39043-39045	NN	denotes	dT
T10731	39046-39052	VBN	denotes	primed
T10736	39053-39056	CC	denotes	and
T10737	39057-39070	RB	denotes	directionally
T10738	39071-39077	VBN	denotes	cloned
T10739	39078-39082	IN	denotes	into
T10740	39083-39086	DT	denotes	the
T10742	39087-39096	NN	denotes	lambdaZAP
T10744	39096-39097	HYPH	denotes	-
T10743	39097-39099	NN	denotes	XR
T10741	39100-39106	NN	denotes	vector
T10745	39107-39108	-LRB-	denotes	(
T10746	39108-39118	NNP	denotes	Stratagene
T10747	39118-39120	,	denotes	,
T10748	39120-39122	NNP	denotes	La
T10749	39123-39128	NNP	denotes	Jolla
T10750	39128-39130	,	denotes	,
T10751	39130-39132	NNP	denotes	CA
T10752	39132-39134	,	denotes	,
T10753	39134-39137	NNP	denotes	USA
T10754	39137-39138	-RRB-	denotes	)
T10755	39138-39139	.	denotes	.
T10756	39139-39261	sentence	denotes	The adult library has a complexity of 6.5 × 106 primary clones, and the embryonic library has a complexity of 1.65 × 106.
T10757	39140-39143	DT	denotes	The
T10759	39144-39149	JJ	denotes	adult
T10758	39150-39157	NN	denotes	library
T10760	39158-39161	VBZ	denotes	has
T10761	39162-39163	DT	denotes	a
T10762	39164-39174	NN	denotes	complexity
T10763	39175-39177	IN	denotes	of
T10764	39178-39181	CD	denotes	6.5
T10766	39182-39183	SYM	denotes	×
T10765	39184-39187	CD	denotes	106
T10768	39188-39195	JJ	denotes	primary
T10767	39196-39202	NNS	denotes	clones
T10769	39202-39204	,	denotes	,
T10770	39204-39207	CC	denotes	and
T10771	39208-39211	DT	denotes	the
T10773	39212-39221	JJ	denotes	embryonic
T10772	39222-39229	NN	denotes	library
T10774	39230-39233	VBZ	denotes	has
T10775	39234-39235	DT	denotes	a
T10776	39236-39246	NN	denotes	complexity
T10777	39247-39249	IN	denotes	of
T10778	39250-39254	CD	denotes	1.65
T10780	39255-39256	SYM	denotes	×
T10779	39257-39260	CD	denotes	106
T10781	39260-39261	.	denotes	.
T10782	39261-39359	sentence	denotes	Libraries were amplified to give titers of 5 × 109 pfu/ml (adult) or 2 × 1010 pfu/ml (embryonic).
T10783	39262-39271	NNS	denotes	Libraries
T10785	39272-39276	VBD	denotes	were
T10784	39277-39286	VBN	denotes	amplified
T10786	39287-39289	TO	denotes	to
T10787	39290-39294	VB	denotes	give
T10788	39295-39301	NNS	denotes	titers
T10789	39302-39304	IN	denotes	of
T10790	39305-39306	CD	denotes	5
T10792	39307-39308	SYM	denotes	×
T10791	39309-39312	CD	denotes	109
T10793	39313-39316	NNS	denotes	pfu
T10794	39316-39317	SYM	denotes	/
T10795	39317-39319	NN	denotes	ml
T10796	39320-39321	-LRB-	denotes	(
T10797	39321-39326	JJ	denotes	adult
T10798	39326-39327	-RRB-	denotes	)
T10799	39328-39330	CC	denotes	or
T10800	39331-39332	CD	denotes	2
T10802	39333-39334	SYM	denotes	×
T10801	39335-39339	CD	denotes	1010
T10803	39340-39343	NNS	denotes	pfu
T10804	39343-39344	SYM	denotes	/
T10805	39344-39346	NN	denotes	ml
T10806	39347-39348	-LRB-	denotes	(
T10807	39348-39357	JJ	denotes	embryonic
T10808	39357-39358	-RRB-	denotes	)
T10809	39358-39359	.	denotes	.
T10810	39359-39549	sentence	denotes	Hybridization probes were made by degenerate PCR of mouse genomic DNA, in a fashion similar to those described previously [1], with primer pairs and annealing temperatures given in Table 2.
T10811	39360-39373	NN	denotes	Hybridization
T10812	39374-39380	NNS	denotes	probes
T10814	39381-39385	VBD	denotes	were
T10813	39386-39390	VBN	denotes	made
T10815	39391-39393	IN	denotes	by
T10816	39394-39404	JJ	denotes	degenerate
T10817	39405-39408	NN	denotes	PCR
T10818	39409-39411	IN	denotes	of
T10819	39412-39417	NN	denotes	mouse
T10821	39418-39425	JJ	denotes	genomic
T10820	39426-39429	NN	denotes	DNA
T10822	39429-39431	,	denotes	,
T10823	39431-39433	IN	denotes	in
T10824	39434-39435	DT	denotes	a
T10825	39436-39443	NN	denotes	fashion
T10826	39444-39451	JJ	denotes	similar
T10827	39452-39454	IN	denotes	to
T10828	39455-39460	DT	denotes	those
T10829	39461-39470	VBN	denotes	described
T10830	39471-39481	RB	denotes	previously
T10831	39482-39483	-LRB-	denotes	[
T10832	39483-39484	CD	denotes	1
T10833	39484-39485	-RRB-	denotes	]
T10834	39485-39487	,	denotes	,
T10835	39487-39491	IN	denotes	with
T10837	39492-39498	NN	denotes	primer
T10838	39499-39504	NNS	denotes	pairs
T10839	39505-39508	CC	denotes	and
T10840	39509-39518	VBG	denotes	annealing
T10841	39519-39531	NNS	denotes	temperatures
T10836	39532-39537	VBN	denotes	given
T10842	39538-39540	IN	denotes	in
T10843	39541-39546	NN	denotes	Table
T10844	39547-39548	CD	denotes	2
T10845	39548-39549	.	denotes	.
T10846	39549-39612	sentence	denotes	Low-stringency hybridization conditions were as described [1].
T10847	39550-39553	JJ	denotes	Low
T10849	39553-39554	HYPH	denotes	-
T10848	39554-39564	NN	denotes	stringency
T10851	39565-39578	NN	denotes	hybridization
T10850	39579-39589	NNS	denotes	conditions
T10852	39590-39594	VBD	denotes	were
T10853	39595-39597	IN	denotes	as
T10854	39598-39607	VBN	denotes	described
T10855	39608-39609	-LRB-	denotes	[
T10856	39609-39610	CD	denotes	1
T10857	39610-39611	-RRB-	denotes	]
T10858	39611-39612	.	denotes	.
T10859	39612-39732	sentence	denotes	Clonally-pure plaques were obtained through secondary screens using the same probe as the corresponding primary screen.
T10860	39613-39621	RB	denotes	Clonally
T10862	39621-39622	HYPH	denotes	-
T10861	39622-39626	JJ	denotes	pure
T10863	39627-39634	NNS	denotes	plaques
T10865	39635-39639	VBD	denotes	were
T10864	39640-39648	VBN	denotes	obtained
T10866	39649-39656	IN	denotes	through
T10867	39657-39666	JJ	denotes	secondary
T10868	39667-39674	NNS	denotes	screens
T10869	39675-39680	VBG	denotes	using
T10870	39681-39684	DT	denotes	the
T10872	39685-39689	JJ	denotes	same
T10871	39690-39695	NN	denotes	probe
T10873	39696-39698	IN	denotes	as
T10874	39699-39702	DT	denotes	the
T10876	39703-39716	VBG	denotes	corresponding
T10877	39717-39724	JJ	denotes	primary
T10875	39725-39731	NN	denotes	screen
T10878	39731-39732	.	denotes	.
T10879	39732-39812	sentence	denotes	PCR with vector primers (M13F/R) was performed to prepare sequencing templates.
T10880	39733-39736	NN	denotes	PCR
T10882	39737-39741	IN	denotes	with
T10883	39742-39748	NN	denotes	vector
T10884	39749-39756	NNS	denotes	primers
T10885	39757-39758	-LRB-	denotes	(
T10887	39758-39762	NN	denotes	M13F
T10888	39762-39763	HYPH	denotes	/
T10886	39763-39764	NN	denotes	R
T10889	39764-39765	-RRB-	denotes	)
T10890	39766-39769	VBD	denotes	was
T10881	39770-39779	VBN	denotes	performed
T10891	39780-39782	TO	denotes	to
T10892	39783-39790	VB	denotes	prepare
T10893	39791-39801	NN	denotes	sequencing
T10894	39802-39811	NNS	denotes	templates
T10895	39811-39812	.	denotes	.
T10896	39812-40049	sentence	denotes	cDNA size estimates were obtained by agarose gel electrophoresis, and inserts were sequenced from the 5' end using the M13R primer and big-dye terminator chemistry according to ABI's protocols (Applied Biosystems, Foster City, CA, USA).
T10897	39813-39817	NN	denotes	cDNA
T10898	39818-39822	NN	denotes	size
T10899	39823-39832	NNS	denotes	estimates
T10901	39833-39837	VBD	denotes	were
T10900	39838-39846	VBN	denotes	obtained
T10902	39847-39849	IN	denotes	by
T10903	39850-39857	NN	denotes	agarose
T10904	39858-39861	NN	denotes	gel
T10905	39862-39877	NN	denotes	electrophoresis
T10906	39877-39879	,	denotes	,
T10907	39879-39882	CC	denotes	and
T10908	39883-39890	NNS	denotes	inserts
T10910	39891-39895	VBD	denotes	were
T10909	39896-39905	VBN	denotes	sequenced
T10911	39906-39910	IN	denotes	from
T10912	39911-39914	DT	denotes	the
T10914	39915-39916	CD	denotes	5
T10915	39916-39917	SYM	denotes	'
T10913	39918-39921	NN	denotes	end
T10916	39922-39927	VBG	denotes	using
T10917	39928-39931	DT	denotes	the
T10919	39932-39936	NN	denotes	M13R
T10918	39937-39943	NN	denotes	primer
T10920	39944-39947	CC	denotes	and
T10921	39948-39951	JJ	denotes	big
T10923	39951-39952	HYPH	denotes	-
T10922	39952-39955	NN	denotes	dye
T10925	39956-39966	NN	denotes	terminator
T10924	39967-39976	NN	denotes	chemistry
T10926	39977-39986	VBG	denotes	according
T10927	39987-39989	IN	denotes	to
T10928	39990-39993	NNP	denotes	ABI
T10930	39993-39995	POS	denotes	's
T10929	39996-40005	NNS	denotes	protocols
T10931	40006-40007	-LRB-	denotes	(
T10933	40007-40014	NNP	denotes	Applied
T10932	40015-40025	NNP	denotes	Biosystems
T10934	40025-40027	,	denotes	,
T10935	40027-40033	NNP	denotes	Foster
T10936	40034-40038	NNP	denotes	City
T10937	40038-40040	,	denotes	,
T10938	40040-40042	NNP	denotes	CA
T10939	40042-40044	,	denotes	,
T10940	40044-40047	NNP	denotes	USA
T10941	40047-40048	-RRB-	denotes	)
T10942	40048-40049	.	denotes	.
T10943	40049-40211	sentence	denotes	In order to obtain 3' sequence, selected phage clones were converted to plasmid stocks following a scaled-down version of Stratagene's in vivo excision protocol.
T10944	40050-40052	IN	denotes	In
T10946	40053-40058	NN	denotes	order
T10947	40059-40061	TO	denotes	to
T10948	40062-40068	VB	denotes	obtain
T10949	40069-40070	CD	denotes	3
T10951	40070-40071	SYM	denotes	'
T10950	40072-40080	NN	denotes	sequence
T10952	40080-40082	,	denotes	,
T10953	40082-40090	VBN	denotes	selected
T10955	40091-40096	NN	denotes	phage
T10954	40097-40103	NNS	denotes	clones
T10956	40104-40108	VBD	denotes	were
T10945	40109-40118	VBN	denotes	converted
T10957	40119-40121	IN	denotes	to
T10958	40122-40129	NN	denotes	plasmid
T10959	40130-40136	NNS	denotes	stocks
T10960	40137-40146	VBG	denotes	following
T10961	40147-40148	DT	denotes	a
T10963	40149-40155	VBN	denotes	scaled
T10964	40155-40156	HYPH	denotes	-
T10965	40156-40160	RP	denotes	down
T10962	40161-40168	NN	denotes	version
T10966	40169-40171	IN	denotes	of
T10967	40172-40182	NNP	denotes	Stratagene
T10969	40182-40184	POS	denotes	's
T10970	40185-40187	FW	denotes	in
T10971	40188-40192	FW	denotes	vivo
T10972	40193-40201	NN	denotes	excision
T10968	40202-40210	NN	denotes	protocol
T10973	40210-40211	.	denotes	.
T10974	40211-40345	sentence	denotes	Plasmid DNA gave better 3'-end sequence than PCR products, which often suffered from polymerase stuttering through the poly(A) tract.
T10975	40212-40219	NN	denotes	Plasmid
T10976	40220-40223	NN	denotes	DNA
T10977	40224-40228	VBD	denotes	gave
T10978	40229-40235	JJR	denotes	better
T10980	40236-40237	CD	denotes	3
T10982	40237-40238	SYM	denotes	'
T10983	40238-40239	HYPH	denotes	-
T10981	40239-40242	NN	denotes	end
T10979	40243-40251	NN	denotes	sequence
T10984	40252-40256	IN	denotes	than
T10985	40257-40260	NN	denotes	PCR
T10986	40261-40269	NNS	denotes	products
T10987	40269-40271	,	denotes	,
T10988	40271-40276	WDT	denotes	which
T10990	40277-40282	RB	denotes	often
T10989	40283-40291	VBD	denotes	suffered
T10991	40292-40296	IN	denotes	from
T10992	40297-40307	NN	denotes	polymerase
T10993	40308-40318	VBG	denotes	stuttering
T10994	40319-40326	IN	denotes	through
T10995	40327-40330	DT	denotes	the
T10997	40331-40335	NN	denotes	poly
T10998	40335-40336	-LRB-	denotes	(
T10999	40336-40337	NN	denotes	A
T11000	40337-40338	-RRB-	denotes	)
T10996	40339-40344	NN	denotes	tract
T11001	40344-40345	.	denotes	.
T11002	40345-40496	sentence	denotes	cDNA sequences and associated information are available through dbEST (Genbank accessions CB172832-CB174569) and our olfactory receptor database [46].
T11003	40346-40350	NN	denotes	cDNA
T11004	40351-40360	NNS	denotes	sequences
T11006	40361-40364	CC	denotes	and
T11007	40365-40375	VBN	denotes	associated
T11008	40376-40387	NN	denotes	information
T11005	40388-40391	VBP	denotes	are
T11009	40392-40401	JJ	denotes	available
T11010	40402-40409	IN	denotes	through
T11011	40410-40415	NN	denotes	dbEST
T11012	40416-40417	-LRB-	denotes	(
T11014	40417-40424	NNP	denotes	Genbank
T11015	40425-40435	NNS	denotes	accessions
T11013	40436-40444	NN	denotes	CB172832
T11016	40444-40445	SYM	denotes	-
T11017	40445-40453	NN	denotes	CB174569
T11018	40453-40454	-RRB-	denotes	)
T11019	40455-40458	CC	denotes	and
T11020	40459-40462	PRP$	denotes	our
T11022	40463-40472	JJ	denotes	olfactory
T11023	40473-40481	NN	denotes	receptor
T11021	40482-40490	NN	denotes	database
T11024	40491-40492	-LRB-	denotes	[
T11025	40492-40494	CD	denotes	46
T11026	40494-40495	-RRB-	denotes	]
T11027	40495-40496	.	denotes	.
T11028	40496-40601	sentence	denotes	The updated olfactory receptor gene catalog is available through Genbank (accessions AY317244-AY318733).
T11029	40497-40500	DT	denotes	The
T11031	40501-40508	JJ	denotes	updated
T11032	40509-40518	JJ	denotes	olfactory
T11033	40519-40527	NN	denotes	receptor
T11034	40528-40532	NN	denotes	gene
T11030	40533-40540	NN	denotes	catalog
T11035	40541-40543	VBZ	denotes	is
T11036	40544-40553	JJ	denotes	available
T11037	40554-40561	IN	denotes	through
T11038	40562-40569	NNP	denotes	Genbank
T11039	40570-40571	-LRB-	denotes	(
T11041	40571-40581	NNS	denotes	accessions
T11040	40582-40590	NN	denotes	AY317244
T11042	40590-40591	SYM	denotes	-
T11043	40591-40599	NN	denotes	AY318733
T11044	40599-40600	-RRB-	denotes	)
T11045	40600-40601	.	denotes	.
T11046	40601-40686	sentence	denotes	Throughout the manuscript, genes are referred to by their Genbank accession numbers.
T11047	40602-40612	IN	denotes	Throughout
T11049	40613-40616	DT	denotes	the
T11050	40617-40627	NN	denotes	manuscript
T11051	40627-40629	,	denotes	,
T11052	40629-40634	NNS	denotes	genes
T11053	40635-40638	VBP	denotes	are
T11048	40639-40647	VBN	denotes	referred
T11054	40648-40650	IN	denotes	to
T11055	40651-40653	IN	denotes	by
T11056	40654-40659	PRP$	denotes	their
T11058	40660-40667	NNP	denotes	Genbank
T11059	40668-40677	NN	denotes	accession
T11057	40678-40685	NNS	denotes	numbers
T11060	40685-40686	.	denotes	.
T11276	40688-40696	NN	denotes	Sequence
T11277	40697-40705	NN	denotes	analysis
T11278	40705-40854	sentence	denotes	cDNA sequences were base-called and quality-trimmed using phred (trim_cutoff = 0.05) [47], and vector sequences were removed using cross_match [48].
T11279	40706-40710	NN	denotes	cDNA
T11280	40711-40720	NNS	denotes	sequences
T11282	40721-40725	VBD	denotes	were
T11283	40726-40730	NN	denotes	base
T11284	40730-40731	HYPH	denotes	-
T11281	40731-40737	VBN	denotes	called
T11285	40738-40741	CC	denotes	and
T11286	40742-40749	NN	denotes	quality
T11288	40749-40750	HYPH	denotes	-
T11287	40750-40757	VBN	denotes	trimmed
T11289	40758-40763	VBG	denotes	using
T11290	40764-40769	NN	denotes	phred
T11291	40770-40771	-LRB-	denotes	(
T11293	40771-40782	NN	denotes	trim_cutoff
T11294	40783-40784	SYM	denotes	=
T11292	40785-40789	CD	denotes	0.05
T11295	40789-40790	-RRB-	denotes	)
T11296	40791-40792	-LRB-	denotes	[
T11297	40792-40794	CD	denotes	47
T11298	40794-40795	-RRB-	denotes	]
T11299	40795-40797	,	denotes	,
T11300	40797-40800	CC	denotes	and
T11301	40801-40807	NN	denotes	vector
T11302	40808-40817	NNS	denotes	sequences
T11304	40818-40822	VBD	denotes	were
T11303	40823-40830	VBN	denotes	removed
T11305	40831-40836	VBG	denotes	using
T11306	40837-40848	NN	denotes	cross_match
T11307	40849-40850	-LRB-	denotes	[
T11308	40850-40852	CD	denotes	48
T11309	40852-40853	-RRB-	denotes	]
T11310	40853-40854	.	denotes	.
T11311	40854-40918	sentence	denotes	Any sequences of less than 50 bp after trimming were discarded.
T11312	40855-40858	DT	denotes	Any
T11313	40859-40868	NNS	denotes	sequences
T11315	40869-40871	IN	denotes	of
T11316	40872-40876	JJR	denotes	less
T11318	40877-40881	IN	denotes	than
T11317	40882-40884	CD	denotes	50
T11319	40885-40887	NNS	denotes	bp
T11320	40888-40893	IN	denotes	after
T11321	40894-40902	NN	denotes	trimming
T11322	40903-40907	VBD	denotes	were
T11314	40908-40917	VBN	denotes	discarded
T11323	40917-40918	.	denotes	.
T11324	40918-41135	sentence	denotes	3' UTR lengths were estimated by combining approximate insert sizes determined by PCR with 5' sequence data where possible (if the 5' sequence did not extend into the coding region we could not estimate 3' UTR size).
T11325	40919-40920	CD	denotes	3
T11327	40920-40921	SYM	denotes	'
T11326	40922-40925	NN	denotes	UTR
T11328	40926-40933	NNS	denotes	lengths
T11330	40934-40938	VBD	denotes	were
T11329	40939-40948	VBN	denotes	estimated
T11331	40949-40951	IN	denotes	by
T11332	40952-40961	VBG	denotes	combining
T11333	40962-40973	JJ	denotes	approximate
T11334	40974-40980	NN	denotes	insert
T11335	40981-40986	VBZ	denotes	sizes
T11336	40987-40997	VBN	denotes	determined
T11337	40998-41000	IN	denotes	by
T11338	41001-41004	NN	denotes	PCR
T11339	41005-41009	IN	denotes	with
T11340	41010-41011	CD	denotes	5
T11342	41011-41012	SYM	denotes	'
T11341	41013-41021	NN	denotes	sequence
T11343	41022-41026	NNS	denotes	data
T11344	41027-41032	WRB	denotes	where
T11345	41033-41041	JJ	denotes	possible
T11346	41042-41043	-LRB-	denotes	(
T11348	41043-41045	IN	denotes	if
T11350	41046-41049	DT	denotes	the
T11352	41050-41051	CD	denotes	5
T11353	41051-41052	SYM	denotes	'
T11351	41053-41061	NN	denotes	sequence
T11354	41062-41065	VBD	denotes	did
T11355	41066-41069	RB	denotes	not
T11349	41070-41076	VB	denotes	extend
T11356	41077-41081	IN	denotes	into
T11357	41082-41085	DT	denotes	the
T11359	41086-41092	VBG	denotes	coding
T11358	41093-41099	NN	denotes	region
T11360	41100-41102	PRP	denotes	we
T11361	41103-41108	MD	denotes	could
T11362	41109-41112	RB	denotes	not
T11347	41113-41121	VB	denotes	estimate
T11363	41122-41123	CD	denotes	3
T11365	41123-41124	SYM	denotes	'
T11364	41125-41128	NN	denotes	UTR
T11366	41129-41133	NN	denotes	size
T11367	41133-41134	-RRB-	denotes	)
T11368	41134-41135	.	denotes	.
T11369	41135-41373	sentence	denotes	We counted cDNAs from a given gene as showing alternative polyadenylation site usage if 3' UTR length estimates varied by at least 400 bp - smaller variation could be real, but may not be distinguishable from error in our size estimates.
T11370	41136-41138	PRP	denotes	We
T11371	41139-41146	VBD	denotes	counted
T11373	41147-41152	NNS	denotes	cDNAs
T11374	41153-41157	IN	denotes	from
T11375	41158-41159	DT	denotes	a
T11377	41160-41165	VBN	denotes	given
T11376	41166-41170	NN	denotes	gene
T11378	41171-41173	IN	denotes	as
T11379	41174-41181	VBG	denotes	showing
T11380	41182-41193	JJ	denotes	alternative
T11382	41194-41209	NN	denotes	polyadenylation
T11381	41210-41214	NN	denotes	site
T11383	41215-41220	NN	denotes	usage
T11384	41221-41223	IN	denotes	if
T11386	41224-41225	CD	denotes	3
T11388	41225-41226	SYM	denotes	'
T11387	41227-41230	NN	denotes	UTR
T11389	41231-41237	NN	denotes	length
T11390	41238-41247	NNS	denotes	estimates
T11385	41248-41254	VBD	denotes	varied
T11391	41255-41257	IN	denotes	by
T11392	41258-41260	RB	denotes	at
T11394	41261-41266	RBS	denotes	least
T11393	41267-41270	CD	denotes	400
T11395	41271-41273	NNS	denotes	bp
T11396	41274-41275	:	denotes	-
T11397	41276-41283	JJR	denotes	smaller
T11398	41284-41293	NN	denotes	variation
T11399	41294-41299	MD	denotes	could
T11372	41300-41302	VB	denotes	be
T11400	41303-41307	JJ	denotes	real
T11401	41307-41309	,	denotes	,
T11402	41309-41312	CC	denotes	but
T11403	41313-41316	MD	denotes	may
T11405	41317-41320	RB	denotes	not
T11404	41321-41323	VB	denotes	be
T11406	41324-41339	JJ	denotes	distinguishable
T11407	41340-41344	IN	denotes	from
T11408	41345-41350	NN	denotes	error
T11409	41351-41353	IN	denotes	in
T11410	41354-41357	PRP$	denotes	our
T11412	41358-41362	NN	denotes	size
T11411	41363-41372	NNS	denotes	estimates
T11413	41372-41373	.	denotes	.
T11414	41373-41718	sentence	denotes	To assign cDNAs to their corresponding olfactory receptor genes, we first defined a genomic 'territory' for each gene, with the following attributes: strand, start position (100 kb upstream of the start codon or 1 kb after the previous gene upstream on the same strand, whichever is closer) and end position (1 kb downstream of the stop codon).
T11415	41374-41376	TO	denotes	To
T11416	41377-41383	VB	denotes	assign
T11418	41384-41389	NNS	denotes	cDNAs
T11419	41390-41392	IN	denotes	to
T11420	41393-41398	PRP$	denotes	their
T11422	41399-41412	VBG	denotes	corresponding
T11423	41413-41422	JJ	denotes	olfactory
T11424	41423-41431	NN	denotes	receptor
T11421	41432-41437	NNS	denotes	genes
T11425	41437-41439	,	denotes	,
T11426	41439-41441	PRP	denotes	we
T11427	41442-41447	RB	denotes	first
T11417	41448-41455	VBD	denotes	defined
T11428	41456-41457	DT	denotes	a
T11430	41458-41465	JJ	denotes	genomic
T11431	41466-41467	``	denotes	'
T11429	41467-41476	NN	denotes	territory
T11432	41476-41477	''	denotes	'
T11433	41478-41481	IN	denotes	for
T11434	41482-41486	DT	denotes	each
T11435	41487-41491	NN	denotes	gene
T11436	41491-41493	,	denotes	,
T11437	41493-41497	IN	denotes	with
T11438	41498-41501	DT	denotes	the
T11440	41502-41511	VBG	denotes	following
T11439	41512-41522	NNS	denotes	attributes
T11441	41522-41524	:	denotes	:
T11442	41524-41530	NN	denotes	strand
T11443	41530-41532	,	denotes	,
T11444	41532-41537	NN	denotes	start
T11445	41538-41546	NN	denotes	position
T11446	41547-41548	-LRB-	denotes	(
T11448	41548-41551	CD	denotes	100
T11449	41552-41554	NNS	denotes	kb
T11450	41555-41563	RB	denotes	upstream
T11451	41564-41566	IN	denotes	of
T11452	41567-41570	DT	denotes	the
T11454	41571-41576	NN	denotes	start
T11453	41577-41582	NN	denotes	codon
T11455	41583-41585	CC	denotes	or
T11456	41586-41587	CD	denotes	1
T11457	41588-41590	NN	denotes	kb
T11458	41591-41596	IN	denotes	after
T11460	41597-41600	DT	denotes	the
T11462	41601-41609	JJ	denotes	previous
T11461	41610-41614	NN	denotes	gene
T11459	41615-41623	RB	denotes	upstream
T11463	41624-41626	IN	denotes	on
T11464	41627-41630	DT	denotes	the
T11466	41631-41635	JJ	denotes	same
T11465	41636-41642	NN	denotes	strand
T11467	41642-41644	,	denotes	,
T11468	41644-41653	WDT	denotes	whichever
T11447	41654-41656	VBZ	denotes	is
T11469	41657-41663	JJR	denotes	closer
T11470	41663-41664	-RRB-	denotes	)
T11471	41665-41668	CC	denotes	and
T11472	41669-41672	NN	denotes	end
T11473	41673-41681	NN	denotes	position
T11474	41682-41683	-LRB-	denotes	(
T11476	41683-41684	CD	denotes	1
T11477	41685-41687	NN	denotes	kb
T11475	41688-41698	RB	denotes	downstream
T11478	41699-41701	IN	denotes	of
T11479	41702-41705	DT	denotes	the
T11481	41706-41710	NN	denotes	stop
T11480	41711-41716	NN	denotes	codon
T11482	41716-41717	-RRB-	denotes	)
T11483	41717-41718	.	denotes	.
T11484	41718-41886	sentence	denotes	Trimmed sequences were compared with genomic sequences using sim4 [30] (settings P = 1 to remove polyA tails and N = 1 to perform an intensive search for small exons).
T11485	41719-41726	JJ	denotes	Trimmed
T11486	41727-41736	NNS	denotes	sequences
T11488	41737-41741	VBD	denotes	were
T11487	41742-41750	VBN	denotes	compared
T11489	41751-41755	IN	denotes	with
T11490	41756-41763	JJ	denotes	genomic
T11491	41764-41773	NNS	denotes	sequences
T11492	41774-41779	VBG	denotes	using
T11493	41780-41784	NN	denotes	sim4
T11494	41785-41786	-LRB-	denotes	[
T11495	41786-41788	CD	denotes	30
T11496	41788-41789	-RRB-	denotes	]
T11497	41790-41791	-LRB-	denotes	(
T11498	41791-41799	NNS	denotes	settings
T11499	41800-41801	NN	denotes	P
T11501	41802-41803	SYM	denotes	=
T11500	41804-41805	CD	denotes	1
T11502	41806-41808	TO	denotes	to
T11503	41809-41815	VB	denotes	remove
T11504	41816-41821	NN	denotes	polyA
T11505	41822-41827	NNS	denotes	tails
T11506	41828-41831	CC	denotes	and
T11507	41832-41833	NN	denotes	N
T11509	41834-41835	SYM	denotes	=
T11508	41836-41837	CD	denotes	1
T11511	41838-41840	TO	denotes	to
T11510	41841-41848	VB	denotes	perform
T11512	41849-41851	DT	denotes	an
T11514	41852-41861	JJ	denotes	intensive
T11513	41862-41868	NN	denotes	search
T11515	41869-41872	IN	denotes	for
T11516	41873-41878	JJ	denotes	small
T11517	41879-41884	NNS	denotes	exons
T11518	41884-41885	-RRB-	denotes	)
T11519	41885-41886	.	denotes	.
T11520	41886-41964	sentence	denotes	The sim4 algorithm uses splice-site consensus sequences to refine alignments.
T11521	41887-41890	DT	denotes	The
T11523	41891-41895	NN	denotes	sim4
T11522	41896-41905	NN	denotes	algorithm
T11524	41906-41910	VBZ	denotes	uses
T11525	41911-41917	NN	denotes	splice
T11527	41917-41918	HYPH	denotes	-
T11526	41918-41922	NN	denotes	site
T11529	41923-41932	NN	denotes	consensus
T11528	41933-41942	NNS	denotes	sequences
T11530	41943-41945	TO	denotes	to
T11531	41946-41952	VB	denotes	refine
T11532	41953-41963	NNS	denotes	alignments
T11533	41963-41964	.	denotes	.
T11534	41964-42032	sentence	denotes	Only matches of 96% or greater nucleotide identity were considered.
T11535	41965-41969	RB	denotes	Only
T11536	41970-41977	NNS	denotes	matches
T11538	41978-41980	IN	denotes	of
T11539	41981-41983	CD	denotes	96
T11540	41983-41984	NN	denotes	%
T11542	41985-41987	CC	denotes	or
T11543	41988-41995	JJR	denotes	greater
T11544	41996-42006	NN	denotes	nucleotide
T11541	42007-42015	NN	denotes	identity
T11545	42016-42020	VBD	denotes	were
T11537	42021-42031	VBN	denotes	considered
T11546	42031-42032	.	denotes	.
T11547	42032-42196	sentence	denotes	RepeatMasked sequences [49] were also compared to genomic sequences; cDNA:genomic sequence pairings not found in both masked and unmasked alignments were rejected.
T11548	42033-42045	JJ	denotes	RepeatMasked
T11549	42046-42055	NNS	denotes	sequences
T11551	42056-42057	-LRB-	denotes	[
T11552	42057-42059	CD	denotes	49
T11553	42059-42060	-RRB-	denotes	]
T11554	42061-42065	VBD	denotes	were
T11555	42066-42070	RB	denotes	also
T11550	42071-42079	VBN	denotes	compared
T11556	42080-42082	IN	denotes	to
T11557	42083-42090	JJ	denotes	genomic
T11558	42091-42100	NNS	denotes	sequences
T11559	42100-42101	:	denotes	;
T11560	42102-42106	NN	denotes	cDNA
T11562	42106-42107	:	denotes	:
T11563	42107-42114	JJ	denotes	genomic
T11564	42115-42123	NN	denotes	sequence
T11565	42124-42132	NNS	denotes	pairings
T11566	42133-42136	RB	denotes	not
T11567	42137-42142	VBN	denotes	found
T11568	42143-42145	IN	denotes	in
T11569	42146-42150	CC	denotes	both
T11570	42151-42157	JJ	denotes	masked
T11572	42158-42161	CC	denotes	and
T11573	42162-42170	JJ	denotes	unmasked
T11571	42171-42181	NNS	denotes	alignments
T11574	42182-42186	VBD	denotes	were
T11561	42187-42195	VBN	denotes	rejected
T11575	42195-42196	.	denotes	.
T11576	42196-42268	sentence	denotes	Coordinates from the unmasked alignment were used for further analysis.
T11577	42197-42208	NNS	denotes	Coordinates
T11579	42209-42213	IN	denotes	from
T11580	42214-42217	DT	denotes	the
T11582	42218-42226	JJ	denotes	unmasked
T11581	42227-42236	NN	denotes	alignment
T11583	42237-42241	VBD	denotes	were
T11578	42242-42246	VBN	denotes	used
T11584	42247-42250	IN	denotes	for
T11585	42251-42258	JJ	denotes	further
T11586	42259-42267	NN	denotes	analysis
T11587	42267-42268	.	denotes	.
T11588	42268-42350	sentence	denotes	Any cDNA sequence matching entirely within a territory was assigned to that gene.
T11589	42269-42272	DT	denotes	Any
T11591	42273-42277	NN	denotes	cDNA
T11590	42278-42286	NN	denotes	sequence
T11593	42287-42295	VBG	denotes	matching
T11594	42296-42304	RB	denotes	entirely
T11595	42305-42311	IN	denotes	within
T11596	42312-42313	DT	denotes	a
T11597	42314-42323	NN	denotes	territory
T11598	42324-42327	VBD	denotes	was
T11592	42328-42336	VBN	denotes	assigned
T11599	42337-42339	IN	denotes	to
T11600	42340-42344	DT	denotes	that
T11601	42345-42349	NN	denotes	gene
T11602	42349-42350	.	denotes	.
T11603	42350-42563	sentence	denotes	If a cDNA matched more than one gene territory, the best match was chosen (that is, the one with highest 'score', where score is the total of all exons' lengths multiplied by their respective percent identities).
T11604	42351-42353	IN	denotes	If
T11606	42354-42355	DT	denotes	a
T11607	42356-42360	NN	denotes	cDNA
T11605	42361-42368	VBD	denotes	matched
T11609	42369-42373	JJR	denotes	more
T11611	42374-42378	IN	denotes	than
T11610	42379-42382	CD	denotes	one
T11613	42383-42387	NN	denotes	gene
T11612	42388-42397	NN	denotes	territory
T11614	42397-42399	,	denotes	,
T11615	42399-42402	DT	denotes	the
T11617	42403-42407	JJS	denotes	best
T11616	42408-42413	NN	denotes	match
T11618	42414-42417	VBD	denotes	was
T11608	42418-42424	VBN	denotes	chosen
T11619	42425-42426	-LRB-	denotes	(
T11621	42426-42430	RB	denotes	that
T11622	42431-42433	RB	denotes	is
T11623	42433-42435	,	denotes	,
T11624	42435-42438	DT	denotes	the
T11620	42439-42442	CD	denotes	one
T11625	42443-42447	IN	denotes	with
T11626	42448-42455	JJS	denotes	highest
T11628	42456-42457	``	denotes	'
T11627	42457-42462	NN	denotes	score
T11629	42462-42463	''	denotes	'
T11630	42463-42465	,	denotes	,
T11631	42465-42470	WRB	denotes	where
T11633	42471-42476	NN	denotes	score
T11632	42477-42479	VBZ	denotes	is
T11634	42480-42483	DT	denotes	the
T11635	42484-42489	NN	denotes	total
T11636	42490-42492	IN	denotes	of
T11637	42493-42496	DT	denotes	all
T11638	42497-42502	NNS	denotes	exons
T11640	42502-42503	POS	denotes	'
T11639	42504-42511	NNS	denotes	lengths
T11641	42512-42522	VBN	denotes	multiplied
T11642	42523-42525	IN	denotes	by
T11643	42526-42531	PRP$	denotes	their
T11645	42532-42542	JJ	denotes	respective
T11646	42543-42550	NN	denotes	percent
T11644	42551-42561	NNS	denotes	identities
T11647	42561-42562	-RRB-	denotes	)
T11648	42562-42563	.	denotes	.
T11649	42563-42685	sentence	denotes	We found 27 cDNAs that spanned a larger genomic range than one gene territory and flagged them for more careful analysis.
T11650	42564-42566	PRP	denotes	We
T11652	42567-42572	VBD	denotes	found
T11653	42573-42575	CD	denotes	27
T11654	42576-42581	NNS	denotes	cDNAs
T11655	42582-42586	WDT	denotes	that
T11656	42587-42594	VBD	denotes	spanned
T11657	42595-42596	DT	denotes	a
T11659	42597-42603	JJR	denotes	larger
T11660	42604-42611	JJ	denotes	genomic
T11658	42612-42617	NN	denotes	range
T11661	42618-42622	IN	denotes	than
T11662	42623-42626	CD	denotes	one
T11664	42627-42631	NN	denotes	gene
T11663	42632-42641	NN	denotes	territory
T11665	42642-42645	CC	denotes	and
T11651	42646-42653	VBD	denotes	flagged
T11666	42654-42658	PRP	denotes	them
T11667	42659-42662	IN	denotes	for
T11668	42663-42667	RBR	denotes	more
T11669	42668-42675	JJ	denotes	careful
T11670	42676-42684	NN	denotes	analysis
T11671	42684-42685	.	denotes	.
T11672	42685-42868	sentence	denotes	Of these, six cDNAs showed unusual splicing within the 3' UTR, but the remaining 'territory violators' were found to be artifacts of the analysis process which fell into three types.
T11673	42686-42688	IN	denotes	Of
T11675	42689-42694	DT	denotes	these
T11676	42694-42696	,	denotes	,
T11677	42696-42699	CD	denotes	six
T11678	42700-42705	NNS	denotes	cDNAs
T11674	42706-42712	VBD	denotes	showed
T11679	42713-42720	JJ	denotes	unusual
T11680	42721-42729	NN	denotes	splicing
T11681	42730-42736	IN	denotes	within
T11682	42737-42740	DT	denotes	the
T11684	42741-42742	CD	denotes	3
T11685	42742-42743	SYM	denotes	'
T11683	42744-42747	NN	denotes	UTR
T11686	42747-42749	,	denotes	,
T11687	42749-42752	CC	denotes	but
T11688	42753-42756	DT	denotes	the
T11690	42757-42766	VBG	denotes	remaining
T11691	42767-42768	``	denotes	'
T11692	42768-42777	NN	denotes	territory
T11689	42778-42787	NNS	denotes	violators
T11694	42787-42788	''	denotes	'
T11695	42789-42793	VBD	denotes	were
T11693	42794-42799	VBN	denotes	found
T11696	42800-42802	TO	denotes	to
T11697	42803-42805	VB	denotes	be
T11698	42806-42815	NNS	denotes	artifacts
T11699	42816-42818	IN	denotes	of
T11700	42819-42822	DT	denotes	the
T11702	42823-42831	NN	denotes	analysis
T11701	42832-42839	NN	denotes	process
T11703	42840-42845	WDT	denotes	which
T11704	42846-42850	VBD	denotes	fell
T11705	42851-42855	IN	denotes	into
T11706	42856-42861	CD	denotes	three
T11707	42862-42867	NNS	denotes	types
T11708	42867-42868	.	denotes	.
T11709	42868-43255	sentence	denotes	These included: cDNAs where the insert appeared to be cloned in the reverse orientation (six cDNAs); sequences from recently duplicated gene pairs, where sim4 assigned coding region and upstream exons to different members of the pair, although exons could equally well have been aligned closer to one another (six cDNAs); and artifacts due to use of sim4's N = 1 parameter (nine cDNAs).
T11710	42869-42874	DT	denotes	These
T11711	42875-42883	VBD	denotes	included
T11712	42883-42885	:	denotes	:
T11713	42885-42890	NNS	denotes	cDNAs
T11714	42891-42896	WRB	denotes	where
T11716	42897-42900	DT	denotes	the
T11717	42901-42907	NN	denotes	insert
T11715	42908-42916	VBD	denotes	appeared
T11718	42917-42919	TO	denotes	to
T11720	42920-42922	VB	denotes	be
T11719	42923-42929	VBN	denotes	cloned
T11721	42930-42932	IN	denotes	in
T11722	42933-42936	DT	denotes	the
T11724	42937-42944	JJ	denotes	reverse
T11723	42945-42956	NN	denotes	orientation
T11725	42957-42958	-LRB-	denotes	(
T11727	42958-42961	CD	denotes	six
T11726	42962-42967	NNS	denotes	cDNAs
T11728	42967-42968	-RRB-	denotes	)
T11729	42968-42969	:	denotes	;
T11730	42970-42979	NNS	denotes	sequences
T11731	42980-42984	IN	denotes	from
T11732	42985-42993	RB	denotes	recently
T11733	42994-43004	VBN	denotes	duplicated
T11735	43005-43009	NN	denotes	gene
T11734	43010-43015	NNS	denotes	pairs
T11736	43015-43017	,	denotes	,
T11737	43017-43022	WRB	denotes	where
T11739	43023-43027	NN	denotes	sim4
T11738	43028-43036	VBN	denotes	assigned
T11740	43037-43043	VBG	denotes	coding
T11741	43044-43050	NN	denotes	region
T11742	43051-43054	CC	denotes	and
T11743	43055-43063	JJ	denotes	upstream
T11744	43064-43069	NNS	denotes	exons
T11745	43070-43072	IN	denotes	to
T11746	43073-43082	JJ	denotes	different
T11747	43083-43090	NNS	denotes	members
T11748	43091-43093	IN	denotes	of
T11749	43094-43097	DT	denotes	the
T11750	43098-43102	NN	denotes	pair
T11751	43102-43104	,	denotes	,
T11752	43104-43112	IN	denotes	although
T11754	43113-43118	NNS	denotes	exons
T11755	43119-43124	MD	denotes	could
T11756	43125-43132	RB	denotes	equally
T11757	43133-43137	RB	denotes	well
T11758	43138-43142	VBP	denotes	have
T11759	43143-43147	VBN	denotes	been
T11753	43148-43155	VBN	denotes	aligned
T11760	43156-43162	RBR	denotes	closer
T11761	43163-43165	IN	denotes	to
T11762	43166-43169	CD	denotes	one
T11763	43170-43177	DT	denotes	another
T11764	43178-43179	-LRB-	denotes	(
T11766	43179-43182	CD	denotes	six
T11765	43183-43188	NNS	denotes	cDNAs
T11767	43188-43189	-RRB-	denotes	)
T11768	43189-43190	:	denotes	;
T11769	43191-43194	CC	denotes	and
T11770	43195-43204	NNS	denotes	artifacts
T11771	43205-43208	IN	denotes	due
T11772	43209-43211	IN	denotes	to
T11773	43212-43215	NN	denotes	use
T11774	43216-43218	IN	denotes	of
T11775	43219-43223	NN	denotes	sim4
T11777	43223-43225	POS	denotes	's
T11778	43226-43227	NN	denotes	N
T11780	43228-43229	SYM	denotes	=
T11779	43230-43231	CD	denotes	1
T11776	43232-43241	NN	denotes	parameter
T11781	43242-43243	-LRB-	denotes	(
T11783	43243-43247	CD	denotes	nine
T11782	43248-43253	NNS	denotes	cDNAs
T11784	43253-43254	-RRB-	denotes	)
T11785	43254-43255	.	denotes	.
T11786	43255-43407	sentence	denotes	This parameter instructs the program to make extra effort to match small upstream exons, allowing a greater total length of EST sequence to be matched.
T11787	43256-43260	DT	denotes	This
T11788	43261-43270	NN	denotes	parameter
T11789	43271-43280	VBZ	denotes	instructs
T11790	43281-43284	DT	denotes	the
T11791	43285-43292	NN	denotes	program
T11792	43293-43295	TO	denotes	to
T11793	43296-43300	VB	denotes	make
T11794	43301-43306	JJ	denotes	extra
T11795	43307-43313	NN	denotes	effort
T11796	43314-43316	TO	denotes	to
T11797	43317-43322	VB	denotes	match
T11798	43323-43328	JJ	denotes	small
T11800	43329-43337	JJ	denotes	upstream
T11799	43338-43343	NNS	denotes	exons
T11801	43343-43345	,	denotes	,
T11802	43345-43353	VBG	denotes	allowing
T11803	43354-43355	DT	denotes	a
T11805	43356-43363	JJR	denotes	greater
T11806	43364-43369	JJ	denotes	total
T11804	43370-43376	NN	denotes	length
T11808	43377-43379	IN	denotes	of
T11809	43380-43383	NN	denotes	EST
T11810	43384-43392	NN	denotes	sequence
T11811	43393-43395	TO	denotes	to
T11812	43396-43398	VB	denotes	be
T11807	43399-43406	VBN	denotes	matched
T11813	43406-43407	.	denotes	.
T11814	43407-43605	sentence	denotes	However, occasionally the N = 1 parameter caused the program to assign very small sequences (1-4 bp) to distant upstream exons, when they probably match nearer to the corresponding coding sequence.
T11815	43408-43415	RB	denotes	However
T11817	43415-43417	,	denotes	,
T11818	43417-43429	RB	denotes	occasionally
T11819	43430-43433	DT	denotes	the
T11821	43434-43435	NN	denotes	N
T11823	43436-43437	SYM	denotes	=
T11822	43438-43439	CD	denotes	1
T11820	43440-43449	NN	denotes	parameter
T11816	43450-43456	VBD	denotes	caused
T11824	43457-43460	DT	denotes	the
T11825	43461-43468	NN	denotes	program
T11827	43469-43471	TO	denotes	to
T11826	43472-43478	VB	denotes	assign
T11828	43479-43483	RB	denotes	very
T11829	43484-43489	JJ	denotes	small
T11830	43490-43499	NNS	denotes	sequences
T11831	43500-43501	-LRB-	denotes	(
T11833	43501-43502	CD	denotes	1
T11835	43502-43503	HYPH	denotes	-
T11834	43503-43504	CD	denotes	4
T11832	43505-43507	NNS	denotes	bp
T11836	43507-43508	-RRB-	denotes	)
T11837	43509-43511	IN	denotes	to
T11838	43512-43519	JJ	denotes	distant
T11840	43520-43528	JJ	denotes	upstream
T11839	43529-43534	NNS	denotes	exons
T11841	43534-43536	,	denotes	,
T11842	43536-43540	WRB	denotes	when
T11844	43541-43545	PRP	denotes	they
T11845	43546-43554	RB	denotes	probably
T11843	43555-43560	VBP	denotes	match
T11846	43561-43567	RBR	denotes	nearer
T11847	43568-43570	IN	denotes	to
T11848	43571-43574	DT	denotes	the
T11850	43575-43588	VBG	denotes	corresponding
T11851	43589-43595	NN	denotes	coding
T11849	43596-43604	NN	denotes	sequence
T11852	43604-43605	.	denotes	.
T11853	43605-43944	sentence	denotes	The expected distribution shown in Figure 2 was calculated using the equation P(x) = e-μμx/x!, where P(x) is the Poisson probability of observing x cDNAs per gene, and μ is the mean number of cDNAs observed per gene (μ = 1,176/983: 1,176 cDNAs matching olfactory receptor genes in our dataset and 983 intact class II olfactory receptors).
T11854	43606-43609	DT	denotes	The
T11856	43610-43618	JJ	denotes	expected
T11855	43619-43631	NN	denotes	distribution
T11858	43632-43637	VBN	denotes	shown
T11859	43638-43640	IN	denotes	in
T11860	43641-43647	NN	denotes	Figure
T11861	43648-43649	CD	denotes	2
T11862	43650-43653	VBD	denotes	was
T11857	43654-43664	VBN	denotes	calculated
T11863	43665-43670	VBG	denotes	using
T11864	43671-43674	DT	denotes	the
T11865	43675-43683	NN	denotes	equation
T11866	43684-43685	NN	denotes	P
T11868	43685-43686	-LRB-	denotes	(
T11867	43686-43687	NN	denotes	x
T11870	43687-43688	-RRB-	denotes	)
T11871	43689-43690	SYM	denotes	=
T11872	43691-43692	NN	denotes	e
T11873	43692-43693	SYM	denotes	-
T11869	43693-43696	NN	denotes	μμx
T11874	43696-43697	SYM	denotes	/
T11875	43697-43698	NN	denotes	x
T11876	43698-43699	SYM	denotes	!
T11877	43699-43701	,	denotes	,
T11878	43701-43706	WRB	denotes	where
T11880	43707-43708	NN	denotes	P
T11882	43708-43709	-LRB-	denotes	(
T11881	43709-43710	NN	denotes	x
T11883	43710-43711	-RRB-	denotes	)
T11879	43712-43714	VBZ	denotes	is
T11884	43715-43718	DT	denotes	the
T11886	43719-43726	NNP	denotes	Poisson
T11885	43727-43738	NN	denotes	probability
T11887	43739-43741	IN	denotes	of
T11888	43742-43751	VBG	denotes	observing
T11889	43752-43753	NN	denotes	x
T11890	43754-43759	NNS	denotes	cDNAs
T11891	43760-43763	IN	denotes	per
T11892	43764-43768	NN	denotes	gene
T11893	43768-43770	,	denotes	,
T11894	43770-43773	CC	denotes	and
T11895	43774-43775	NN	denotes	μ
T11896	43776-43778	VBZ	denotes	is
T11897	43779-43782	DT	denotes	the
T11899	43783-43787	JJ	denotes	mean
T11898	43788-43794	NN	denotes	number
T11900	43795-43797	IN	denotes	of
T11901	43798-43803	NNS	denotes	cDNAs
T11902	43804-43812	VBN	denotes	observed
T11903	43813-43816	IN	denotes	per
T11904	43817-43821	NN	denotes	gene
T11905	43822-43823	-LRB-	denotes	(
T11907	43823-43824	NN	denotes	μ
T11909	43825-43826	SYM	denotes	=
T11908	43827-43828	CD	denotes	1
T11910	43828-43829	,	denotes	,
T11911	43829-43832	CD	denotes	176
T11913	43832-43833	SYM	denotes	/
T11912	43833-43836	CD	denotes	983
T11914	43836-43838	:	denotes	:
T11915	43838-43843	CD	denotes	1,176
T11906	43844-43849	NNS	denotes	cDNAs
T11916	43850-43858	VBG	denotes	matching
T11917	43859-43868	JJ	denotes	olfactory
T11918	43869-43877	NN	denotes	receptor
T11919	43878-43883	NNS	denotes	genes
T11920	43884-43886	IN	denotes	in
T11921	43887-43890	PRP$	denotes	our
T11922	43891-43898	NN	denotes	dataset
T11923	43899-43902	CC	denotes	and
T11924	43903-43906	CD	denotes	983
T11926	43907-43913	JJ	denotes	intact
T11927	43914-43919	NN	denotes	class
T11928	43920-43922	CD	denotes	II
T11929	43923-43932	JJ	denotes	olfactory
T11925	43933-43942	NNS	denotes	receptors
T11930	43942-43943	-RRB-	denotes	)
T11931	43943-43944	.	denotes	.
T11932	43944-44113	sentence	denotes	In our analysis of expressed pseudogenes, we ignored two olfactory receptor pseudogenes found very near the ends of genomic sequences and thus likely to be error-prone.
T11933	43945-43947	IN	denotes	In
T11935	43948-43951	PRP$	denotes	our
T11936	43952-43960	NN	denotes	analysis
T11937	43961-43963	IN	denotes	of
T11938	43964-43973	JJ	denotes	expressed
T11939	43974-43985	NNS	denotes	pseudogenes
T11940	43985-43987	,	denotes	,
T11941	43987-43989	PRP	denotes	we
T11934	43990-43997	VBD	denotes	ignored
T11942	43998-44001	CD	denotes	two
T11944	44002-44011	JJ	denotes	olfactory
T11945	44012-44020	NN	denotes	receptor
T11943	44021-44032	NNS	denotes	pseudogenes
T11946	44033-44038	VBN	denotes	found
T11947	44039-44043	RB	denotes	very
T11948	44044-44048	IN	denotes	near
T11949	44049-44052	DT	denotes	the
T11950	44053-44057	NNS	denotes	ends
T11951	44058-44060	IN	denotes	of
T11952	44061-44068	JJ	denotes	genomic
T11953	44069-44078	NNS	denotes	sequences
T11954	44079-44082	CC	denotes	and
T11955	44083-44087	RB	denotes	thus
T11956	44088-44094	JJ	denotes	likely
T11957	44095-44097	TO	denotes	to
T11958	44098-44100	VB	denotes	be
T11959	44101-44106	NN	denotes	error
T11961	44106-44107	HYPH	denotes	-
T11960	44107-44112	JJ	denotes	prone
T11962	44112-44113	.	denotes	.
T11963	44113-44385	sentence	denotes	A protein sequence alignment of intact mouse olfactory receptors was generated using CLUSTALW [50], edited by hand, and used to produce the phylogenetic tree shown in Figure 1 using PAUP's neighbor-joining algorithm (v4.0b6 Version 4, Sinauer Associates, Sunderland, MA).
T11964	44114-44115	DT	denotes	A
T11966	44116-44123	NN	denotes	protein
T11967	44124-44132	NN	denotes	sequence
T11965	44133-44142	NN	denotes	alignment
T11969	44143-44145	IN	denotes	of
T11970	44146-44152	JJ	denotes	intact
T11972	44153-44158	NN	denotes	mouse
T11973	44159-44168	JJ	denotes	olfactory
T11971	44169-44178	NNS	denotes	receptors
T11974	44179-44182	VBD	denotes	was
T11968	44183-44192	VBN	denotes	generated
T11975	44193-44198	VBG	denotes	using
T11976	44199-44207	NN	denotes	CLUSTALW
T11977	44208-44209	-LRB-	denotes	[
T11978	44209-44211	CD	denotes	50
T11979	44211-44212	-RRB-	denotes	]
T11980	44212-44214	,	denotes	,
T11981	44214-44220	VBN	denotes	edited
T11982	44221-44223	IN	denotes	by
T11983	44224-44228	NN	denotes	hand
T11984	44228-44230	,	denotes	,
T11985	44230-44233	CC	denotes	and
T11986	44234-44238	VBN	denotes	used
T11987	44239-44241	TO	denotes	to
T11988	44242-44249	VB	denotes	produce
T11989	44250-44253	DT	denotes	the
T11991	44254-44266	JJ	denotes	phylogenetic
T11990	44267-44271	NN	denotes	tree
T11992	44272-44277	VBN	denotes	shown
T11993	44278-44280	IN	denotes	in
T11994	44281-44287	NN	denotes	Figure
T11995	44288-44289	CD	denotes	1
T11996	44290-44295	VBG	denotes	using
T11997	44296-44300	NN	denotes	PAUP
T11999	44300-44302	POS	denotes	's
T12000	44303-44311	NN	denotes	neighbor
T12002	44311-44312	HYPH	denotes	-
T12001	44312-44319	VBG	denotes	joining
T11998	44320-44329	NN	denotes	algorithm
T12003	44330-44331	-LRB-	denotes	(
T12005	44331-44337	NN	denotes	v4.0b6
T12006	44338-44345	NN	denotes	Version
T12007	44346-44347	CD	denotes	4
T12008	44347-44349	,	denotes	,
T12009	44349-44356	NNP	denotes	Sinauer
T12004	44357-44367	NNPS	denotes	Associates
T12010	44367-44369	,	denotes	,
T12011	44369-44379	NNP	denotes	Sunderland
T12012	44379-44381	,	denotes	,
T12013	44381-44383	NNP	denotes	MA
T12014	44383-44384	-RRB-	denotes	)
T12015	44384-44385	.	denotes	.
T12016	44385-44429	sentence	denotes	The tree was colored using a custom script.
T12017	44386-44389	DT	denotes	The
T12018	44390-44394	NN	denotes	tree
T12020	44395-44398	VBD	denotes	was
T12019	44399-44406	VBN	denotes	colored
T12021	44407-44412	VBG	denotes	using
T12022	44413-44414	DT	denotes	a
T12024	44415-44421	NN	denotes	custom
T12023	44422-44428	NN	denotes	script
T12025	44428-44429	.	denotes	.
T12026	44429-44574	sentence	denotes	Information content (the measure of sequence conservation shown in Figure 6) was calculated for each position in the alignment using alpro [51].
T12027	44430-44441	NN	denotes	Information
T12028	44442-44449	NN	denotes	content
T12030	44450-44451	-LRB-	denotes	(
T12031	44451-44454	DT	denotes	the
T12032	44455-44462	NN	denotes	measure
T12033	44463-44465	IN	denotes	of
T12034	44466-44474	NN	denotes	sequence
T12035	44475-44487	NN	denotes	conservation
T12036	44488-44493	VBN	denotes	shown
T12037	44494-44496	IN	denotes	in
T12038	44497-44503	NN	denotes	Figure
T12039	44504-44505	CD	denotes	6
T12040	44505-44506	-RRB-	denotes	)
T12041	44507-44510	VBD	denotes	was
T12029	44511-44521	VBN	denotes	calculated
T12042	44522-44525	IN	denotes	for
T12043	44526-44530	DT	denotes	each
T12044	44531-44539	NN	denotes	position
T12045	44540-44542	IN	denotes	in
T12046	44543-44546	DT	denotes	the
T12047	44547-44556	NN	denotes	alignment
T12048	44557-44562	VBG	denotes	using
T12049	44563-44568	NN	denotes	alpro
T12050	44569-44570	-LRB-	denotes	[
T12051	44570-44572	CD	denotes	51
T12052	44572-44573	-RRB-	denotes	]
T12053	44573-44574	.	denotes	.
T12054	44574-44733	sentence	denotes	To determine the number of transcriptional isoforms for each gene, we examined the sim4 output for every matching cDNA in decreasing order of number of exons.
T12055	44575-44577	TO	denotes	To
T12056	44578-44587	VB	denotes	determine
T12058	44588-44591	DT	denotes	the
T12059	44592-44598	NN	denotes	number
T12060	44599-44601	IN	denotes	of
T12061	44602-44617	JJ	denotes	transcriptional
T12062	44618-44626	NNS	denotes	isoforms
T12063	44627-44630	IN	denotes	for
T12064	44631-44635	DT	denotes	each
T12065	44636-44640	NN	denotes	gene
T12066	44640-44642	,	denotes	,
T12067	44642-44644	PRP	denotes	we
T12057	44645-44653	VBD	denotes	examined
T12068	44654-44657	DT	denotes	the
T12070	44658-44662	NN	denotes	sim4
T12069	44663-44669	NN	denotes	output
T12071	44670-44673	IN	denotes	for
T12072	44674-44679	DT	denotes	every
T12074	44680-44688	VBG	denotes	matching
T12073	44689-44693	NN	denotes	cDNA
T12075	44694-44696	IN	denotes	in
T12076	44697-44707	JJ	denotes	decreasing
T12077	44708-44713	NN	denotes	order
T12078	44714-44716	IN	denotes	of
T12079	44717-44723	NN	denotes	number
T12080	44724-44726	IN	denotes	of
T12081	44727-44732	NNS	denotes	exons
T12082	44732-44733	.	denotes	.
T12083	44733-44899	sentence	denotes	The first cDNA was counted as one splice form, and for each subsequent cDNA, we determined whether exon structure was mutually exclusive to isoforms already counted.
T12084	44734-44737	DT	denotes	The
T12086	44738-44743	JJ	denotes	first
T12085	44744-44748	NN	denotes	cDNA
T12088	44749-44752	VBD	denotes	was
T12087	44753-44760	VBN	denotes	counted
T12089	44761-44763	IN	denotes	as
T12090	44764-44767	CD	denotes	one
T12092	44768-44774	NN	denotes	splice
T12091	44775-44779	NN	denotes	form
T12093	44779-44781	,	denotes	,
T12094	44781-44784	CC	denotes	and
T12095	44785-44788	IN	denotes	for
T12097	44789-44793	DT	denotes	each
T12099	44794-44804	JJ	denotes	subsequent
T12098	44805-44809	NN	denotes	cDNA
T12100	44809-44811	,	denotes	,
T12101	44811-44813	PRP	denotes	we
T12096	44814-44824	VBD	denotes	determined
T12102	44825-44832	IN	denotes	whether
T12104	44833-44837	NN	denotes	exon
T12105	44838-44847	NN	denotes	structure
T12103	44848-44851	VBD	denotes	was
T12106	44852-44860	RB	denotes	mutually
T12107	44861-44870	JJ	denotes	exclusive
T12108	44871-44873	IN	denotes	to
T12109	44874-44882	NNS	denotes	isoforms
T12110	44883-44890	RB	denotes	already
T12111	44891-44898	VBN	denotes	counted
T12112	44898-44899	.	denotes	.
T12113	44899-45058	sentence	denotes	We were conservative in our definition of mutually exclusive, and thus our count represents the minimum number of isoforms represented in the cDNA collection.
T12114	44900-44902	PRP	denotes	We
T12115	44903-44907	VBD	denotes	were
T12116	44908-44920	JJ	denotes	conservative
T12117	44921-44923	IN	denotes	in
T12118	44924-44927	PRP$	denotes	our
T12119	44928-44938	NN	denotes	definition
T12120	44939-44941	IN	denotes	of
T12121	44942-44950	RB	denotes	mutually
T12122	44951-44960	JJ	denotes	exclusive
T12123	44960-44962	,	denotes	,
T12124	44962-44965	CC	denotes	and
T12125	44966-44970	RB	denotes	thus
T12127	44971-44974	PRP$	denotes	our
T12128	44975-44980	NN	denotes	count
T12126	44981-44991	VBZ	denotes	represents
T12129	44992-44995	DT	denotes	the
T12131	44996-45003	JJ	denotes	minimum
T12130	45004-45010	NN	denotes	number
T12132	45011-45013	IN	denotes	of
T12133	45014-45022	NNS	denotes	isoforms
T12134	45023-45034	VBN	denotes	represented
T12135	45035-45037	IN	denotes	in
T12136	45038-45041	DT	denotes	the
T12138	45042-45046	NN	denotes	cDNA
T12137	45047-45057	NN	denotes	collection
T12139	45057-45058	.	denotes	.
T12269	45060-45062	NN	denotes	RT
T12271	45062-45063	HYPH	denotes	-
T12270	45063-45066	NN	denotes	PCR
T12272	45066-45195	sentence	denotes	The olfactory epithelia were dissected from three adult female C57BL/6 mice, including tissues attached to the skull and septum.
T12273	45067-45070	DT	denotes	The
T12275	45071-45080	JJ	denotes	olfactory
T12274	45081-45090	NNS	denotes	epithelia
T12277	45091-45095	VBD	denotes	were
T12276	45096-45105	VBN	denotes	dissected
T12278	45106-45110	IN	denotes	from
T12279	45111-45116	CD	denotes	three
T12281	45117-45122	JJ	denotes	adult
T12282	45123-45129	JJ	denotes	female
T12283	45130-45135	NN	denotes	C57BL
T12284	45135-45136	HYPH	denotes	/
T12285	45136-45137	CD	denotes	6
T12280	45138-45142	NNS	denotes	mice
T12286	45142-45144	,	denotes	,
T12287	45144-45153	VBG	denotes	including
T12288	45154-45161	NNS	denotes	tissues
T12289	45162-45170	VBN	denotes	attached
T12290	45171-45173	IN	denotes	to
T12291	45174-45177	DT	denotes	the
T12292	45178-45183	NN	denotes	skull
T12293	45184-45187	CC	denotes	and
T12294	45188-45194	NN	denotes	septum
T12295	45194-45195	.	denotes	.
T12296	45195-45308	sentence	denotes	RNA was isolated using the Qiagen RNeasy midi kit (Qiagen, Valencia, CA, USA), including a DNase treatment step.
T12297	45196-45199	NN	denotes	RNA
T12299	45200-45203	VBD	denotes	was
T12298	45204-45212	VBN	denotes	isolated
T12300	45213-45218	VBG	denotes	using
T12301	45219-45222	DT	denotes	the
T12303	45223-45229	NNP	denotes	Qiagen
T12304	45230-45236	NNP	denotes	RNeasy
T12305	45237-45241	NN	denotes	midi
T12302	45242-45245	NN	denotes	kit
T12306	45246-45247	-LRB-	denotes	(
T12307	45247-45253	NNP	denotes	Qiagen
T12308	45253-45255	,	denotes	,
T12309	45255-45263	NNP	denotes	Valencia
T12310	45263-45265	,	denotes	,
T12311	45265-45267	NNP	denotes	CA
T12312	45267-45269	,	denotes	,
T12313	45269-45272	NNP	denotes	USA
T12314	45272-45273	-RRB-	denotes	)
T12315	45273-45275	,	denotes	,
T12316	45275-45284	VBG	denotes	including
T12317	45285-45286	DT	denotes	a
T12319	45287-45292	NN	denotes	DNase
T12320	45293-45302	NN	denotes	treatment
T12318	45303-45307	NN	denotes	step
T12321	45307-45308	.	denotes	.
T12322	45308-45537	sentence	denotes	First-strand cDNA was produced from 2.5 μg of RNA in a volume of 50 μl using random hexamers and Invitrogen's Superscript II reverse transcriptase (Invitrogen, Carlsbad, CA, USA), according to the manufacturer's recommendations.
T12323	45309-45314	JJ	denotes	First
T12325	45314-45315	HYPH	denotes	-
T12324	45315-45321	NN	denotes	strand
T12326	45322-45326	NN	denotes	cDNA
T12328	45327-45330	VBD	denotes	was
T12327	45331-45339	VBN	denotes	produced
T12329	45340-45344	IN	denotes	from
T12330	45345-45348	CD	denotes	2.5
T12331	45349-45351	NNS	denotes	μg
T12332	45352-45354	IN	denotes	of
T12333	45355-45358	NN	denotes	RNA
T12334	45359-45361	IN	denotes	in
T12335	45362-45363	DT	denotes	a
T12336	45364-45370	NN	denotes	volume
T12337	45371-45373	IN	denotes	of
T12338	45374-45376	CD	denotes	50
T12339	45377-45379	NNS	denotes	μl
T12340	45380-45385	VBG	denotes	using
T12341	45386-45392	JJ	denotes	random
T12342	45393-45401	NNS	denotes	hexamers
T12343	45402-45405	CC	denotes	and
T12344	45406-45416	NNP	denotes	Invitrogen
T12346	45416-45418	POS	denotes	's
T12347	45419-45430	NN	denotes	Superscript
T12348	45431-45433	CD	denotes	II
T12349	45434-45441	JJ	denotes	reverse
T12345	45442-45455	NN	denotes	transcriptase
T12350	45456-45457	-LRB-	denotes	(
T12351	45457-45467	NNP	denotes	Invitrogen
T12352	45467-45469	,	denotes	,
T12353	45469-45477	NNP	denotes	Carlsbad
T12354	45477-45479	,	denotes	,
T12355	45479-45481	NNP	denotes	CA
T12356	45481-45483	,	denotes	,
T12357	45483-45486	NNP	denotes	USA
T12358	45486-45487	-RRB-	denotes	)
T12359	45487-45489	,	denotes	,
T12360	45489-45498	VBG	denotes	according
T12361	45499-45501	IN	denotes	to
T12362	45502-45505	DT	denotes	the
T12363	45506-45518	NN	denotes	manufacturer
T12365	45518-45520	POS	denotes	's
T12364	45521-45536	NNS	denotes	recommendations
T12366	45536-45537	.	denotes	.
T12367	45537-45626	sentence	denotes	One-twenty-fifth of the resulting cDNA was used as template in subsequent PCR reactions.
T12368	45538-45541	CD	denotes	One
T12370	45541-45542	HYPH	denotes	-
T12371	45542-45548	CD	denotes	twenty
T12372	45548-45549	HYPH	denotes	-
T12369	45549-45554	NN	denotes	fifth
T12374	45555-45557	IN	denotes	of
T12375	45558-45561	DT	denotes	the
T12377	45562-45571	VBG	denotes	resulting
T12376	45572-45576	NN	denotes	cDNA
T12378	45577-45580	VBD	denotes	was
T12373	45581-45585	VBN	denotes	used
T12379	45586-45588	IN	denotes	as
T12380	45589-45597	NN	denotes	template
T12381	45598-45600	IN	denotes	in
T12382	45601-45611	JJ	denotes	subsequent
T12384	45612-45615	NN	denotes	PCR
T12383	45616-45625	NNS	denotes	reactions
T12385	45625-45626	.	denotes	.
T12386	45626-45814	sentence	denotes	PCR amplification biased towards class I olfactory receptors was performed using degenerate primers P26 [17] and classI_R1 (5'-GGRTTIADIRYIGGNGG-3') with an annealing temperature of 44°C.
T12387	45627-45630	NN	denotes	PCR
T12388	45631-45644	NN	denotes	amplification
T12390	45645-45651	JJ	denotes	biased
T12391	45652-45659	IN	denotes	towards
T12392	45660-45665	NN	denotes	class
T12394	45666-45667	CD	denotes	I
T12395	45668-45677	JJ	denotes	olfactory
T12393	45678-45687	NNS	denotes	receptors
T12396	45688-45691	VBD	denotes	was
T12389	45692-45701	VBN	denotes	performed
T12397	45702-45707	VBG	denotes	using
T12398	45708-45718	JJ	denotes	degenerate
T12400	45719-45726	NNS	denotes	primers
T12399	45727-45730	NN	denotes	P26
T12401	45731-45732	-LRB-	denotes	[
T12402	45732-45734	CD	denotes	17
T12403	45734-45735	-RRB-	denotes	]
T12404	45736-45739	CC	denotes	and
T12405	45740-45749	NN	denotes	classI_R1
T12406	45750-45751	-LRB-	denotes	(
T12408	45751-45752	CD	denotes	5
T12409	45752-45753	SYM	denotes	'
T12410	45753-45754	HYPH	denotes	-
T12407	45754-45771	NN	denotes	GGRTTIADIRYIGGNGG
T12411	45771-45772	HYPH	denotes	-
T12412	45772-45773	CD	denotes	3
T12413	45773-45774	SYM	denotes	'
T12414	45774-45775	-RRB-	denotes	)
T12415	45776-45780	IN	denotes	with
T12416	45781-45783	DT	denotes	an
T12418	45784-45793	JJ	denotes	annealing
T12417	45794-45805	NN	denotes	temperature
T12419	45806-45808	IN	denotes	of
T12420	45809-45811	CD	denotes	44
T12421	45811-45813	NNS	denotes	°C
T12422	45813-45814	.	denotes	.
T12423	45814-45905	sentence	denotes	The product was cloned (TA cloning kit, Invitrogen), and individual clones were sequenced.
T12424	45815-45818	DT	denotes	The
T12425	45819-45826	NN	denotes	product
T12427	45827-45830	VBD	denotes	was
T12426	45831-45837	VBN	denotes	cloned
T12428	45838-45839	-LRB-	denotes	(
T12430	45839-45841	NN	denotes	TA
T12432	45842-45849	NN	denotes	cloning
T12431	45850-45853	NN	denotes	kit
T12433	45853-45855	,	denotes	,
T12429	45855-45865	NNP	denotes	Invitrogen
T12434	45865-45866	-RRB-	denotes	)
T12435	45866-45868	,	denotes	,
T12436	45868-45871	CC	denotes	and
T12437	45872-45882	JJ	denotes	individual
T12438	45883-45889	NNS	denotes	clones
T12440	45890-45894	VBD	denotes	were
T12439	45895-45904	VBN	denotes	sequenced
T12441	45904-45905	.	denotes	.
T12442	45905-46029	sentence	denotes	Specific PCR primers used to confirm expression of individual olfactory receptor genes are given in Additional data file 1.
T12443	45906-45914	JJ	denotes	Specific
T12445	45915-45918	NN	denotes	PCR
T12444	45919-45926	NNS	denotes	primers
T12447	45927-45931	VBN	denotes	used
T12448	45932-45934	TO	denotes	to
T12449	45935-45942	VB	denotes	confirm
T12450	45943-45953	NN	denotes	expression
T12451	45954-45956	IN	denotes	of
T12452	45957-45967	JJ	denotes	individual
T12454	45968-45977	JJ	denotes	olfactory
T12455	45978-45986	NN	denotes	receptor
T12453	45987-45992	NNS	denotes	genes
T12456	45993-45996	VBP	denotes	are
T12446	45997-46002	VBN	denotes	given
T12457	46003-46005	IN	denotes	in
T12458	46006-46016	JJ	denotes	Additional
T12460	46017-46021	NNS	denotes	data
T12459	46022-46026	NN	denotes	file
T12461	46027-46028	CD	denotes	1
T12462	46028-46029	.	denotes	.
T12463	46029-46128	sentence	denotes	Each PCR product was sequenced to confirm that the expected gene and no others had been amplified.
T12464	46030-46034	DT	denotes	Each
T12466	46035-46038	NN	denotes	PCR
T12465	46039-46046	NN	denotes	product
T12468	46047-46050	VBD	denotes	was
T12467	46051-46060	VBN	denotes	sequenced
T12469	46061-46063	TO	denotes	to
T12470	46064-46071	VB	denotes	confirm
T12471	46072-46076	IN	denotes	that
T12473	46077-46080	DT	denotes	the
T12475	46081-46089	VBN	denotes	expected
T12474	46090-46094	NN	denotes	gene
T12476	46095-46098	CC	denotes	and
T12477	46099-46101	DT	denotes	no
T12478	46102-46108	NNS	denotes	others
T12479	46109-46112	VBD	denotes	had
T12480	46113-46117	VBN	denotes	been
T12472	46118-46127	VBN	denotes	amplified
T12481	46127-46128	.	denotes	.
T12482	46128-46287	sentence	denotes	Control reactions on a template made by omitting reverse transcriptase gave no product, indicating that the RNA preparation was uncontaminated by genomic DNA.
T12483	46129-46136	NN	denotes	Control
T12484	46137-46146	NNS	denotes	reactions
T12486	46147-46149	IN	denotes	on
T12487	46150-46151	DT	denotes	a
T12488	46152-46160	NN	denotes	template
T12489	46161-46165	VBN	denotes	made
T12490	46166-46168	IN	denotes	by
T12491	46169-46177	VBG	denotes	omitting
T12492	46178-46185	JJ	denotes	reverse
T12493	46186-46199	NN	denotes	transcriptase
T12485	46200-46204	VBD	denotes	gave
T12494	46205-46207	DT	denotes	no
T12495	46208-46215	NN	denotes	product
T12496	46215-46217	,	denotes	,
T12497	46217-46227	VBG	denotes	indicating
T12498	46228-46232	IN	denotes	that
T12500	46233-46236	DT	denotes	the
T12502	46237-46240	NN	denotes	RNA
T12501	46241-46252	NN	denotes	preparation
T12499	46253-46256	VBD	denotes	was
T12503	46257-46271	JJ	denotes	uncontaminated
T12504	46272-46274	IN	denotes	by
T12505	46275-46282	JJ	denotes	genomic
T12506	46283-46286	NN	denotes	DNA
T12507	46286-46287	.	denotes	.
T12508	46287-46490	sentence	denotes	Relative transcript levels were estimated using real-time PCR according to Applied Biosystems' protocols, with magnesium concentration, primer pair and fluorescent probe given in Additional data file 2.
T12509	46288-46296	JJ	denotes	Relative
T12511	46297-46307	NN	denotes	transcript
T12510	46308-46314	NNS	denotes	levels
T12513	46315-46319	VBD	denotes	were
T12512	46320-46329	VBN	denotes	estimated
T12514	46330-46335	VBG	denotes	using
T12515	46336-46340	JJ	denotes	real
T12517	46340-46341	HYPH	denotes	-
T12516	46341-46345	NN	denotes	time
T12518	46346-46349	NN	denotes	PCR
T12519	46350-46359	VBG	denotes	according
T12520	46360-46362	IN	denotes	to
T12521	46363-46370	NNP	denotes	Applied
T12522	46371-46381	NNP	denotes	Biosystems
T12524	46381-46382	POS	denotes	'
T12523	46383-46392	NNS	denotes	protocols
T12525	46392-46394	,	denotes	,
T12526	46394-46398	IN	denotes	with
T12528	46399-46408	NN	denotes	magnesium
T12529	46409-46422	NN	denotes	concentration
T12530	46422-46424	,	denotes	,
T12531	46424-46430	NN	denotes	primer
T12532	46431-46435	NN	denotes	pair
T12533	46436-46439	CC	denotes	and
T12534	46440-46451	JJ	denotes	fluorescent
T12535	46452-46457	NN	denotes	probe
T12527	46458-46463	VBN	denotes	given
T12536	46464-46466	IN	denotes	in
T12537	46467-46477	JJ	denotes	Additional
T12539	46478-46482	NNS	denotes	data
T12538	46483-46487	NN	denotes	file
T12540	46488-46489	CD	denotes	2
T12541	46489-46490	.	denotes	.
T12542	46490-46577	sentence	denotes	The increase in fluorescence during thermocycling was measured on an ABI PRISM 7900HT.
T12543	46491-46494	DT	denotes	The
T12544	46495-46503	NN	denotes	increase
T12546	46504-46506	IN	denotes	in
T12547	46507-46519	NN	denotes	fluorescence
T12548	46520-46526	IN	denotes	during
T12549	46527-46540	NN	denotes	thermocycling
T12550	46541-46544	VBD	denotes	was
T12545	46545-46553	VBN	denotes	measured
T12551	46554-46556	IN	denotes	on
T12552	46557-46559	DT	denotes	an
T12554	46560-46563	NN	denotes	ABI
T12555	46564-46569	NN	denotes	PRISM
T12553	46570-46576	NN	denotes	7900HT
T12556	46576-46577	.	denotes	.
T12557	46577-46775	sentence	denotes	Standard curves were constructed for each primer pair using triplicate samples of mouse genomic DNA of nine known concentrations (range 0.01-100 ng, or about 3-30,000 copies of the haploid genome).
T12558	46578-46586	JJ	denotes	Standard
T12559	46587-46593	NNS	denotes	curves
T12561	46594-46598	VBD	denotes	were
T12560	46599-46610	VBN	denotes	constructed
T12562	46611-46614	IN	denotes	for
T12563	46615-46619	DT	denotes	each
T12565	46620-46626	NN	denotes	primer
T12564	46627-46631	NN	denotes	pair
T12566	46632-46637	VBG	denotes	using
T12567	46638-46648	NN	denotes	triplicate
T12568	46649-46656	NNS	denotes	samples
T12569	46657-46659	IN	denotes	of
T12570	46660-46665	NN	denotes	mouse
T12572	46666-46673	JJ	denotes	genomic
T12571	46674-46677	NN	denotes	DNA
T12573	46678-46680	IN	denotes	of
T12574	46681-46685	CD	denotes	nine
T12576	46686-46691	VBN	denotes	known
T12575	46692-46706	NNS	denotes	concentrations
T12577	46707-46708	-LRB-	denotes	(
T12579	46708-46713	NN	denotes	range
T12580	46714-46718	CD	denotes	0.01
T12582	46718-46719	SYM	denotes	-
T12581	46719-46722	CD	denotes	100
T12578	46723-46725	NNS	denotes	ng
T12583	46725-46727	,	denotes	,
T12584	46727-46729	CC	denotes	or
T12585	46730-46735	RB	denotes	about
T12587	46736-46737	CD	denotes	3
T12588	46737-46738	SYM	denotes	-
T12586	46738-46744	CD	denotes	30,000
T12589	46745-46751	NNS	denotes	copies
T12590	46752-46754	IN	denotes	of
T12591	46755-46758	DT	denotes	the
T12593	46759-46766	NN	denotes	haploid
T12592	46767-46773	NN	denotes	genome
T12594	46773-46774	-RRB-	denotes	)
T12595	46774-46775	.	denotes	.
T12596	46775-47034	sentence	denotes	Relative expression level of each gene was determined by comparing the mean Ct (cycle where fluorescence reaches an arbitrary threshold value) obtained with six replicate samples of reverse-transcribed RNA to the standard curve for the corresponding primers.
T12597	46776-46784	JJ	denotes	Relative
T12599	46785-46795	NN	denotes	expression
T12598	46796-46801	NN	denotes	level
T12601	46802-46804	IN	denotes	of
T12602	46805-46809	DT	denotes	each
T12603	46810-46814	NN	denotes	gene
T12604	46815-46818	VBD	denotes	was
T12600	46819-46829	VBN	denotes	determined
T12605	46830-46832	IN	denotes	by
T12606	46833-46842	VBG	denotes	comparing
T12607	46843-46846	DT	denotes	the
T12609	46847-46851	JJ	denotes	mean
T12608	46852-46854	NN	denotes	Ct
T12610	46855-46856	-LRB-	denotes	(
T12611	46856-46861	NN	denotes	cycle
T12612	46862-46867	WRB	denotes	where
T12614	46868-46880	NN	denotes	fluorescence
T12613	46881-46888	VBZ	denotes	reaches
T12615	46889-46891	DT	denotes	an
T12617	46892-46901	JJ	denotes	arbitrary
T12618	46902-46911	NN	denotes	threshold
T12616	46912-46917	NN	denotes	value
T12619	46917-46918	-RRB-	denotes	)
T12620	46919-46927	VBN	denotes	obtained
T12621	46928-46932	IN	denotes	with
T12622	46933-46936	CD	denotes	six
T12624	46937-46946	NN	denotes	replicate
T12623	46947-46954	NNS	denotes	samples
T12625	46955-46957	IN	denotes	of
T12626	46958-46965	JJ	denotes	reverse
T12628	46965-46966	HYPH	denotes	-
T12627	46966-46977	JJ	denotes	transcribed
T12629	46978-46981	NN	denotes	RNA
T12630	46982-46984	IN	denotes	to
T12631	46985-46988	DT	denotes	the
T12633	46989-46997	JJ	denotes	standard
T12632	46998-47003	NN	denotes	curve
T12634	47004-47007	IN	denotes	for
T12635	47008-47011	DT	denotes	the
T12637	47012-47025	VBG	denotes	corresponding
T12636	47026-47033	NNS	denotes	primers
T12638	47033-47034	.	denotes	.
T12639	47034-47137	sentence	denotes	Relative RNA levels of a housekeeping gene, ribosomal S16, were measured as previously described [52].
T12640	47035-47043	JJ	denotes	Relative
T12642	47044-47047	NN	denotes	RNA
T12641	47048-47054	NNS	denotes	levels
T12644	47055-47057	IN	denotes	of
T12645	47058-47059	DT	denotes	a
T12647	47060-47072	NN	denotes	housekeeping
T12646	47073-47077	NN	denotes	gene
T12648	47077-47079	,	denotes	,
T12649	47079-47088	JJ	denotes	ribosomal
T12650	47089-47092	NN	denotes	S16
T12651	47092-47094	,	denotes	,
T12652	47094-47098	VBD	denotes	were
T12643	47099-47107	VBN	denotes	measured
T12653	47108-47110	IN	denotes	as
T12655	47111-47121	RB	denotes	previously
T12654	47122-47131	VBN	denotes	described
T12656	47132-47133	-LRB-	denotes	[
T12657	47133-47135	CD	denotes	52
T12658	47135-47136	-RRB-	denotes	]
T12659	47136-47137	.	denotes	.
T12660	47137-47280	sentence	denotes	Control reactions on template prepared by omitting reverse transcriptase amplified at a relative level of 0.03 ± 0.01 ng or less in each case.
T12661	47138-47145	NN	denotes	Control
T12662	47146-47155	NNS	denotes	reactions
T12664	47156-47158	IN	denotes	on
T12665	47159-47167	NN	denotes	template
T12666	47168-47176	VBN	denotes	prepared
T12667	47177-47179	IN	denotes	by
T12668	47180-47188	VBG	denotes	omitting
T12669	47189-47196	JJ	denotes	reverse
T12670	47197-47210	NN	denotes	transcriptase
T12663	47211-47220	VBN	denotes	amplified
T12671	47221-47223	IN	denotes	at
T12672	47224-47225	DT	denotes	a
T12674	47226-47234	JJ	denotes	relative
T12673	47235-47240	NN	denotes	level
T12675	47241-47243	IN	denotes	of
T12676	47244-47248	CD	denotes	0.03
T12678	47249-47250	SYM	denotes	±
T12677	47251-47255	CD	denotes	0.01
T12679	47256-47258	NNS	denotes	ng
T12680	47259-47261	CC	denotes	or
T12681	47262-47266	JJR	denotes	less
T12682	47267-47269	IN	denotes	in
T12683	47270-47274	DT	denotes	each
T12684	47275-47279	NN	denotes	case
T12685	47279-47280	.	denotes	.
T12686	47280-47408	sentence	denotes	Expression measurements of the seven genes were normalized for each mouse so that S16 levels were equal to 1 (arbitrary units).
T12687	47281-47291	NN	denotes	Expression
T12688	47292-47304	NNS	denotes	measurements
T12690	47305-47307	IN	denotes	of
T12691	47308-47311	DT	denotes	the
T12693	47312-47317	CD	denotes	seven
T12692	47318-47323	NNS	denotes	genes
T12694	47324-47328	VBD	denotes	were
T12689	47329-47339	VBN	denotes	normalized
T12695	47340-47343	IN	denotes	for
T12696	47344-47348	DT	denotes	each
T12697	47349-47354	NN	denotes	mouse
T12698	47355-47357	IN	denotes	so
T12700	47358-47362	IN	denotes	that
T12701	47363-47366	NN	denotes	S16
T12702	47367-47373	NNS	denotes	levels
T12699	47374-47378	VBD	denotes	were
T12703	47379-47384	JJ	denotes	equal
T12704	47385-47387	IN	denotes	to
T12705	47388-47389	CD	denotes	1
T12706	47390-47391	-LRB-	denotes	(
T12708	47391-47400	JJ	denotes	arbitrary
T12707	47401-47406	NNS	denotes	units
T12709	47406-47407	-RRB-	denotes	)
T12710	47407-47408	.	denotes	.
T12805	47410-47412	FW	denotes	In
T12806	47413-47417	FW	denotes	situ
T12807	47418-47431	NN	denotes	hybridization
T12808	47431-47547	sentence	denotes	Coronal sections were cut from the olfactory epithelia of an adult mouse (Figure 4) and a young (P6) C57BL/6 mouse.
T12809	47432-47439	JJ	denotes	Coronal
T12810	47440-47448	NNS	denotes	sections
T12812	47449-47453	VBD	denotes	were
T12811	47454-47457	VBN	denotes	cut
T12813	47458-47462	IN	denotes	from
T12814	47463-47466	DT	denotes	the
T12816	47467-47476	JJ	denotes	olfactory
T12815	47477-47486	NNS	denotes	epithelia
T12817	47487-47489	IN	denotes	of
T12818	47490-47492	DT	denotes	an
T12820	47493-47498	JJ	denotes	adult
T12819	47499-47504	NN	denotes	mouse
T12821	47505-47506	-LRB-	denotes	(
T12822	47506-47512	NN	denotes	Figure
T12823	47513-47514	CD	denotes	4
T12824	47514-47515	-RRB-	denotes	)
T12825	47516-47519	CC	denotes	and
T12826	47520-47521	DT	denotes	a
T12828	47522-47527	JJ	denotes	young
T12829	47528-47529	-LRB-	denotes	(
T12830	47529-47531	NN	denotes	P6
T12831	47531-47532	-RRB-	denotes	)
T12832	47533-47538	NN	denotes	C57BL
T12833	47538-47539	HYPH	denotes	/
T12834	47539-47540	CD	denotes	6
T12827	47541-47546	NN	denotes	mouse
T12835	47546-47547	.	denotes	.
T12836	47547-47742	sentence	denotes	RNA in situ hybridization was carried out as described previously [15,53] with digoxigenin-labeled antisense riboprobes specific for the 3' UTRs of genes AY318555 (0.5 kb) and AY317365 (0.5 kb).
T12837	47548-47551	NN	denotes	RNA
T12839	47552-47554	FW	denotes	in
T12840	47555-47559	FW	denotes	situ
T12838	47560-47573	NN	denotes	hybridization
T12842	47574-47577	VBD	denotes	was
T12841	47578-47585	VBN	denotes	carried
T12843	47586-47589	RP	denotes	out
T12844	47590-47592	IN	denotes	as
T12845	47593-47602	VBN	denotes	described
T12846	47603-47613	RB	denotes	previously
T12847	47614-47615	-LRB-	denotes	[
T12849	47615-47617	CD	denotes	15
T12850	47617-47618	,	denotes	,
T12848	47618-47620	CD	denotes	53
T12851	47620-47621	-RRB-	denotes	]
T12852	47622-47626	IN	denotes	with
T12853	47627-47638	NN	denotes	digoxigenin
T12855	47638-47639	HYPH	denotes	-
T12854	47639-47646	VBN	denotes	labeled
T12857	47647-47656	JJ	denotes	antisense
T12856	47657-47667	NNS	denotes	riboprobes
T12858	47668-47676	JJ	denotes	specific
T12859	47677-47680	IN	denotes	for
T12860	47681-47684	DT	denotes	the
T12862	47685-47686	CD	denotes	3
T12863	47686-47687	SYM	denotes	'
T12861	47688-47692	NNS	denotes	UTRs
T12864	47693-47695	IN	denotes	of
T12865	47696-47701	NNS	denotes	genes
T12866	47702-47710	NN	denotes	AY318555
T12867	47711-47712	-LRB-	denotes	(
T12869	47712-47715	CD	denotes	0.5
T12868	47716-47718	NN	denotes	kb
T12870	47718-47719	-RRB-	denotes	)
T12871	47720-47723	CC	denotes	and
T12872	47724-47732	NN	denotes	AY317365
T12873	47733-47734	-LRB-	denotes	(
T12875	47734-47737	CD	denotes	0.5
T12874	47738-47740	NN	denotes	kb
T12876	47740-47741	-RRB-	denotes	)
T12877	47741-47742	.	denotes	.
T12878	47742-47976	sentence	denotes	Riboprobe sequences were generated by PCR using primer pairs 5'-TCTTCCAAACCTGGACCCCCC-3' and 5'-ATCTCTCCAGCACCTTACTTG-3' for AY318555 and primer pairs 5'-TAAGATGTAAGTGATAATTTAGATTACAGG-3' and 5'-TTTCTGCCTCAGCTATGACAG-3' for AY317365.
T12879	47743-47752	NN	denotes	Riboprobe
T12880	47753-47762	NNS	denotes	sequences
T12882	47763-47767	VBD	denotes	were
T12881	47768-47777	VBN	denotes	generated
T12883	47778-47780	IN	denotes	by
T12884	47781-47784	NN	denotes	PCR
T12885	47785-47790	VBG	denotes	using
T12886	47791-47797	NN	denotes	primer
T12887	47798-47803	NNS	denotes	pairs
T12888	47804-47805	CD	denotes	5
T12890	47805-47806	SYM	denotes	'
T12891	47806-47807	HYPH	denotes	-
T12889	47807-47828	NN	denotes	TCTTCCAAACCTGGACCCCCC
T12892	47828-47829	HYPH	denotes	-
T12893	47829-47830	CD	denotes	3
T12894	47830-47831	SYM	denotes	'
T12895	47832-47835	CC	denotes	and
T12896	47836-47837	CD	denotes	5
T12898	47837-47838	SYM	denotes	'
T12899	47838-47839	HYPH	denotes	-
T12897	47839-47860	NN	denotes	ATCTCTCCAGCACCTTACTTG
T12900	47860-47861	HYPH	denotes	-
T12901	47861-47862	CD	denotes	3
T12902	47862-47863	SYM	denotes	'
T12903	47864-47867	IN	denotes	for
T12904	47868-47876	NN	denotes	AY318555
T12905	47877-47880	CC	denotes	and
T12906	47881-47887	NN	denotes	primer
T12907	47888-47893	NNS	denotes	pairs
T12908	47894-47895	CD	denotes	5
T12910	47895-47896	SYM	denotes	'
T12911	47896-47897	HYPH	denotes	-
T12909	47897-47927	NN	denotes	TAAGATGTAAGTGATAATTTAGATTACAGG
T12912	47927-47928	HYPH	denotes	-
T12913	47928-47929	CD	denotes	3
T12914	47929-47930	SYM	denotes	'
T12915	47931-47934	CC	denotes	and
T12916	47935-47936	CD	denotes	5
T12918	47936-47937	SYM	denotes	'
T12919	47937-47938	HYPH	denotes	-
T12917	47938-47959	NN	denotes	TTTCTGCCTCAGCTATGACAG
T12920	47959-47960	HYPH	denotes	-
T12921	47960-47961	CD	denotes	3
T12922	47961-47962	SYM	denotes	'
T12923	47963-47966	IN	denotes	for
T12924	47967-47975	NN	denotes	AY317365
T12925	47975-47976	.	denotes	.
T12926	47976-48093	sentence	denotes	Hybridization was carried out in 50% formamide at 65°C, and slides were washed at high stringency (65°C, 0.2 × SSC).
T12927	47977-47990	NN	denotes	Hybridization
T12929	47991-47994	VBD	denotes	was
T12928	47995-48002	VBN	denotes	carried
T12930	48003-48006	RP	denotes	out
T12931	48007-48009	IN	denotes	in
T12932	48010-48012	CD	denotes	50
T12933	48012-48013	NN	denotes	%
T12934	48014-48023	NN	denotes	formamide
T12935	48024-48026	IN	denotes	at
T12936	48027-48029	CD	denotes	65
T12937	48029-48031	NNS	denotes	°C
T12938	48031-48033	,	denotes	,
T12939	48033-48036	CC	denotes	and
T12940	48037-48043	NNS	denotes	slides
T12942	48044-48048	VBD	denotes	were
T12941	48049-48055	VBN	denotes	washed
T12943	48056-48058	IN	denotes	at
T12944	48059-48063	JJ	denotes	high
T12945	48064-48074	NN	denotes	stringency
T12946	48075-48076	-LRB-	denotes	(
T12948	48076-48078	CD	denotes	65
T12947	48078-48080	NNS	denotes	°C
T12949	48080-48082	,	denotes	,
T12950	48082-48085	CD	denotes	0.2
T12952	48086-48087	SYM	denotes	×
T12951	48088-48091	NN	denotes	SSC
T12953	48091-48092	-RRB-	denotes	)
T12954	48092-48093	.	denotes	.
T12955	48093-48225	sentence	denotes	The probes each hybridize to only one band on a Southern blot, indicating that each probe only detects one olfactory receptor gene.
T12956	48094-48097	DT	denotes	The
T12957	48098-48104	NNS	denotes	probes
T12959	48105-48109	DT	denotes	each
T12958	48110-48119	VBP	denotes	hybridize
T12960	48120-48122	IN	denotes	to
T12961	48123-48127	RB	denotes	only
T12963	48128-48131	CD	denotes	one
T12962	48132-48136	NN	denotes	band
T12964	48137-48139	IN	denotes	on
T12965	48140-48141	DT	denotes	a
T12967	48142-48150	NNP	denotes	Southern
T12966	48151-48155	NN	denotes	blot
T12968	48155-48157	,	denotes	,
T12969	48157-48167	VBG	denotes	indicating
T12970	48168-48172	IN	denotes	that
T12972	48173-48177	DT	denotes	each
T12973	48178-48183	NN	denotes	probe
T12974	48184-48188	RB	denotes	only
T12971	48189-48196	VBZ	denotes	detects
T12975	48197-48200	CD	denotes	one
T12977	48201-48210	JJ	denotes	olfactory
T12978	48211-48219	NN	denotes	receptor
T12976	48220-48224	NN	denotes	gene
T12979	48224-48225	.	denotes	.
T12980	48225-48400	sentence	denotes	BLAST analyses show that the AY318555 probe is unique in Celera's mouse genome assembly (Release 13), and that the AY317365 probe is similar to only one other genomic region.
T12981	48226-48231	NN	denotes	BLAST
T12982	48232-48240	NNS	denotes	analyses
T12983	48241-48245	VBP	denotes	show
T12984	48246-48250	IN	denotes	that
T12986	48251-48254	DT	denotes	the
T12988	48255-48263	NN	denotes	AY318555
T12987	48264-48269	NN	denotes	probe
T12985	48270-48272	VBZ	denotes	is
T12989	48273-48279	JJ	denotes	unique
T12990	48280-48282	IN	denotes	in
T12991	48283-48289	NNP	denotes	Celera
T12993	48289-48291	POS	denotes	's
T12994	48292-48297	NN	denotes	mouse
T12995	48298-48304	NN	denotes	genome
T12992	48305-48313	NN	denotes	assembly
T12996	48314-48315	-LRB-	denotes	(
T12997	48315-48322	NN	denotes	Release
T12998	48323-48325	CD	denotes	13
T12999	48325-48326	-RRB-	denotes	)
T13000	48326-48328	,	denotes	,
T13001	48328-48331	CC	denotes	and
T13002	48332-48336	IN	denotes	that
T13004	48337-48340	DT	denotes	the
T13006	48341-48349	NN	denotes	AY317365
T13005	48350-48355	NN	denotes	probe
T13003	48356-48358	VBZ	denotes	is
T13007	48359-48366	JJ	denotes	similar
T13008	48367-48369	IN	denotes	to
T13009	48370-48374	RB	denotes	only
T13011	48375-48378	CD	denotes	one
T13012	48379-48384	JJ	denotes	other
T13013	48385-48392	JJ	denotes	genomic
T13010	48393-48399	NN	denotes	region
T13014	48399-48400	.	denotes	.
T13015	48400-48585	sentence	denotes	This potential cross-hybridizing region is over 10 Mb from the nearest olfactory receptor coding region and is thus highly unlikely to be included in any olfactory receptor transcript.
T13016	48401-48405	DT	denotes	This
T13018	48406-48415	JJ	denotes	potential
T13019	48416-48433	JJ	denotes	cross-hybridizing
T13017	48434-48440	NN	denotes	region
T13020	48441-48443	VBZ	denotes	is
T13021	48444-48448	IN	denotes	over
T13022	48449-48451	CD	denotes	10
T13023	48452-48454	NNS	denotes	Mb
T13024	48455-48459	IN	denotes	from
T13025	48460-48463	DT	denotes	the
T13027	48464-48471	JJS	denotes	nearest
T13028	48472-48481	JJ	denotes	olfactory
T13029	48482-48490	NN	denotes	receptor
T13030	48491-48497	NN	denotes	coding
T13026	48498-48504	NN	denotes	region
T13031	48505-48508	CC	denotes	and
T13032	48509-48511	VBZ	denotes	is
T13033	48512-48516	RB	denotes	thus
T13034	48517-48523	RB	denotes	highly
T13035	48524-48532	JJ	denotes	unlikely
T13036	48533-48535	TO	denotes	to
T13038	48536-48538	VB	denotes	be
T13037	48539-48547	VBN	denotes	included
T13039	48548-48550	IN	denotes	in
T13040	48551-48554	DT	denotes	any
T13042	48555-48564	JJ	denotes	olfactory
T13043	48565-48573	NN	denotes	receptor
T13041	48574-48584	NN	denotes	transcript
T13044	48584-48585	.	denotes	.
T13045	48585-48688	sentence	denotes	Low-power images are composed of three overlapping micrographs (40×) assembled in Adobe Photoshop 7.0.
T13046	48586-48589	JJ	denotes	Low
T13048	48589-48590	HYPH	denotes	-
T13047	48590-48595	NN	denotes	power
T13049	48596-48602	NNS	denotes	images
T13051	48603-48606	VBP	denotes	are
T13050	48607-48615	VBN	denotes	composed
T13052	48616-48618	IN	denotes	of
T13053	48619-48624	CD	denotes	three
T13055	48625-48636	VBG	denotes	overlapping
T13054	48637-48648	NNS	denotes	micrographs
T13056	48649-48650	-LRB-	denotes	(
T13057	48650-48652	CD	denotes	40
T13058	48652-48653	SYM	denotes	×
T13059	48653-48654	-RRB-	denotes	)
T13060	48655-48664	VBN	denotes	assembled
T13061	48665-48667	IN	denotes	in
T13062	48668-48673	NNP	denotes	Adobe
T13063	48674-48683	NN	denotes	Photoshop
T13064	48684-48687	CD	denotes	7.0
T13065	48687-48688	.	denotes	.
T13084	48712-48713	DT	denotes	A
T13085	48714-48718	NN	denotes	list
T13087	48719-48721	IN	denotes	of
T13088	48722-48725	DT	denotes	the
T13089	48726-48733	NNS	denotes	primers
T13090	48734-48738	VBN	denotes	used
T13091	48739-48741	TO	denotes	to
T13092	48742-48749	VB	denotes	confirm
T13093	48750-48753	DT	denotes	the
T13094	48754-48764	NN	denotes	expression
T13095	48765-48767	IN	denotes	of
T13096	48768-48777	JJ	denotes	olfactory
T13097	48778-48786	NN	denotes	receptor
T13098	48787-48792	NNS	denotes	genes
T13099	48793-48795	IN	denotes	by
T13100	48796-48798	NN	denotes	RT
T13102	48798-48799	HYPH	denotes	-
T13101	48799-48802	NN	denotes	PCR
T13103	48803-48806	CC	denotes	and
T13104	48807-48810	NN	denotes	PCR
T13105	48811-48815	IN	denotes	from
T13106	48816-48820	NN	denotes	cDNA
T13108	48821-48828	NN	denotes	library
T13107	48829-48838	NNS	denotes	templates
T13109	48839-48842	MD	denotes	can
T13110	48843-48845	VB	denotes	be
T13086	48846-48851	VBN	denotes	found
T13111	48852-48854	IN	denotes	in
T13112	48855-48865	JJ	denotes	Additional
T13114	48866-48870	NNS	denotes	data
T13113	48871-48875	NN	denotes	file
T13115	48876-48877	CD	denotes	1
T13116	48877-48878	.	denotes	.
T13117	48878-48969	sentence	denotes	The experimental conditions used for real-time PCR can be found in Additional data file 2.
T13118	48879-48882	DT	denotes	The
T13120	48883-48895	JJ	denotes	experimental
T13119	48896-48906	NNS	denotes	conditions
T13122	48907-48911	VBN	denotes	used
T13123	48912-48915	IN	denotes	for
T13124	48916-48920	JJ	denotes	real
T13126	48920-48921	HYPH	denotes	-
T13125	48921-48925	NN	denotes	time
T13127	48926-48929	NN	denotes	PCR
T13128	48930-48933	MD	denotes	can
T13129	48934-48936	VB	denotes	be
T13121	48937-48942	VBN	denotes	found
T13130	48943-48945	IN	denotes	in
T13131	48946-48956	JJ	denotes	Additional
T13133	48957-48961	NNS	denotes	data
T13132	48962-48966	NN	denotes	file
T13134	48967-48968	CD	denotes	2
T13135	48968-48969	.	denotes	.
R10	T306	T301	prep	for,is
R100	T405	T406	nsubj	Differences,are
R1000	T1915	T1884	conj	feedback,send
R1001	T1916	T1917	aux	to,enhance
R1002	T1917	T1915	advcl	enhance,feedback
R1003	T1918	T1919	poss	its,expression
R1004	T1919	T1917	dobj	expression,enhance
R1005	T1920	T1919	amod	own,expression
R1006	T1921	T1884	punct	.,send
R1007	T2010	T2008	pobj	genes,for
R1008	T1923	T1924	prep	In,share
R1009	T2011	T2001	cc	and,sequences
R101	T407	T405	prep	for,Differences
R1010	T1925	T1926	det	the,models
R1011	T1926	T1923	pobj	models,In
R1012	T1927	T1926	amod	latter,models
R1013	T2012	T2001	punct	", ",sequences
R1014	T1928	T1926	nummod	three,models
R1015	T2013	T2014	advmod	therefore,dataset
R1016	T1929	T1924	punct	", ",share
R1017	T2014	T2001	conj	dataset,sequences
R1018	T1930	T1924	nsubj	some,share
R1019	T1931	T1930	cc	or,some
R102	T408	T409	nummod	one,pair
R1020	T1932	T1933	nmod	all,genes
R1021	T2015	T2014	punct	", ",dataset
R1022	T1933	T1930	conj	genes,some
R1023	T1934	T1935	amod	olfactory,receptor
R1024	T1935	T1933	compound	receptor,genes
R1025	T2016	T2014	det	a,dataset
R1026	T1936	T1924	aux	might,share
R1027	T1937	T1938	compound	transcription,factor
R1028	T1938	T1939	compound	factor,binding
R1029	T2017	T2014	amod	large,dataset
R103	T409	T407	pobj	pair,for
R1030	T1939	T1940	compound	binding,motifs
R1031	T1940	T1924	dobj	motifs,share
R1032	T2018	T2014	prep	of,dataset
R1033	T1941	T1924	punct	", ",share
R1034	T1942	T1924	cc	and,share
R1035	T1943	T1944	prep	in,share
R1036	T2019	T2020	compound	candidate,promoter
R1037	T1944	T1924	conj	share,share
R1038	T1945	T1946	det	the,model
R1039	T2020	T2021	compound	promoter,regions
R104	T410	T409	compound	gene,pair
R1040	T1946	T1943	pobj	model,in
R1041	T1947	T1946	amod	first,model
R1042	T2021	T2018	pobj	regions,of
R1043	T1948	T1944	punct	", ",share
R1044	T1949	T1950	amod	olfactory,receptor
R1045	T2022	T1999	punct	.,provides
R1046	T1950	T1951	compound	receptor,genes
R1047	T1951	T1944	nsubj	genes,share
R1048	T2024	T2025	amod	Olfactory,receptor
R1049	T1952	T1944	aux	might,share
R105	T411	T406	advmod	apparently,are
R1050	T1953	T1954	det	a,signal
R1051	T1954	T1944	dobj	signal,share
R1052	T1955	T1954	amod	common,signal
R1053	T1956	T1954	compound	recombination,signal
R1054	T1957	T1944	punct	.,share
R1055	T2025	T2026	compound	receptor,genes
R1056	T1959	T1960	prep	In,is
R1057	T1961	T1959	pobj	order,In
R1058	T2026	T2027	nsubj	genes,have
R1059	T1962	T1963	aux	to,perform
R106	T412	T406	prep	due,are
R1060	T1963	T1961	acl	perform,order
R1061	T1964	T1965	amod	computational,searches
R1062	T2028	T2029	det	an,region
R1063	T1965	T1963	dobj	searches,perform
R1064	T1966	T1964	cc	and,computational
R1065	T1967	T1964	conj	experimental,computational
R1066	T1968	T1965	prep	for,searches
R1067	T2029	T2027	dobj	region,have
R1068	T1969	T1970	amod	such,signals
R1069	T1970	T1968	pobj	signals,for
R107	T413	T412	pcomp	to,due
R1070	T1971	T1960	punct	", ",is
R1071	T2030	T2029	amod	intronless,region
R1072	T1972	T1960	nsubj	it,is
R1073	T1973	T1960	acomp	important,is
R1074	T2031	T2029	amod	coding,region
R1075	T1974	T1975	aux	to,have
R1076	T1975	T1960	xcomp	have,is
R1077	T1976	T1977	det	a,idea
R1078	T2032	T2027	punct	", ",have
R1079	T1977	T1975	dobj	idea,have
R108	T414	T415	preconj	both,numbers
R1080	T1978	T1977	amod	better,idea
R1081	T1979	T1977	prep	of,idea
R1082	T1980	T1981	det	the,site
R1083	T2033	T2027	advcl	simplifying,have
R1084	T1981	T1979	pobj	site,of
R1085	T1982	T1981	amod	transcriptional,site
R1086	T2034	T2035	preconj	both,computational
R1087	T1983	T1981	compound	start,site
R1088	T1984	T1981	prep	of,site
R1089	T1985	T1986	det	a,number
R109	T415	T412	pobj	numbers,due
R1090	T2035	T2036	amod	computational,identification
R1091	T1986	T1984	pobj	number,of
R1092	T1987	T1986	amod	large,number
R1093	T1988	T1986	prep	of,number
R1094	T2036	T2033	dobj	identification,simplifying
R1095	T1989	T1990	amod	olfactory,receptor
R1096	T1990	T1991	compound	receptor,genes
R1097	T2037	T2035	cc	and,computational
R1098	T1991	T1988	pobj	genes,of
R1099	T1992	T1960	punct	.,is
R11	T307	T308	advmod	very,few
R110	T416	T415	amod	unequal,numbers
R1100	T2038	T2035	conj	experimental,computational
R1101	T1994	T1995	poss	Our,collection
R1102	T1995	T1999	nsubj	collection,provides
R1103	T1996	T1997	amod	olfactory,receptor
R1104	T1997	T1995	compound	receptor,collection
R1105	T1998	T1995	compound	EST,collection
R1106	T2039	T2040	amod	olfactory,receptor
R1107	T2000	T2001	nummod	5,sequences
R1108	T2001	T1999	dobj	sequences,provides
R1109	T2040	T2036	compound	receptor,identification
R111	T417	T415	prep	of,numbers
R1110	T2002	T2000	punct	',5
R1111	T2003	T2004	amod	untranslated,region
R1112	T2004	T2001	nmod	region,sequences
R1113	T2041	T2027	punct	.,have
R1114	T2005	T2004	punct	(,region
R1115	T2006	T2004	appos	UTR,region
R1116	T2007	T2001	punct	),sequences
R1117	T2008	T2001	prep	for,sequences
R1118	T2043	T2044	prep	For,determined
R1119	T2045	T2046	det	a,number
R112	T418	T419	amod	expressing,cells
R1120	T2046	T2043	pobj	number,For
R1121	T2115	T2116	punct	-,27
R1122	T2047	T2046	amod	small,number
R1123	T2116	T2114	prep	27,25
R1124	T2117	T2114	punct	],25
R1125	T2118	T2094	punct	.,described
R1126	T2048	T2046	prep	of,number
R1127	T2120	T2121	compound	cDNA,identification
R1128	T2049	T2050	amod	olfactory,receptors
R1129	T2121	T2122	nsubjpass	identification,used
R113	T419	T417	pobj	cells,of
R1130	T2123	T2121	cc	and,identification
R1131	T2050	T2048	pobj	receptors,of
R1132	T2124	T2125	compound	RACE,data
R1133	T2125	T2121	conj	data,identification
R1134	T2126	T2122	aux	have,used
R1135	T2051	T2044	punct	", ",determined
R1136	T2127	T2122	auxpass	been,used
R1137	T2128	T2129	aux	to,determine
R1138	T2129	T2122	advcl	determine,used
R1139	T2052	T2053	compound	gene,structure
R114	T420	T415	cc	and,numbers
R1140	T2130	T2131	compound	gene,structure
R1141	T2131	T2129	dobj	structure,determine
R1142	T2132	T2129	prep	for,determine
R1143	T2053	T2044	nsubjpass	structure,determined
R1144	T2133	T2134	advmod	about,30
R1145	T2134	T2135	nummod	30,genes
R1146	T2054	T2044	aux	has,determined
R1147	T2135	T2132	pobj	genes,for
R1148	T2136	T2137	punct	", ",see
R1149	T2137	T2122	parataxis	see,used
R115	T421	T422	amod	unequal,levels
R1150	T2055	T2044	auxpass	been,determined
R1151	T2138	T2137	punct	", ",see
R1152	T2139	T2140	prep	for,23
R1153	T2140	T2137	dobj	23,see
R1154	T2056	T2044	punct	.,determined
R1155	T2141	T2139	pobj	example,for
R1156	T2142	T2140	punct	", ",23
R1157	T2143	T2140	punct	[,23
R1158	T2144	T2140	nummod	19,23
R1159	T2145	T2140	punct	",",23
R116	T422	T415	conj	levels,numbers
R1160	T2058	T2059	amod	Additional,exons
R1161	T2146	T2137	punct	],see
R1162	T2147	T2122	punct	.,used
R1163	T2149	T2150	advmod	However,been
R1164	T2059	T2063	nsubj	exons,lie
R1165	T2151	T2150	punct	", ",been
R1166	T2152	T2153	amod	computational,prediction
R1167	T2153	T2150	nsubj	prediction,been
R1168	T2060	T2059	nummod	5,exons
R1169	T2154	T2153	prep	of,prediction
R117	T423	T422	compound	transcript,levels
R1170	T2155	T2156	det	the,location
R1171	T2156	T2154	pobj	location,of
R1172	T2061	T2060	punct	',5
R1173	T2157	T2156	prep	of,location
R1174	T2158	T2159	nummod	5,exons
R1175	T2159	T2157	pobj	exons,of
R1176	T2062	T2059	amod	untranslated,exons
R1177	T2160	T2158	punct	',5
R1178	T2161	T2159	amod	upstream,exons
R1179	T2162	T2153	cc	and,prediction
R118	T424	T422	prep	per,levels
R1180	T2064	T2063	advmod	upstream,lie
R1181	T2163	T2164	det	the,extent
R1182	T2164	T2153	conj	extent,prediction
R1183	T2165	T2164	prep	of,extent
R1184	T2166	T2167	det	the,UTR
R1185	T2065	T2064	prep	of,upstream
R1186	T2167	T2165	pobj	UTR,of
R1187	T2168	T2167	nummod	3,UTR
R1188	T2066	T2067	det	the,region
R1189	T2169	T2168	punct	',3
R119	T425	T426	amod	expressing,cell
R1190	T2170	T2153	prep	from,prediction
R1191	T2171	T2172	amod	genomic,sequence
R1192	T2067	T2065	pobj	region,of
R1193	T2172	T2170	pobj	sequence,from
R1194	T2173	T2150	aux	has,been
R1195	T2174	T2175	advmod	extremely,difficult
R1196	T2068	T2067	amod	coding,region
R1197	T2175	T2150	acomp	difficult,been
R1198	T2176	T2150	punct	.,been
R1199	T2069	T2063	cc	and,lie
R12	T308	T309	amod	few,receptors
R120	T426	T424	pobj	cell,per
R1200	T2178	T2179	det	A,combination
R1201	T2179	T2180	nsubj	combination,allowed
R1202	T2070	T2071	aux	can,spliced
R1203	T2181	T2179	prep	of,combination
R1204	T2182	T2183	compound	splice,site
R1205	T2071	T2063	conj	spliced,lie
R1206	T2183	T2184	compound	site,predictions
R1207	T2184	T2181	pobj	predictions,of
R1208	T2185	T2184	cc	and,predictions
R1209	T2186	T2184	conj	similarity,predictions
R121	T427	T406	punct	.,are
R1210	T2187	T2186	prep	to,similarity
R1211	T2188	T2189	amod	other,receptors
R1212	T2072	T2071	auxpass	be,spliced
R1213	T2189	T2187	pobj	receptors,to
R1214	T2190	T2189	amod	olfactory,receptors
R1215	T2073	T2071	advmod	alternatively,spliced
R1216	T2074	T2075	punct	[,23
R1217	T2191	T2180	aux	has,allowed
R1218	T2075	T2071	parataxis	23,spliced
R1219	T2192	T2193	det	some,investigators
R122	T429	T430	advmod	At,two
R1220	T2076	T2075	dep	19,23
R1221	T2193	T2194	nsubj	investigators,predict
R1222	T2194	T2180	ccomp	predict,allowed
R1223	T2077	T2075	punct	",",23
R1224	T2195	T2194	aux	to,predict
R1225	T2196	T2197	nummod	5,exon
R1226	T2197	T2199	compound	exon,locations
R1227	T2078	T2079	punct	-,26
R1228	T2198	T2196	punct	',5
R1229	T2199	T2194	dobj	locations,predict
R123	T430	T432	nsubj	two,exhibit
R1230	T2200	T2194	prep	for,predict
R1231	T2079	T2075	prep	26,23
R1232	T2201	T2202	advmod	around,15
R1233	T2202	T2203	nummod	15,genes
R1234	T2080	T2075	punct	],23
R1235	T2203	T2200	pobj	genes,for
R1236	T2204	T2205	punct	[,28
R1237	T2205	T2180	parataxis	28,allowed
R1238	T2206	T2205	nummod	25,28
R1239	T2081	T2063	punct	.,lie
R124	T431	T430	advmod	least,two
R1240	T2207	T2205	punct	",",28
R1241	T2208	T2205	punct	],28
R1242	T2209	T2180	punct	.,allowed
R1243	T2083	T2084	det	The,region
R1244	T2211	T2212	amod	Experimental,validation
R1245	T2212	T2213	nsubj	validation,shows
R1246	T2084	T2088	nsubj	region,is
R1247	T2214	T2215	mark	that,are
R1248	T2215	T2213	ccomp	are,shows
R1249	T2085	T2084	nummod	3,region
R125	T433	T430	punct	-,two
R1250	T2216	T2215	nsubj	some,are
R1251	T2217	T2216	punct	", ",some
R1252	T2218	T2216	cc	but,some
R1253	T2086	T2085	punct	',3
R1254	T2219	T2218	neg	not,but
R1255	T2220	T2216	conj	all,some
R1256	T2087	T2084	amod	untranslated,region
R1257	T2089	T2088	advmod	typically,is
R1258	T2090	T2088	acomp	intronless,is
R1259	T2221	T2220	punct	", ",all
R126	T434	T430	quantmod	thirds,two
R1260	T2222	T2220	conj	predictions,all
R1261	T2223	T2215	acomp	accurate,are
R1262	T2224	T2225	punct	[,25
R1263	T2091	T2088	punct	.,is
R1264	T2225	T2213	parataxis	25,shows
R1265	T2226	T2225	nummod	24,25
R1266	T2227	T2225	punct	",",25
R1267	T2093	T2094	nsubjpass	Exceptions,described
R1268	T2228	T2225	punct	],25
R1269	T2229	T2213	punct	.,shows
R127	T435	T430	prep	of,two
R1270	T2231	T2232	det	The,number
R1271	T2095	T2093	prep	to,Exceptions
R1272	T2232	T2234	nsubjpass	number,increased
R1273	T2233	T2232	amod	total,number
R1274	T2235	T2232	prep	of,number
R1275	T2096	T2097	det	this,structure
R1276	T2236	T2237	amod	olfactory,receptors
R1277	T2237	T2235	pobj	receptors,of
R1278	T2238	T2239	prep	for,known
R1279	T2097	T2095	pobj	structure,to
R128	T436	T437	amod	olfactory,receptors
R1280	T2239	T2232	relcl	known,number
R1281	T2240	T2238	pobj	which,for
R1282	T2098	T2097	amod	stereotyped,structure
R1283	T2241	T2242	compound	gene,structure
R1284	T2242	T2239	nsubjpass	structure,known
R1285	T2243	T2239	auxpass	is,known
R1286	T2099	T2094	aux	have,described
R1287	T2244	T2234	auxpass	is,increased
R1288	T2245	T2234	advmod	vastly,increased
R1289	T2246	T2234	agent	by,increased
R129	T437	T435	pobj	receptors,of
R1290	T2100	T2094	auxpass	been,described
R1291	T2247	T2248	poss	our,study
R1292	T2248	T2246	pobj	study,by
R1293	T2249	T2234	punct	.,increased
R1294	T2101	T2094	prep	for,described
R1295	T2251	T2252	prep	In,describe
R1296	T2102	T2103	det	some,receptors
R1297	T2253	T2254	det	this,report
R1298	T2254	T2251	pobj	report,In
R1299	T2103	T2101	pobj	receptors,for
R13	T309	T306	pobj	receptors,for
R130	T438	T439	amod	multiple,variants
R1300	T2255	T2252	punct	", ",describe
R1301	T2256	T2252	nsubj	we,describe
R1302	T2104	T2103	amod	human,receptors
R1303	T2257	T2258	det	the,isolation
R1304	T2258	T2252	dobj	isolation,describe
R1305	T2259	T2258	cc	and,isolation
R1306	T2105	T2103	amod	olfactory,receptors
R1307	T2260	T2258	conj	analysis,isolation
R1308	T2261	T2258	prep	of,isolation
R1309	T2262	T2263	quantmod	over,"1,200"
R131	T439	T432	dobj	variants,exhibit
R1310	T2106	T2094	punct	", ",described
R1311	T2263	T2264	nummod	"1,200",cDNAs
R1312	T2264	T2261	pobj	cDNAs,of
R1313	T2265	T2264	acl	representing,cDNAs
R1314	T2266	T2267	nummod	419,genes
R1315	T2267	T2265	dobj	genes,representing
R1316	T2268	T2269	amod	odorant,receptor
R1317	T2107	T2094	cc	but,described
R1318	T2269	T2267	compound	receptor,genes
R1319	T2270	T2252	punct	.,describe
R132	T440	T439	amod	transcriptional,variants
R1320	T2108	T2109	auxpass	are,thought
R1321	T2272	T2273	nsubj	We,screened
R1322	T2109	T2094	conj	thought,described
R1323	T2274	T2275	det	a,library
R1324	T2275	T2273	dobj	library,screened
R1325	T2110	T2111	aux	to,be
R1326	T2276	T2277	nmod	mouse,epithelium
R1327	T2277	T2275	compound	epithelium,library
R1328	T2278	T2277	amod	olfactory,epithelium
R1329	T2111	T2109	xcomp	be,thought
R133	T441	T439	punct	", ",variants
R1330	T2279	T2273	prep	with,screened
R1331	T2280	T2281	amod	degenerate,probes
R1332	T2112	T2111	acomp	rare,be
R1333	T2113	T2114	punct	[,25
R1334	T2281	T2279	pobj	probes,with
R1335	T2114	T2109	parataxis	25,thought
R1336	T2282	T2283	amod	olfactory,receptor
R1337	T2283	T2281	compound	receptor,probes
R1338	T2284	T2273	cc	and,screened
R1339	T2327	T2314	conj	show,demonstrate
R134	T442	T439	prep	with,variants
R1340	T2285	T2273	conj	obtained,screened
R1341	T2286	T2285	dobj	5,obtained
R1342	T2287	T2286	punct	',5
R1343	T2288	T2289	compound	end,sequences
R1344	T2328	T2329	mark	that,have
R1345	T2289	T2286	appos	sequences,5
R1346	T2290	T2289	punct	(,sequences
R1347	T2329	T2327	ccomp	have,show
R1348	T2291	T2289	appos	ESTs,sequences
R1349	T2292	T2285	punct	),obtained
R135	T443	T444	amod	alternative,isoforms
R1350	T2330	T2331	amod	most,genes
R1351	T2293	T2285	prep	from,obtained
R1352	T2294	T2295	amod	purified,cDNAs
R1353	T2295	T2293	pobj	cDNAs,from
R1354	T2331	T2329	nsubj	genes,have
R1355	T2296	T2273	punct	.,screened
R1356	T2332	T2333	amod	olfactory,receptor
R1357	T2298	T2299	det	These,clones
R1358	T2299	T2300	nsubj	clones,confirm
R1359	T2333	T2331	compound	receptor,genes
R136	T444	T442	pobj	isoforms,with
R1360	T2301	T2302	amod	olfactory,expression
R1361	T2302	T2300	dobj	expression,confirm
R1362	T2334	T2335	amod	multiple,isoforms
R1363	T2303	T2302	prep	of,expression
R1364	T2304	T2305	quantmod	over,400
R1365	T2305	T2306	nummod	400,receptors
R1366	T2306	T2303	pobj	receptors,of
R1367	T2307	T2306	amod	olfactory,receptors
R1368	T2308	T2300	punct	", ",confirm
R1369	T2309	T2300	conj	provide,confirm
R137	T445	T444	prep	of,isoforms
R1370	T2335	T2329	dobj	isoforms,have
R1371	T2310	T2311	poss	their,structure
R1372	T2311	T2309	dobj	structure,provide
R1373	T2336	T2335	amod	transcriptional,isoforms
R1374	T2312	T2311	compound	gene,structure
R1375	T2313	T2309	punct	", ",provide
R1376	T2314	T2309	conj	demonstrate,provide
R1377	T2337	T2300	punct	.,confirm
R1378	T2315	T2316	mark	that,expressed
R1379	T2316	T2314	ccomp	expressed,demonstrate
R138	T446	T447	preconj	both,5
R1380	T2317	T2318	neg	not,receptors
R1381	T2318	T2316	nsubjpass	receptors,expressed
R1382	T2319	T2318	det	all,receptors
R1383	T2320	T2318	amod	olfactory,receptors
R1384	T2321	T2316	auxpass	are,expressed
R1385	T2322	T2316	prep	at,expressed
R1386	T2323	T2324	det	the,level
R1387	T2324	T2322	pobj	level,at
R1388	T2325	T2324	amod	same,level
R1389	T2326	T2314	cc	and,demonstrate
R139	T447	T448	nummod	5,regions
R1390	T2624	T2625	advmod	At,419
R1391	T2625	T2627	nummod	419,genes
R1392	T2626	T2625	advmod	least,419
R1393	T2627	T2631	nsubjpass	genes,expressed
R1394	T2628	T2627	nmod	mouse,genes
R1395	T2629	T2630	amod	olfactory,receptor
R1396	T2630	T2627	compound	receptor,genes
R1397	T2632	T2631	auxpass	are,expressed
R1398	T2633	T2631	prep	in,expressed
R1399	T2634	T2635	det	the,epithelium
R14	T310	T309	amod	olfactory,receptors
R140	T448	T445	pobj	regions,of
R1400	T2635	T2633	pobj	epithelium,in
R1401	T2636	T2635	amod	olfactory,epithelium
R1402	T2638	T2639	nsubj	We,isolated
R1403	T2640	T2639	aux	have,isolated
R1404	T2641	T2642	nummod	"1,264",clones
R1405	T2642	T2639	dobj	clones,isolated
R1406	T2643	T2644	amod	olfactory,receptor
R1407	T2644	T2645	compound	receptor,cDNA
R1408	T2645	T2642	compound	cDNA,clones
R1409	T2646	T2639	punct	", ",isolated
R141	T449	T447	punct	',5
R1410	T2647	T2648	dep	which,confirm
R1411	T2648	T2639	ccomp	confirm,isolated
R1412	T2649	T2648	advmod	together,confirm
R1413	T2650	T2651	det	the,expression
R1414	T2651	T2648	dobj	expression,confirm
R1415	T2652	T2653	amod	olfactory,epithelial
R1416	T2653	T2651	amod	epithelial,expression
R1417	T2654	T2651	prep	of,expression
R1418	T2655	T2656	nummod	419,genes
R1419	T2656	T2654	pobj	genes,of
R142	T450	T447	cc	and,5
R1420	T2657	T2656	amod	annotated,genes
R1421	T2658	T2659	amod	olfactory,receptor
R1422	T2659	T2656	compound	receptor,genes
R1423	T2660	T2639	punct	.,isolated
R1424	T2662	T2663	nsubj	We,used
R1425	T2664	T2665	amod	low,stringency
R1426	T2665	T2667	compound	stringency,hybridization
R1427	T2666	T2665	punct	-,stringency
R1428	T2667	T2663	dobj	hybridization,used
R1429	T2668	T2667	prep	with,hybridization
R143	T451	T447	conj	3,5
R1430	T2669	T2670	amod	degenerate,probes
R1431	T2670	T2668	pobj	probes,with
R1432	T2671	T2672	amod	olfactory,receptor
R1433	T2672	T2670	compound	receptor,probes
R1434	T2673	T2670	compound	DNA,probes
R1435	T2674	T2675	aux	to,screen
R1436	T2675	T2663	advcl	screen,used
R1437	T2676	T2677	advmod	around,"4,100,000"
R1438	T2677	T2678	nummod	"4,100,000",units
R1439	T2678	T2675	dobj	units,screen
R144	T452	T451	punct	',3
R1440	T2679	T2680	npadvmod	plaque,forming
R1441	T2680	T2678	amod	forming,units
R1442	T2681	T2680	punct	-,forming
R1443	T2682	T2678	punct	(,units
R1444	T2683	T2678	appos	pfu,units
R1445	T2684	T2678	punct	),units
R1446	T2685	T2678	prep	of,units
R1447	T2686	T2687	det	an,library
R1448	T2687	T2685	pobj	library,of
R1449	T2688	T2689	amod	adult,mouse
R145	T453	T448	amod	untranslated,regions
R1450	T2689	T2690	nmod	mouse,cDNA
R1451	T2690	T2687	compound	cDNA,library
R1452	T2691	T2692	amod	olfactory,epithelium
R1453	T2692	T2690	compound	epithelium,cDNA
R1454	T2693	T2678	cc	and,units
R1455	T2694	T2695	advmod	around,"640,000"
R1456	T2695	T2696	nummod	"640,000",pfu
R1457	T2696	T2678	conj	pfu,units
R1458	T2697	T2696	prep	of,pfu
R1459	T2698	T2699	det	an,library
R146	T454	T432	punct	.,exhibit
R1460	T2699	T2697	pobj	library,of
R1461	T2700	T2701	amod	embryonic,epithelium
R1462	T2701	T2699	compound	epithelium,library
R1463	T2702	T2701	amod	olfactory,epithelium
R1464	T2703	T2663	punct	.,used
R1465	T2705	T2706	nsubj	We,obtained
R1466	T2707	T2706	dobj	sequences,obtained
R1467	T2708	T2706	prep	from,obtained
R1468	T2709	T2710	nummod	"1,715",cDNAs
R1469	T2710	T2708	pobj	cDNAs,from
R147	T456	T457	det	Some,transcripts
R1470	T2711	T2712	npadvmod	hybridization,positive
R1471	T2712	T2710	amod	positive,cDNAs
R1472	T2713	T2712	punct	-,positive
R1473	T2714	T2706	prep	following,obtained
R1474	T2715	T2716	amod	secondary,screens
R1475	T2716	T2714	pobj	screens,following
R1476	T2717	T2718	aux	to,isolate
R1477	T2718	T2716	advcl	isolate,screens
R1478	T2719	T2720	amod	single,clones
R1479	T2720	T2718	dobj	clones,isolate
R148	T457	T458	nsubj	transcripts,utilize
R1480	T2721	T2706	punct	.,obtained
R1481	T2723	T2724	prep	Of,yielded
R1482	T2725	T2726	det	these,clones
R1483	T2726	T2723	pobj	clones,Of
R1484	T2727	T2724	punct	", ",yielded
R1485	T2728	T2724	nsubj	"1,264",yielded
R1486	T2729	T2730	amod	olfactory,receptor
R1487	T2730	T2731	npadvmod	receptor,containing
R1488	T2731	T2733	amod	containing,sequences
R1489	T2732	T2731	punct	-,containing
R149	T459	T460	punct	(,%
R1490	T2733	T2724	dobj	sequences,yielded
R1491	T2734	T2724	punct	.,yielded
R1492	T2736	T2737	det	The,rate
R1493	T2737	T2743	nsubj	rate,is
R1494	T2738	T2739	nummod	26,%
R1495	T2739	T2737	nmod	%,rate
R1496	T2740	T2741	amod	false,positive
R1497	T2741	T2737	amod	positive,rate
R1498	T2742	T2741	punct	-,positive
R1499	T2744	T2745	det	a,consequence
R15	T311	T301	punct	.,is
R150	T460	T457	parataxis	%,transcripts
R1500	T2745	T2743	attr	consequence,is
R1501	T2746	T2745	prep	of,consequence
R1502	T2747	T2746	pcomp	using,of
R1503	T2748	T2749	amod	low,stringency
R1504	T2749	T2751	compound	stringency,hybridization
R1505	T2750	T2749	punct	-,stringency
R1506	T2751	T2747	dobj	hybridization,using
R1507	T2752	T2753	aux	to,obtain
R1508	T2753	T2747	advcl	obtain,using
R1509	T2754	T2755	amod	maximal,sensitivity
R151	T461	T460	nummod	5,%
R1510	T2755	T2753	dobj	sensitivity,obtain
R1511	T2756	T2743	punct	.,is
R1512	T2758	T2759	csubj	Continuing,resulted
R1513	T2759	T2764	ccomp	resulted,represented
R1514	T2760	T2761	det	the,screen
R1515	T2761	T2758	dobj	screen,Continuing
R1516	T2762	T2759	aux	would,resulted
R1517	T2763	T2759	aux	have,resulted
R1518	T2765	T2759	prep	in,resulted
R1519	T2766	T2765	pobj	cDNAs,in
R152	T462	T460	punct	),%
R1520	T2767	T2766	prep	from,cDNAs
R1521	T2768	T2769	amod	additional,receptors
R1522	T2769	T2767	pobj	receptors,from
R1523	T2770	T2769	amod	olfactory,receptors
R1524	T2771	T2759	punct	", ",resulted
R1525	T2772	T2759	cc	but,resulted
R1526	T2773	T2774	nsubj	we,reached
R1527	T2774	T2759	conj	reached,resulted
R1528	T2775	T2776	det	a,point
R1529	T2776	T2774	dobj	point,reached
R153	T463	T464	compound	splice,sites
R1530	T2777	T2776	prep	of,point
R1531	T2778	T2779	amod	limiting,returns
R1532	T2779	T2777	pobj	returns,of
R1533	T2780	T2764	punct	: ,represented
R1534	T2781	T2782	poss	our,batch
R1535	T2782	T2764	nsubj	batch,represented
R1536	T2783	T2782	amod	final,batch
R1537	T2784	T2782	prep	of,batch
R1538	T2785	T2786	nummod	45,sequences
R1539	T2786	T2784	pobj	sequences,of
R154	T464	T458	dobj	sites,utilize
R1540	T2787	T2788	amod	olfactory,receptor
R1541	T2788	T2789	npadvmod	receptor,positive
R1542	T2789	T2786	amod	positive,sequences
R1543	T2790	T2789	punct	-,positive
R1544	T2791	T2792	nummod	33,receptors
R1545	T2792	T2764	dobj	receptors,represented
R1546	T2793	T2792	amod	different,receptors
R1547	T2794	T2792	amod	olfactory,receptors
R1548	T2795	T2792	punct	", ",receptors
R1549	T2796	T2797	prep	of,encountered
R155	T465	T464	prep	within,sites
R1550	T2797	T2792	relcl	encountered,receptors
R1551	T2798	T2796	pobj	which,of
R1552	T2799	T2800	advmod	only,four
R1553	T2800	T2797	nsubjpass	four,encountered
R1554	T2801	T2797	aux	had,encountered
R1555	T2802	T2797	neg	not,encountered
R1556	T2803	T2797	auxpass	been,encountered
R1557	T2804	T2797	advmod	previously,encountered
R1558	T2805	T2797	prep	in,encountered
R1559	T2806	T2807	poss	our,screen
R156	T466	T467	det	the,region
R1560	T2807	T2805	pobj	screen,in
R1561	T2808	T2764	punct	.,represented
R1562	T2810	T2811	compound	Sequence,analysis
R1563	T2811	T2812	nsubj	analysis,shows
R1564	T2813	T2814	mark	that,are
R1565	T2814	T2812	ccomp	are,shows
R1566	T2815	T2816	det	the,libraries
R1567	T2816	T2814	nsubj	libraries,are
R1568	T2817	T2814	prep	of,are
R1569	T2818	T2819	amod	high,quality
R157	T467	T465	pobj	region,within
R1570	T2819	T2817	pobj	quality,of
R1571	T2820	T2812	punct	.,shows
R1572	T2822	T2823	advmod	Firstly,matched
R1573	T2824	T2823	punct	", ",matched
R1574	T2825	T2826	amod	directional,cloning
R1575	T2826	T2827	nsubj	cloning,was
R1576	T2827	T2823	ccomp	was,matched
R1577	T2828	T2827	acomp	successful,was
R1578	T2829	T2823	punct	: ,matched
R1579	T2830	T2831	advmod	only,"1,430"
R158	T468	T467	amod	coding,region
R1580	T2831	T2835	nummod	"1,430",sequences
R1581	T2832	T2831	quantmod	eight,"1,430"
R1582	T2833	T2831	quantmod	out,"1,430"
R1583	T2834	T2831	quantmod	of,"1,430"
R1584	T2835	T2823	nsubj	sequences,matched
R1585	T2836	T2835	compound	cDNA,sequences
R1586	T2837	T2835	prep	with,sequences
R1587	T2838	T2839	det	any,homology
R1588	T2839	T2837	pobj	homology,with
R1589	T2840	T2839	compound	protein,homology
R159	T469	T458	punct	", ",utilize
R1590	T2841	T2842	det	that,protein
R1591	T2842	T2823	dobj	protein,matched
R1592	T2843	T2823	prep	on,matched
R1593	T2844	T2845	det	the,strand
R1594	T2845	T2843	pobj	strand,on
R1595	T2846	T2845	amod	reverse,strand
R1596	T2847	T2823	punct	.,matched
R1597	T2849	T2850	advmod	Secondly,spliced
R1598	T2851	T2850	punct	", ",spliced
R1599	T2852	T2853	amod	genomic,contamination
R16	T313	T314	nsubj	We,screened
R160	T470	T458	advcl	contrary,utilize
R1600	T2853	T2854	nsubj	contamination,is
R1601	T2854	T2850	ccomp	is,spliced
R1602	T2855	T2854	acomp	rare,is
R1603	T2856	T2850	punct	: ,spliced
R1604	T2857	T2858	advmod	when,aligned
R1605	T2858	T2850	advcl	aligned,spliced
R1606	T2859	T2860	det	the,sequences
R1607	T2860	T2858	nsubjpass	sequences,aligned
R1608	T2861	T2860	nummod	83,sequences
R1609	T2862	T2863	amod	olfactory,receptor
R161	T471	T470	prep	to,contrary
R1610	T2863	T2864	npadvmod	receptor,containing
R1611	T2864	T2860	amod	containing,sequences
R1612	T2865	T2864	punct	-,containing
R1613	T2866	T2867	dep	that,had
R1614	T2867	T2860	relcl	had,sequences
R1615	T2868	T2869	det	a,UTR
R1616	T2869	T2867	dobj	UTR,had
R1617	T2870	T2869	nummod	5,UTR
R1618	T2871	T2870	punct	',5
R1619	T2872	T2869	prep	of,UTR
R162	T472	T473	det	the,structure
R1620	T2873	T2874	nummod	400,bp
R1621	T2874	T2872	pobj	bp,of
R1622	T2875	T2876	cc	or,longer
R1623	T2876	T2874	nummod	longer,bp
R1624	T2877	T2858	auxpass	were,aligned
R1625	T2878	T2858	prep	to,aligned
R1626	T2879	T2880	amod	genomic,sequence
R1627	T2880	T2878	pobj	sequence,to
R1628	T2881	T2850	punct	", ",spliced
R1629	T2882	T2850	nsubj	80,spliced
R163	T473	T471	pobj	structure,to
R1630	T2883	T2850	prep	across,spliced
R1631	T2884	T2885	advmod	at,one
R1632	T2885	T2887	nummod	one,intron
R1633	T2886	T2885	advmod	least,one
R1634	T2887	T2883	pobj	intron,across
R1635	T2888	T2850	punct	", ",spliced
R1636	T2889	T2850	advcl	leaving,spliced
R1637	T2890	T2891	det	a,maximum
R1638	T2891	T2889	dobj	maximum,leaving
R1639	T2892	T2891	prep	of,maximum
R164	T474	T473	amod	stereotyped,structure
R1640	T2893	T2894	nummod	three,clones
R1641	T2894	T2892	pobj	clones,of
R1642	T2895	T2896	punct	(,%
R1643	T2896	T2894	parataxis	%,clones
R1644	T2897	T2896	nummod	3.6,%
R1645	T2898	T2896	punct	),%
R1646	T2899	T2900	dep	that,represent
R1647	T2900	T2894	relcl	represent,clones
R1648	T2901	T2900	advmod	potentially,represent
R1649	T2902	T2903	amod	genomic,contamination
R165	T475	T476	amod	olfactory,receptor
R1650	T2903	T2900	dobj	contamination,represent
R1651	T2904	T2903	prep	of,contamination
R1652	T2905	T2906	det	the,libraries
R1653	T2906	T2904	pobj	libraries,of
R1654	T2907	T2850	punct	.,spliced
R1655	T2909	T2910	advmod	Thirdly,contain
R1656	T2911	T2910	punct	", ",contain
R1657	T2912	T2913	amod	most,clones
R1658	T2913	T2914	nsubj	clones,are
R1659	T2914	T2910	ccomp	are,contain
R166	T476	T477	compound	receptor,gene
R1660	T2915	T2914	prep	of,are
R1661	T2916	T2917	det	a,length
R1662	T2917	T2915	pobj	length,of
R1663	T2918	T2917	amod	reasonable,length
R1664	T2919	T2910	punct	: ,contain
R1665	T2920	T2921	mark	although,determine
R1666	T2921	T2910	advcl	determine,contain
R1667	T2922	T2921	nsubj	we,determine
R1668	T2923	T2921	aux	did,determine
R1669	T2924	T2921	neg	not,determine
R167	T477	T473	compound	gene,structure
R1670	T2925	T2926	mark	whether,are
R1671	T2926	T2921	ccomp	are,determine
R1672	T2927	T2926	nsubj	clones,are
R1673	T2928	T2929	amod	full,length
R1674	T2929	T2926	attr	length,are
R1675	T2930	T2929	punct	-,length
R1676	T2931	T2910	punct	", ",contain
R1677	T2932	T2933	quantmod	881,"1,264"
R1678	T2933	T2935	nummod	"1,264",cDNAs
R1679	T2934	T2933	quantmod	of,"1,264"
R168	T478	T458	punct	.,utilize
R1680	T2935	T2910	nsubj	cDNAs,contain
R1681	T2936	T2937	punct	(,%
R1682	T2937	T2933	parataxis	%,"1,264"
R1683	T2938	T2937	nummod	70,%
R1684	T2939	T2937	punct	),%
R1685	T2940	T2941	amod	olfactory,receptor
R1686	T2941	T2935	compound	receptor,cDNAs
R1687	T2942	T2943	det	the,gene
R1688	T2943	T2944	poss	gene,codon
R1689	T2944	T2910	dobj	codon,contain
R169	T480	T481	amod	Most,transcripts
R1690	T2945	T2943	case	's,gene
R1691	T2946	T2944	compound	start,codon
R1692	T2947	T2944	cc	and,codon
R1693	T2948	T2949	advmod	at,least
R1694	T2949	T2950	advmod	least,sequence
R1695	T2950	T2944	conj	sequence,codon
R1696	T2951	T2950	det	some,sequence
R1697	T2952	T2953	nummod	5,UTR
R1698	T2953	T2950	compound	UTR,sequence
R1699	T2954	T2952	punct	',5
R17	T315	T314	advmod	therefore,screened
R170	T481	T483	nsubj	transcripts,encode
R1700	T2955	T2910	punct	.,contain
R1701	T2957	T2958	prep	In,updated
R1702	T2959	T2957	pobj	order,In
R1703	T2960	T2961	aux	to,match
R1704	T2961	T2959	acl	match,order
R1705	T2962	T2961	dobj	cDNAs,match
R1706	T2963	T2961	prep	to,match
R1707	T2964	T2965	poss	their,counterparts
R1708	T2965	T2963	pobj	counterparts,to
R1709	T2966	T2965	amod	genomic,counterparts
R171	T482	T481	amod	atypical,transcripts
R1710	T2967	T2958	punct	", ",updated
R1711	T2968	T2958	advmod	first,updated
R1712	T2969	T2958	nsubj	we,updated
R1713	T2970	T2971	poss	our,catalog
R1714	T2971	T2958	dobj	catalog,updated
R1715	T2972	T2971	prep	of,catalog
R1716	T2973	T2974	nmod	mouse,genes
R1717	T2974	T2972	pobj	genes,of
R1718	T2975	T2976	amod	olfactory,receptor
R1719	T2976	T2974	compound	receptor,genes
R172	T484	T483	amod	nonfunctional,encode
R1720	T2977	T2978	punct	[,1
R1721	T2978	T2971	parataxis	1,catalog
R1722	T2979	T2978	punct	],1
R1723	T2980	T2958	prep	based,updated
R1724	T2981	T2980	prep	on,based
R1725	T2982	T2983	poss	Celera,assembly
R1726	T2983	T2981	pobj	assembly,on
R1727	T2984	T2982	case	's,Celera
R1728	T2985	T2986	advmod	most,recent
R1729	T2986	T2983	amod	recent,assembly
R173	T485	T483	amod	olfactory,encode
R1730	T2987	T2983	compound	genome,assembly
R1731	T2988	T2989	punct	(,Release
R1732	T2989	T2983	parataxis	Release,assembly
R1733	T2990	T2989	nummod	13,Release
R1734	T2991	T2989	punct	),Release
R1735	T2992	T2993	punct	[,29
R1736	T2993	T2983	parataxis	29,assembly
R1737	T2994	T2993	punct	],29
R1738	T2995	T2958	punct	.,updated
R1739	T2997	T2998	amod	Previous,reports
R174	T486	T483	dep	receptors,encode
R1740	T2998	T2999	nsubjpass	reports,based
R1741	T3000	T2998	prep	of,reports
R1742	T3001	T3002	det	the,repertoire
R1743	T3002	T3000	pobj	repertoire,of
R1744	T3003	T3004	nmod	mouse,receptor
R1745	T3004	T3002	compound	receptor,repertoire
R1746	T3005	T3004	amod	olfactory,receptor
R1747	T3006	T3007	punct	[,2
R1748	T3007	T2998	parataxis	2,reports
R1749	T3008	T3007	nummod	1,2
R175	T487	T483	punct	", ",encode
R1750	T3009	T3007	punct	",",2
R1751	T3010	T3007	punct	],2
R1752	T3011	T2999	auxpass	were,based
R1753	T3012	T2999	prep	on,based
R1754	T3013	T3014	det	the,assembly
R1755	T3014	T3012	pobj	assembly,on
R1756	T3015	T3014	nmod	Release,assembly
R1757	T3016	T3015	nummod	12,Release
R1758	T3017	T2999	punct	.,based
R1759	T3019	T3020	nsubj	Release,consists
R176	T488	T483	cc	but,encode
R1760	T3021	T3019	nummod	13,Release
R1761	T3022	T3020	prep	of,consists
R1762	T3023	T3024	amod	fewer,longer
R1763	T3024	T3026	amod	longer,sequences
R1764	T3025	T3024	punct	", ",longer
R1765	T3026	T3022	pobj	sequences,of
R1766	T3027	T3026	compound	scaffold,sequences
R1767	T3028	T3026	acl	containing,sequences
R1768	T3029	T3030	amod	fewer,smaller
R1769	T3030	T3032	amod	smaller,gaps
R177	T489	T490	aux	can,increase
R1770	T3031	T3030	punct	", ",smaller
R1771	T3032	T3028	dobj	gaps,containing
R1772	T3033	T3032	prep	than,gaps
R1773	T3034	T3033	pobj	Release,than
R1774	T3035	T3034	nummod	12,Release
R1775	T3036	T3020	punct	.,consists
R1776	T3038	T3039	advcl	Using,identified
R1777	T3040	T3041	det	the,methods
R1778	T3041	T3038	dobj	methods,Using
R1779	T3042	T3043	npadvmod	BLAST,based
R178	T490	T483	conj	increase,encode
R1780	T3043	T3041	amod	based,methods
R1781	T3044	T3043	punct	-,based
R1782	T3045	T3041	acl	detailed,methods
R1783	T3046	T3045	advmod	previously,detailed
R1784	T3047	T3048	punct	[,1
R1785	T3048	T3045	parataxis	1,detailed
R1786	T3049	T3048	punct	],1
R1787	T3050	T3039	punct	", ",identified
R1788	T3051	T3039	nsubj	we,identified
R1789	T3052	T3053	nummod	"1,490",sequences
R179	T491	T490	advmod	occasionally,increase
R1790	T3053	T3039	dobj	sequences,identified
R1791	T3054	T3055	amod	olfactory,receptor
R1792	T3055	T3053	compound	receptor,sequences
R1793	T3056	T3039	prep	in,identified
R1794	T3057	T3058	det	the,assembly
R1795	T3058	T3056	pobj	assembly,in
R1796	T3059	T3058	amod	new,assembly
R1797	T3060	T3039	punct	", ",identified
R1798	T3061	T3039	prep	including,identified
R1799	T3062	T3063	nummod	"1,107",genes
R18	T316	T317	det	a,library
R180	T492	T493	compound	receptor,diversity
R1800	T3063	T3061	pobj	genes,including
R1801	T3064	T3063	amod	intact,genes
R1802	T3065	T3066	amod	olfactory,receptor
R1803	T3066	T3063	compound	receptor,genes
R1804	T3067	T3039	punct	(,identified
R1805	T3068	T3039	prep	compared,identified
R1806	T3069	T3068	prep	to,compared
R1807	T3070	T3071	nummod	866,receptors
R1808	T3071	T3069	pobj	receptors,to
R1809	T3072	T3071	amod	intact,receptors
R181	T493	T490	dobj	diversity,increase
R1810	T3073	T3071	amod	olfactory,receptors
R1811	T3074	T3071	prep	in,receptors
R1812	T3075	T3076	det	the,assembly
R1813	T3076	T3074	pobj	assembly,in
R1814	T3077	T3076	amod	old,assembly
R1815	T3078	T3039	punct	),identified
R1816	T3079	T3039	advcl	reflecting,identified
R1817	T3080	T3081	det	the,rate
R1818	T3081	T3079	dobj	rate,reflecting
R1819	T3082	T3081	amod	reduced,rate
R182	T494	T483	punct	.,encode
R1820	T3083	T3084	compound	sequence,error
R1821	T3084	T3081	compound	error,rate
R1822	T3085	T3081	cc	and,rate
R1823	T3086	T3087	amod	increased,coverage
R1824	T3087	T3081	conj	coverage,rate
R1825	T3088	T3087	prep	of,coverage
R1826	T3089	T3090	det	the,assembly
R1827	T3090	T3088	pobj	assembly,of
R1828	T3091	T3090	amod	new,assembly
R1829	T3092	T3093	punct	(,Table
R183	T498	T499	poss	Our,collection
R1830	T3093	T3079	parataxis	Table,reflecting
R1831	T3094	T3093	nummod	1,Table
R1832	T3095	T3093	punct	),Table
R1833	T3096	T3039	punct	.,identified
R1834	T3098	T3099	nsubj	We,created
R1835	T3100	T3101	det	a,database
R1836	T3101	T3099	dobj	database,created
R1837	T3102	T3101	amod	local,database
R1838	T3103	T3101	prep	of,database
R1839	T3104	T3105	amod	genomic,sequences
R184	T499	T501	nsubj	collection,confirms
R1840	T3105	T3103	pobj	sequences,of
R1841	T3106	T3101	prep	including,database
R1842	T3107	T3108	det	all,loci
R1843	T3108	T3106	pobj	loci,including
R1844	T3109	T3110	amod	olfactory,receptor
R1845	T3110	T3108	compound	receptor,loci
R1846	T3111	T3108	cc	and,loci
R1847	T3112	T3113	nummod	0.5,Mb
R1848	T3113	T3114	compound	Mb,sequences
R1849	T3114	T3108	conj	sequences,loci
R185	T500	T499	compound	cDNA,collection
R1850	T3115	T3114	compound	flanking,sequences
R1851	T3116	T3117	punct	(,available
R1852	T3117	T3114	parataxis	available,sequences
R1853	T3118	T3117	mark	if,available
R1854	T3119	T3117	punct	),available
R1855	T3120	T3099	cc	and,created
R1856	T3121	T3099	conj	compared,created
R1857	T3122	T3123	det	each,sequence
R1858	T3123	T3121	dobj	sequence,compared
R1859	T3124	T3123	compound	cDNA,sequence
R186	T502	T503	amod	olfactory,function
R1860	T3125	T3121	prep	to,compared
R1861	T3126	T3127	det	this,database
R1862	T3127	T3125	pobj	database,to
R1863	T3128	T3127	punct	',database
R1864	T3129	T3130	amod	olfactory,subgenome
R1865	T3130	T3127	nmod	subgenome,database
R1866	T3131	T3127	punct	',database
R1867	T3132	T3121	advcl	using,compared
R1868	T3133	T3132	dobj	sim4,using
R1869	T3134	T3135	punct	[,30
R187	T503	T501	dobj	function,confirms
R1870	T3135	T3121	parataxis	30,compared
R1871	T3136	T3135	punct	],30
R1872	T3137	T3099	punct	.,created
R1873	T3139	T3140	nsubjpass	cDNAs,assigned
R1874	T3141	T3140	auxpass	were,assigned
R1875	T3142	T3140	prep	to,assigned
R1876	T3143	T3144	amod	individual,genes
R1877	T3144	T3142	pobj	genes,to
R1878	T3145	T3140	prep	based,assigned
R1879	T3146	T3145	prep	on,based
R188	T504	T503	prep	of,function
R1880	T3147	T3148	poss	their,match
R1881	T3148	T3146	pobj	match,on
R1882	T3149	T3148	amod	best,match
R1883	T3150	T3148	prep	to,match
R1884	T3151	T3152	det	an,region
R1885	T3152	T3150	pobj	region,to
R1886	T3153	T3154	amod	olfactory,receptor
R1887	T3154	T3152	compound	receptor,region
R1888	T3155	T3152	compound	coding,region
R1889	T3156	T3152	cc	or,region
R189	T505	T506	quantmod	over,one
R1890	T3157	T3158	poss	its,region
R1891	T3158	T3152	conj	region,region
R1892	T3159	T3158	amod	upstream,region
R1893	T3160	T3161	punct	(,see
R1894	T3161	T3140	parataxis	see,assigned
R1895	T3162	T3161	dobj	Materials,see
R1896	T3163	T3162	cc	and,Materials
R1897	T3164	T3162	conj	methods,Materials
R1898	T3165	T3161	punct	),see
R1899	T3166	T3140	punct	.,assigned
R19	T317	T314	dobj	library,screened
R190	T506	T504	pobj	one,of
R1900	T3168	T3169	prep	Of,matched
R1901	T3169	T3177	ccomp	matched,matched
R1902	T3170	T3171	det	the,cDNAs
R1903	T3171	T3168	pobj	cDNAs,Of
R1904	T3172	T3171	nummod	"1,264",cDNAs
R1905	T3173	T3174	amod	olfactory,receptor
R1906	T3174	T3171	compound	receptor,cDNAs
R1907	T3175	T3169	punct	", ",matched
R1908	T3176	T3169	nsubj	"1,176",matched
R1909	T3178	T3179	det	a,total
R191	T507	T506	punct	-,one
R1910	T3179	T3169	dobj	total,matched
R1911	T3180	T3179	prep	of,total
R1912	T3181	T3182	nummod	419,genes
R1913	T3182	T3180	pobj	genes,of
R1914	T3183	T3184	amod	olfactory,receptor
R1915	T3184	T3182	compound	receptor,genes
R1916	T3185	T3177	punct	;,matched
R1917	T3186	T3187	det	the,cDNAs
R1918	T3187	T3177	nsubj	cDNAs,matched
R1919	T3188	T3187	amod	remaining,cDNAs
R192	T508	T506	quantmod	third,one
R1920	T3189	T3177	preconj	either,matched
R1921	T3190	T3191	det	an,receptor
R1922	T3191	T3177	dobj	receptor,matched
R1923	T3192	T3191	amod	olfactory,receptor
R1924	T3193	T3177	prep	below,matched
R1925	T3194	T3195	poss	our,threshold
R1926	T3195	T3193	pobj	threshold,below
R1927	T3196	T3197	nummod	96,%
R1928	T3197	T3198	compound	%,identity
R1929	T3198	T3195	compound	identity,threshold
R193	T509	T506	prep	of,one
R1930	T3199	T3198	compound	nucleotide,identity
R1931	T3200	T3177	cc	or,matched
R1932	T3201	T3177	conj	had,matched
R1933	T3202	T3203	amod	ambiguous,matches
R1934	T3203	T3201	dobj	matches,had
R1935	T3204	T3203	acl	encompassing,matches
R1936	T3205	T3206	amod	more,one
R1937	T3206	T3208	nummod	one,region
R1938	T3207	T3206	quantmod	than,one
R1939	T3208	T3204	dobj	region,encompassing
R194	T510	T511	det	the,receptors
R1940	T3209	T3210	amod	olfactory,receptor
R1941	T3210	T3211	compound	receptor,gene
R1942	T3211	T3208	compound	gene,region
R1943	T3212	T3213	punct	(,see
R1944	T3213	T3201	parataxis	see,had
R1945	T3214	T3213	advmod	below,see
R1946	T3215	T3213	punct	),see
R1947	T3216	T3177	punct	.,matched
R195	T511	T509	pobj	receptors,of
R1950	T3520	T3521	det	A,primer
R1951	T3521	T3527	nsubj	primer,broadens
R1952	T3522	T3521	nmod	class,primer
R1953	T3523	T3522	nummod	I,class
R1954	T3524	T3525	amod	olfactory,receptor
R1955	T3525	T3521	nmod	receptor,primer
R1956	T3526	T3521	amod	degenerate,primer
R1957	T3528	T3529	amod	phylogenetic,distribution
R1958	T3529	T3527	dobj	distribution,broadens
R1959	T3530	T3529	prep	of,distribution
R196	T512	T511	amod	intact,receptors
R1960	T3531	T3532	amod	confirmed,genes
R1961	T3532	T3530	pobj	genes,of
R1962	T3533	T3534	amod	olfactory,receptor
R1963	T3534	T3532	compound	receptor,genes
R1964	T3536	T3537	amod	Previous,analyses
R1965	T3537	T3538	nsubj	analyses,define
R1966	T3539	T3537	prep	of,analyses
R1967	T3540	T3541	det	the,family
R1968	T3541	T3539	pobj	family,of
R1969	T3542	T3541	amod	mammalian,family
R197	T513	T511	nmod	mouse,receptors
R1970	T3543	T3544	amod	olfactory,receptor
R1971	T3544	T3541	compound	receptor,family
R1972	T3545	T3546	nummod	two,clades
R1973	T3546	T3538	dobj	clades,define
R1974	T3547	T3546	amod	major,clades
R1975	T3548	T3546	amod	phylogenetic,clades
R1976	T3549	T3546	punct	", ",clades
R1977	T3550	T3546	acl	referred,clades
R1978	T3551	T3550	prep	to,referred
R1979	T3552	T3550	prep	as,referred
R198	T514	T511	amod	olfactory,receptors
R1980	T3553	T3554	nmod	class,I
R1981	T3554	T3555	nummod	I,receptors
R1982	T3555	T3552	pobj	receptors,as
R1983	T3556	T3554	cc	and,I
R1984	T3557	T3554	conj	II,I
R1985	T3558	T3555	amod	olfactory,receptors
R1986	T3559	T3538	punct	", ",define
R1987	T3560	T3538	cc	and,define
R1988	T3561	T3538	conj	suggest,define
R1989	T3562	T3563	mark	that,are
R199	T515	T501	punct	.,confirms
R1990	T3563	T3561	ccomp	are,suggest
R1991	T3564	T3565	nmod	class,receptors
R1992	T3565	T3563	nsubj	receptors,are
R1993	T3566	T3564	nummod	I,class
R1994	T3567	T3565	amod	olfactory,receptors
R1995	T3568	T3569	advmod	more,similar
R1996	T3569	T3563	acomp	similar,are
R1997	T3570	T3569	prep	to,similar
R1998	T3571	T3572	nmod	fish,receptors
R1999	T3572	T3570	pobj	receptors,to
R20	T318	T319	nmod	mouse,epithelium
R200	T517	T518	nsubjpass	Most,annotated
R2000	T3573	T3572	amod	olfactory,receptors
R2001	T3574	T3575	mark	than,are
R2002	T3575	T3569	advcl	are,similar
R2003	T3576	T3577	compound	class,IIs
R2004	T3577	T3575	nsubj	IIs,are
R2005	T3578	T3579	punct	[,5
R2006	T3579	T3561	parataxis	5,suggest
R2007	T3580	T3579	punct	],5
R2008	T3581	T3538	punct	.,define
R2009	T3583	T3584	nsubj	Figure,illustrates
R201	T519	T517	prep	of,Most
R2010	T3585	T3583	nummod	1,Figure
R2011	T3586	T3587	det	the,diversity
R2012	T3587	T3584	dobj	diversity,illustrates
R2013	T3588	T3587	amod	phylogenetic,diversity
R2014	T3589	T3587	prep	of,diversity
R2015	T3590	T3591	poss	our,collection
R2016	T3591	T3589	pobj	collection,of
R2017	T3592	T3591	compound	cDNA,collection
R2018	T3593	T3584	punct	", ",illustrates
R2019	T3594	T3584	advcl	showing,illustrates
R202	T520	T521	det	these,genes
R2020	T3595	T3596	mark	that,confirmed
R2021	T3596	T3594	ccomp	confirmed,showing
R2022	T3597	T3596	nsubj	we,confirmed
R2023	T3598	T3596	aux	have,confirmed
R2024	T3599	T3596	dobj	expression,confirmed
R2025	T3600	T3599	prep	of,expression
R2026	T3601	T3602	advmod	at,one
R2027	T3602	T3604	nummod	one,gene
R2028	T3603	T3602	advmod	least,one
R2029	T3604	T3600	pobj	gene,of
R203	T521	T519	pobj	genes,of
R2030	T3605	T3606	amod	olfactory,receptor
R2031	T3606	T3604	compound	receptor,gene
R2032	T3607	T3599	prep	in,expression
R2033	T3608	T3609	det	each,clade
R2034	T3609	T3607	pobj	clade,in
R2035	T3610	T3609	amod	major,clade
R2036	T3611	T3609	prep	of,clade
R2037	T3612	T3613	det	the,genes
R2038	T3613	T3611	pobj	genes,of
R2039	T3614	T3613	nmod	class,genes
R204	T522	T518	auxpass	were,annotated
R2040	T3615	T3614	nummod	II,class
R2041	T3616	T3617	amod	olfactory,receptor
R2042	T3617	T3613	compound	receptor,genes
R2043	T3618	T3599	punct	", ",expression
R2044	T3619	T3599	cc	or,expression
R2045	T3620	T3621	quantmod	391,983
R2046	T3621	T3599	conj	983,expression
R2047	T3622	T3621	quantmod	out,983
R2048	T3623	T3621	quantmod	of,983
R2049	T3624	T3625	punct	(,%
R205	T523	T518	advmod	previously,annotated
R2050	T3625	T3621	parataxis	%,983
R2051	T3626	T3625	nummod	40,%
R2052	T3627	T3625	punct	),%
R2053	T3628	T3621	prep	of,983
R2054	T3629	T3630	det	all,genes
R2055	T3630	T3628	pobj	genes,of
R2056	T3631	T3630	amod	intact,genes
R2057	T3632	T3630	nmod	class,genes
R2058	T3633	T3632	nummod	II,class
R2059	T3634	T3635	amod	olfactory,receptor
R206	T524	T518	prep	as,annotated
R2060	T3635	T3630	compound	receptor,genes
R2061	T3636	T3637	advmod	where,are
R2062	T3637	T3630	relcl	are,genes
R2063	T3638	T3639	amod	full,length
R2064	T3639	T3641	nmod	length,data
R2065	T3640	T3639	punct	-,length
R2066	T3641	T3637	nsubj	data,are
R2067	T3642	T3643	amod	genomic,sequence
R2068	T3643	T3641	compound	sequence,data
R2069	T3644	T3637	acomp	available,are
R207	T525	T526	amod	olfactory,receptors
R2070	T3645	T3646	punct	(,branches
R2071	T3646	T3596	parataxis	branches,confirmed
R2072	T3647	T3646	amod	blue,branches
R2073	T3648	T3646	punct	),branches
R2074	T3649	T3584	punct	.,illustrates
R2075	T3651	T3652	det	The,screen
R2076	T3652	T3653	nsubj	screen,appears
R2077	T3654	T3653	advmod	thus,appears
R2078	T3655	T3656	advmod	relatively,unbiased
R2079	T3656	T3653	oprd	unbiased,appears
R208	T526	T524	pobj	receptors,as
R2080	T3657	T3653	prep	in,appears
R2081	T3658	T3659	poss	its,coverage
R2082	T3659	T3657	pobj	coverage,in
R2083	T3660	T3659	prep	of,coverage
R2084	T3661	T3662	nmod	class,receptors
R2085	T3662	T3660	pobj	receptors,of
R2086	T3663	T3661	nummod	II,class
R2087	T3664	T3662	amod	olfactory,receptors
R2088	T3665	T3653	punct	.,appears
R2089	T3667	T3668	advmod	However,provided
R209	T527	T518	prep	based,annotated
R2090	T3669	T3668	punct	", ",provided
R2091	T3670	T3671	poss	our,screen
R2092	T3671	T3668	nsubj	screen,provided
R2093	T3672	T3671	amod	random,screen
R2094	T3673	T3668	dobj	cDNAs,provided
R2095	T3674	T3668	prep	for,provided
R2096	T3675	T3676	advmod	only,124
R2097	T3676	T3680	nummod	124,receptors
R2098	T3677	T3676	quantmod	two,124
R2099	T3678	T3676	quantmod	out,124
R21	T319	T317	compound	epithelium,library
R210	T528	T529	advmod	solely,on
R2100	T3679	T3676	quantmod	of,124
R2101	T3680	T3674	pobj	receptors,for
R2102	T3681	T3680	amod	intact,receptors
R2103	T3682	T3680	punct	", ",receptors
R2104	T3683	T3684	amod	full,length
R2105	T3684	T3680	nmod	length,receptors
R2106	T3685	T3684	punct	-,length
R2107	T3686	T3680	nmod	class,receptors
R2108	T3687	T3686	nummod	I,class
R2109	T3688	T3680	amod	olfactory,receptors
R211	T529	T527	prep	on,based
R2110	T3689	T3668	punct	.,provided
R2111	T3691	T3692	prep	In,used
R2112	T3693	T3694	det	an,attempt
R2113	T3694	T3691	pobj	attempt,In
R2114	T3695	T3696	aux	to,broaden
R2115	T3696	T3694	acl	broaden,attempt
R2116	T3697	T3698	det	the,coverage
R2117	T3698	T3696	dobj	coverage,broaden
R2118	T3699	T3698	amod	phylogenetic,coverage
R2119	T3700	T3698	prep	of,coverage
R212	T530	T531	compound	sequence,similarity
R2120	T3701	T3702	poss	our,screen
R2121	T3702	T3700	pobj	screen,of
R2122	T3703	T3702	compound	hybridization,screen
R2123	T3704	T3692	punct	", ",used
R2124	T3705	T3692	nsubj	we,used
R2125	T3706	T3707	amod	additional,probes
R2126	T3707	T3692	dobj	probes,used
R2127	T3708	T3707	amod	degenerate,probes
R2128	T3709	T3692	prep	on,used
R2129	T3710	T3711	det	the,library
R213	T531	T529	pobj	similarity,on
R2130	T3711	T3709	pobj	library,on
R2131	T3712	T3711	amod	adult,library
R2132	T3713	T3692	cc	and,used
R2133	T3714	T3692	conj	screened,used
R2134	T3715	T3716	det	an,library
R2135	T3716	T3714	dobj	library,screened
R2136	T3717	T3716	amod	embryonic,library
R2137	T3718	T3719	punct	(,Table
R2138	T3719	T3714	parataxis	Table,screened
R2139	T3720	T3719	nummod	2,Table
R214	T532	T518	punct	.,annotated
R2140	T3721	T3719	punct	),Table
R2141	T3722	T3692	punct	.,used
R2142	T3724	T3725	det	These,experiments
R2143	T3725	T3726	nsubj	experiments,increase
R2144	T3727	T3726	aux	did,increase
R2145	T3728	T3726	neg	not,increase
R2146	T3729	T3730	det	the,diversity
R2147	T3730	T3726	dobj	diversity,increase
R2148	T3731	T3730	prep	of,diversity
R2149	T3732	T3731	pobj	clones,of
R215	T534	T535	poss	Our,finding
R2150	T3733	T3732	acl	identified,clones
R2151	T3734	T3735	punct	(,shown
R2152	T3735	T3726	parataxis	shown,increase
R2153	T3736	T3735	neg	not,shown
R2154	T3737	T3735	punct	),shown
R2155	T3738	T3726	punct	.,increase
R2156	T3740	T3741	det	This,underrepresentation
R2157	T3741	T3745	nsubj	underrepresentation,be
R2158	T3742	T3741	amod	severe,underrepresentation
R2159	T3743	T3741	nmod	class,underrepresentation
R216	T535	T536	nsubj	finding,is
R2160	T3744	T3743	nummod	I,class
R2161	T3746	T3745	aux	could,be
R2162	T3747	T3745	prep	due,be
R2163	T3748	T3747	pcomp	to,due
R2164	T3749	T3750	amod	experimental,bias
R2165	T3750	T3747	pobj	bias,due
R2166	T3751	T3750	punct	-,bias
R2167	T3752	T3753	det	a,consequence
R2168	T3753	T3750	appos	consequence,bias
R2169	T3754	T3753	prep	of,consequence
R217	T537	T538	mark	that,have
R2170	T3755	T3754	pcomp	using,of
R2171	T3756	T3757	amod	degenerate,primers
R2172	T3757	T3755	dobj	primers,using
R2173	T3758	T3759	aux	to,create
R2174	T3759	T3755	advcl	create,using
R2175	T3760	T3761	poss	our,probe
R2176	T3761	T3759	dobj	probe,create
R2177	T3762	T3761	compound	hybridization,probe
R2178	T3763	T3745	punct	.,be
R2179	T3765	T3766	advmod	Alternatively,expressed
R218	T538	T535	acl	have,finding
R2180	T3767	T3766	punct	", ",expressed
R2181	T3768	T3769	nmod	class,genes
R2182	T3769	T3766	nsubjpass	genes,expressed
R2183	T3770	T3768	nummod	I,class
R2184	T3771	T3766	aux	might,expressed
R2185	T3772	T3766	auxpass	be,expressed
R2186	T3773	T3766	prep	at,expressed
R2187	T3774	T3775	advmod	extremely,low
R2188	T3775	T3776	amod	low,levels
R2189	T3776	T3773	pobj	levels,at
R219	T539	T540	amod	different,receptors
R2190	T3777	T3766	prep	in,expressed
R2191	T3778	T3779	det	the,epithelium
R2192	T3779	T3777	pobj	epithelium,in
R2193	T3780	T3779	amod	olfactory,epithelium
R2194	T3781	T3766	punct	.,expressed
R2195	T3783	T3784	prep	In,designed
R2196	T3785	T3783	pobj	order,In
R2197	T3786	T3787	aux	to,determine
R2198	T3787	T3785	acl	determine,order
R2199	T3788	T3789	mark	whether,expressed
R22	T320	T319	amod	olfactory,epithelium
R220	T540	T538	nsubj	receptors,have
R2200	T3789	T3787	ccomp	expressed,determine
R2201	T3790	T3791	nmod	class,receptors
R2202	T3791	T3789	nsubjpass	receptors,expressed
R2203	T3792	T3790	nummod	I,class
R2204	T3793	T3791	amod	olfactory,receptors
R2205	T3794	T3789	auxpass	are,expressed
R2206	T3795	T3789	prep	in,expressed
R2207	T3796	T3797	det	the,epithelium
R2208	T3797	T3795	pobj	epithelium,in
R2209	T3798	T3797	amod	olfactory,epithelium
R221	T541	T540	amod	olfactory,receptors
R2210	T3799	T3784	punct	", ",designed
R2211	T3800	T3784	nsubj	we,designed
R2212	T3801	T3802	det	a,primer
R2213	T3802	T3784	dobj	primer,designed
R2214	T3803	T3804	amod	reverse,strand
R2215	T3804	T3802	nmod	strand,primer
R2216	T3805	T3804	punct	-,strand
R2217	T3806	T3802	amod	degenerate,primer
R2218	T3807	T3808	aux	to,recognize
R2219	T3808	T3784	advcl	recognize,designed
R222	T542	T543	amod	different,levels
R2220	T3809	T3810	det	a,motif
R2221	T3810	T3808	dobj	motif,recognize
R2222	T3811	T3810	prep	in,motif
R2223	T3812	T3813	compound	transmembrane,domain
R2224	T3813	T3811	pobj	domain,in
R2225	T3814	T3813	nummod	7,domain
R2226	T3815	T3816	punct	(,NP
R2227	T3816	T3813	parataxis	NP,domain
R2228	T3817	T3816	dep	PP,NP
R2229	T3818	T3816	punct	{,NP
R223	T543	T538	dobj	levels,have
R2230	T3819	T3820	compound	V,T
R2231	T3820	T3816	dep	T,NP
R2232	T3821	T3820	punct	/,T
R2233	T3822	T3820	compound	M,T
R2234	T3823	T3820	punct	/,T
R2235	T3824	T3820	compound	A,T
R2236	T3825	T3820	punct	/,T
R2237	T3826	T3816	advmod	},NP
R2238	T3827	T3816	punct	{,NP
R2239	T3828	T3829	compound	F,M
R224	T544	T543	compound	expression,levels
R2240	T3829	T3816	dep	M,NP
R2241	T3830	T3829	punct	/,M
R2242	T3831	T3829	compound	L,M
R2243	T3832	T3829	punct	/,M
R2244	T3833	T3829	compound	I,M
R2245	T3834	T3829	punct	/,M
R2246	T3835	T3816	advmod	},NP
R2247	T3836	T3816	punct	),NP
R2248	T3837	T3810	acl	enriched,motif
R2249	T3838	T3837	prep	among,enriched
R225	T545	T536	acomp	intriguing,is
R2250	T3839	T3840	nmod	class,sequences
R2251	T3840	T3838	pobj	sequences,among
R2252	T3841	T3839	nummod	I,class
R2253	T3842	T3843	amod	olfactory,receptor
R2254	T3843	T3840	compound	receptor,sequences
R2255	T3844	T3784	punct	.,designed
R2256	T3846	T3847	nsubjpass	Most,shared
R2257	T3848	T3846	prep	of,Most
R2258	T3849	T3850	det	the,motif
R2259	T3850	T3848	pobj	motif,of
R226	T546	T536	prep	given,is
R2260	T3851	T3847	auxpass	is,shared
R2261	T3852	T3847	prep	among,shared
R2262	T3853	T3854	det	all,receptors
R2263	T3854	T3852	pobj	receptors,among
R2264	T3855	T3854	amod	olfactory,receptors
R2265	T3856	T3847	punct	", ",shared
R2266	T3857	T3847	cc	but,shared
R2267	T3858	T3859	det	the,residue
R2268	T3859	T3862	nsubjpass	residue,found
R2269	T3860	T3859	amod	first,residue
R227	T547	T548	det	the,regime
R2270	T3861	T3859	compound	proline,residue
R2271	T3862	T3847	conj	found,shared
R2272	T3863	T3859	punct	(,residue
R2273	T3864	T3859	prep	at,residue
R2274	T3865	T3866	det	the,primer
R2275	T3866	T3867	poss	primer,end
R2276	T3867	T3864	pobj	end,at
R2277	T3868	T3866	case	's,primer
R2278	T3869	T3867	nummod	3,end
R2279	T3870	T3869	punct	',3
R228	T548	T546	pobj	regime,given
R2280	T3871	T3862	punct	),found
R2281	T3872	T3862	auxpass	is,found
R2282	T3873	T3862	prep	in,found
R2283	T3874	T3875	quantmod	121,124
R2284	T3875	T3878	nummod	124,genes
R2285	T3876	T3875	quantmod	out,124
R2286	T3877	T3875	quantmod	of,124
R2287	T3878	T3873	pobj	genes,in
R2288	T3879	T3880	punct	(,%
R2289	T3880	T3875	parataxis	%,124
R229	T549	T550	nummod	one,neuron
R2290	T3881	T3880	nummod	98,%
R2291	T3882	T3880	punct	),%
R2292	T3883	T3878	amod	intact,genes
R2293	T3884	T3878	nmod	class,genes
R2294	T3885	T3884	nummod	I,class
R2295	T3886	T3862	prep	compared,found
R2296	T3887	T3886	prep	to,compared
R2297	T3888	T3889	advmod	only,983
R2298	T3889	T3893	nummod	983,genes
R2299	T3890	T3889	quantmod	37,983
R23	T321	T317	compound	cDNA,library
R230	T550	T548	nmod	neuron,regime
R2300	T3891	T3889	quantmod	out,983
R2301	T3892	T3889	quantmod	of,983
R2302	T3893	T3887	pobj	genes,to
R2303	T3894	T3895	punct	(,%
R2304	T3895	T3889	parataxis	%,983
R2305	T3896	T3895	nummod	4,%
R2306	T3897	T3895	punct	),%
R2307	T3898	T3893	amod	intact,genes
R2308	T3899	T3893	nmod	class,genes
R2309	T3900	T3899	nummod	II,class
R231	T551	T550	punct	-,neuron
R2310	T3901	T3862	punct	.,found
R2311	T3903	T3904	advmod	When,combined
R2312	T3904	T3905	advcl	combined,amplifies
R2313	T3905	T3921	ccomp	amplifies,represented
R2314	T3906	T3904	prep	with,combined
R2315	T3907	T3908	det	another,primer
R2316	T3908	T3906	pobj	primer,with
R2317	T3909	T3910	amod	olfactory,receptor
R2318	T3910	T3908	nmod	receptor,primer
R2319	T3911	T3908	amod	degenerate,primer
R232	T552	T550	punct	", ",neuron
R2320	T3912	T3908	punct	", ",primer
R2321	T3913	T3908	appos	P26,primer
R2322	T3914	T3915	punct	[,17
R2323	T3915	T3904	parataxis	17,combined
R2324	T3916	T3915	punct	],17
R2325	T3917	T3905	punct	", ",amplifies
R2326	T3918	T3919	det	this,primer
R2327	T3919	T3905	nsubj	primer,amplifies
R2328	T3920	T3905	advmod	preferentially,amplifies
R2329	T3922	T3923	nmod	class,receptors
R233	T553	T554	nummod	one,gene
R2330	T3923	T3905	dobj	receptors,amplifies
R2331	T3924	T3922	nummod	I,class
R2332	T3925	T3923	amod	olfactory,receptors
R2333	T3926	T3923	prep	from,receptors
R2334	T3927	T3928	nmod	mouse,DNA
R2335	T3928	T3926	pobj	DNA,from
R2336	T3929	T3928	amod	genomic,DNA
R2337	T3930	T3921	punct	: ,represented
R2338	T3931	T3921	prep	of,represented
R2339	T3932	T3933	nummod	33,products
R234	T554	T550	appos	gene,neuron
R2340	T3933	T3931	pobj	products,of
R2341	T3934	T3933	amod	sequenced,products
R2342	T3935	T3933	punct	", ",products
R2343	T3936	T3933	amod	cloned,products
R2344	T3937	T3933	compound	PCR,products
R2345	T3938	T3921	punct	", ",represented
R2346	T3939	T3921	nsubj	17,represented
R2347	T3940	T3941	nummod	seven,receptors
R2348	T3941	T3921	dobj	receptors,represented
R2349	T3942	T3941	amod	different,receptors
R235	T555	T554	punct	-,gene
R2350	T3943	T3941	nmod	class,receptors
R2351	T3944	T3943	nummod	I,class
R2352	T3945	T3941	amod	olfactory,receptors
R2353	T3946	T3921	punct	", ",represented
R2354	T3947	T3948	nsubj	six,represented
R2355	T3948	T3921	conj	represented,represented
R2356	T3949	T3950	nummod	three,receptors
R2357	T3950	T3948	dobj	receptors,represented
R2358	T3951	T3950	amod	different,receptors
R2359	T3952	T3950	nmod	class,receptors
R236	T556	T548	compound	expression,regime
R2360	T3953	T3952	nummod	II,class
R2361	T3954	T3950	amod	olfactory,receptors
R2362	T3955	T3948	punct	", ",represented
R2363	T3956	T3948	cc	and,represented
R2364	T3957	T3958	nsubj	ten,represented
R2365	T3958	T3948	conj	represented,represented
R2366	T3959	T3960	nummod	five,contaminants
R2367	T3960	T3958	dobj	contaminants,represented
R2368	T3961	T3960	amod	different,contaminants
R2369	T3962	T3963	amod	non-olfactory,receptor
R237	T557	T548	prep	of,regime
R2370	T3963	T3960	compound	receptor,contaminants
R2371	T3964	T3921	punct	.,represented
R2372	T3966	T3967	amod	Degenerate,PCR
R2373	T3967	T3968	nsubj	PCR,showed
R2374	T3969	T3967	punct	", ",PCR
R2375	T3970	T3967	conj	cloning,PCR
R2376	T3971	T3970	punct	", ",cloning
R2377	T3972	T3970	cc	and,cloning
R2378	T3973	T3970	conj	sequencing,cloning
R2379	T3974	T3973	prep	from,sequencing
R238	T558	T559	amod	olfactory,receptors
R2380	T3975	T3976	amod	reverse,transcribed
R2381	T3976	T3978	amod	transcribed,RNA
R2382	T3977	T3976	punct	-,transcribed
R2383	T3978	T3974	pobj	RNA,from
R2384	T3979	T3980	amod	olfactory,epithelium
R2385	T3980	T3978	compound	epithelium,RNA
R2386	T3981	T3982	mark	that,expressed
R2387	T3982	T3968	ccomp	expressed,showed
R2388	T3983	T3984	advmod	at,seven
R2389	T3984	T3986	nummod	seven,receptors
R239	T559	T557	pobj	receptors,of
R2390	T3985	T3984	advmod	least,seven
R2391	T3986	T3982	nsubjpass	receptors,expressed
R2392	T3987	T3986	nmod	class,receptors
R2393	T3988	T3987	nummod	I,class
R2394	T3989	T3986	amod	olfactory,receptors
R2395	T3990	T3982	auxpass	are,expressed
R2396	T3991	T3982	punct	", ",expressed
R2397	T3992	T3993	advmod	as,as
R2398	T3993	T3995	cc	as,gene
R2399	T3994	T3993	advmod	well,as
R24	T322	T323	aux	to,obtain
R240	T560	T536	punct	.,is
R2400	T3995	T3982	dep	gene,expressed
R2401	T3996	T3995	nummod	one,gene
R2402	T3997	T3995	amod	additional,gene
R2403	T3998	T3995	nmod	class,gene
R2404	T3999	T3998	nummod	II,class
R2405	T4000	T3995	punct	(,gene
R2406	T4001	T3995	acl	colored,gene
R2407	T4002	T4001	oprd	red,colored
R2408	T4003	T4001	prep	in,colored
R2409	T4004	T4003	pobj	Figure,in
R241	T562	T563	nsubj	We,provide
R2410	T4005	T4004	nummod	1,Figure
R2411	T4006	T3995	punct	),gene
R2412	T4007	T3968	punct	.,showed
R2413	T4009	T4010	advmod	However,obtained
R2414	T4011	T4010	punct	", ",obtained
R2415	T4012	T4013	det	no,products
R2416	T4013	T4010	nsubjpass	products,obtained
R2417	T4014	T4010	aux	could,obtained
R2418	T4015	T4010	auxpass	be,obtained
R2419	T4016	T4010	prep	from,obtained
R242	T564	T565	nummod	5,region
R2420	T4017	T4018	det	the,adult
R2421	T4018	T4016	pobj	adult,from
R2422	T4019	T4018	cc	or,adult
R2423	T4020	T4021	det	the,fetal
R2424	T4021	T4022	nmod	fetal,libraries
R2425	T4022	T4018	conj	libraries,adult
R2426	T4023	T4024	amod	olfactory,epithelium
R2427	T4024	T4022	compound	epithelium,libraries
R2428	T4025	T4022	compound	cDNA,libraries
R2429	T4026	T4010	advcl	using,obtained
R243	T565	T568	compound	region,sequences
R2430	T4027	T4028	det	the,primer
R2431	T4028	T4026	dobj	primer,using
R2432	T4029	T4028	nmod	class,primer
R2433	T4030	T4029	nummod	I,class
R2434	T4031	T4010	punct	", ",obtained
R2435	T4032	T4010	advcl	suggesting,obtained
R2436	T4033	T4034	mark	that,contain
R2437	T4034	T4032	ccomp	contain,suggesting
R2438	T4035	T4036	det	the,libraries
R2439	T4036	T4034	nsubj	libraries,contain
R244	T566	T564	punct	',5
R2440	T4037	T4038	advmod	very,low
R2441	T4038	T4039	amod	low,levels
R2442	T4039	T4034	dobj	levels,contain
R2443	T4040	T4039	prep	of,levels
R2444	T4041	T4042	nmod	class,receptors
R2445	T4042	T4040	pobj	receptors,of
R2446	T4043	T4041	nummod	I,class
R2447	T4044	T4042	amod	olfactory,receptors
R2448	T4045	T4010	punct	.,obtained
R2449	T4047	T4048	nsubj	We,confirmed
R245	T567	T565	amod	untranslated,region
R2450	T4049	T4048	advmod	also,confirmed
R2451	T4050	T4048	dobj	expression,confirmed
R2452	T4051	T4050	prep	of,expression
R2453	T4052	T4053	nummod	nine,receptors
R2454	T4053	T4051	pobj	receptors,of
R2455	T4054	T4053	amod	additional,receptors
R2456	T4055	T4053	amod	olfactory,receptors
R2457	T4056	T4053	punct	(,receptors
R2458	T4057	T4058	nummod	three,class
R2459	T4058	T4053	appos	class,receptors
R246	T568	T563	dobj	sequences,provide
R2460	T4059	T4058	nummod	I,class
R2461	T4060	T4058	cc	and,class
R2462	T4061	T4062	nummod	six,class
R2463	T4062	T4058	conj	class,class
R2464	T4063	T4062	nummod	II,class
R2465	T4064	T4053	punct	", ",receptors
R2466	T4065	T4053	acl	colored,receptors
R2467	T4066	T4065	oprd	red,colored
R2468	T4067	T4065	prep	in,colored
R2469	T4068	T4067	pobj	Figure,in
R247	T569	T568	cc	and,sequences
R2470	T4069	T4068	nummod	1,Figure
R2471	T4070	T4053	punct	),receptors
R2472	T4071	T4048	prep	from,confirmed
R2473	T4072	T4071	pobj	subclades,from
R2474	T4073	T4074	dep	that,represented
R2475	T4074	T4072	relcl	represented,subclades
R2476	T4075	T4074	auxpass	were,represented
R2477	T4076	T4074	advmod	poorly,represented
R2478	T4077	T4074	prep	in,represented
R2479	T4078	T4079	poss	our,screen
R248	T570	T571	compound	candidate,regions
R2480	T4079	T4077	pobj	screen,in
R2481	T4080	T4079	compound	cDNA,screen
R2482	T4081	T4048	advcl	using,confirmed
R2483	T4082	T4083	npadvmod	gene,specific
R2484	T4083	T4085	amod	specific,pairs
R2485	T4084	T4083	punct	-,specific
R2486	T4085	T4081	dobj	pairs,using
R2487	T4086	T4085	compound	primer,pairs
R2488	T4087	T4088	aux	to,amplify
R2489	T4088	T4081	advcl	amplify,using
R249	T571	T568	conj	regions,sequences
R2490	T4089	T4090	compound	cDNA,library
R2491	T4090	T4088	dobj	library,amplify
R2492	T4091	T4090	cc	or,library
R2493	T4092	T4093	amod	reverse,transcribed
R2494	T4093	T4095	amod	transcribed,templates
R2495	T4094	T4093	punct	-,transcribed
R2496	T4095	T4090	conj	templates,library
R2497	T4096	T4095	compound	RNA,templates
R2498	T4097	T4048	punct	.,confirmed
R2499	T4099	T4100	prep	For,determined
R25	T323	T314	advcl	obtain,screened
R250	T572	T571	compound	promoter,regions
R2500	T4101	T4099	pobj	two,For
R2501	T4102	T4101	prep	of,two
R2502	T4103	T4104	det	the,genes
R2503	T4104	T4102	pobj	genes,of
R2504	T4105	T4104	nmod	class,genes
R2505	T4106	T4105	nummod	I,class
R2506	T4107	T4108	nsubj	we,shown
R2507	T4108	T4104	advcl	shown,genes
R2508	T4109	T4108	aux	had,shown
R2509	T4110	T4111	aux	to,expressed
R251	T573	T568	prep	for,sequences
R2510	T4111	T4108	xcomp	expressed,shown
R2511	T4112	T4111	auxpass	be,expressed
R2512	T4113	T4100	punct	", ",determined
R2513	T4114	T4100	nsubj	we,determined
R2514	T4115	T4116	amod	relative,levels
R2515	T4116	T4100	dobj	levels,determined
R2516	T4117	T4116	compound	transcript,levels
R2517	T4118	T4100	advcl	using,determined
R2518	T4119	T4120	amod	quantitative,PCR
R2519	T4120	T4118	dobj	PCR,using
R252	T574	T575	amod	more,300
R2520	T4121	T4120	compound	RT,PCR
R2521	T4122	T4120	punct	-,PCR
R2522	T4123	T4124	punct	(,see
R2523	T4124	T4100	parataxis	see,determined
R2524	T4125	T4124	advmod	below,see
R2525	T4126	T4124	punct	),see
R2526	T4127	T4100	punct	.,determined
R2527	T4129	T4130	compound	Expression,levels
R2528	T4130	T4131	nsubj	levels,were
R2529	T4132	T4131	acomp	similar,were
R253	T575	T577	nummod	300,receptors
R2530	T4133	T4132	prep	to,similar
R2531	T4134	T4133	pobj	those,to
R2532	T4135	T4134	acl	observed,those
R2533	T4136	T4135	prep	for,observed
R2534	T4137	T4136	pobj	genes,for
R2535	T4138	T4139	dep	that,represented
R2536	T4139	T4137	relcl	represented,genes
R2537	T4140	T4139	auxpass	were,represented
R2538	T4141	T4139	prep	in,represented
R2539	T4142	T4143	poss	our,collection
R254	T576	T575	quantmod	than,300
R2540	T4143	T4141	pobj	collection,in
R2541	T4144	T4143	compound	cDNA,collection
R2542	T4145	T4131	punct	", ",were
R2543	T4146	T4131	advcl	suggesting,were
R2544	T4147	T4148	mark	that,represented
R2545	T4148	T4146	advcl	represented,suggesting
R2546	T4149	T4150	nmod	class,receptors
R2547	T4150	T4148	nsubjpass	receptors,represented
R2548	T4151	T4149	nummod	I,class
R2549	T4152	T4150	amod	olfactory,receptors
R255	T577	T573	pobj	receptors,for
R2550	T4153	T4148	auxpass	are,represented
R2551	T4154	T4148	neg	not,represented
R2552	T4155	T4148	advmod	under,represented
R2553	T4156	T4148	punct	-,represented
R2554	T4157	T4148	prep	in,represented
R2555	T4158	T4159	det	the,epithelium
R2556	T4159	T4157	pobj	epithelium,in
R2557	T4160	T4159	amod	olfactory,epithelium
R2558	T4161	T4148	punct	", ",represented
R2559	T4162	T4148	cc	and,represented
R256	T578	T577	amod	olfactory,receptors
R2560	T4163	T4164	mark	that,is
R2561	T4164	T4148	conj	is,represented
R2562	T4165	T4166	det	the,dearth
R2563	T4166	T4164	nsubj	dearth,is
R2564	T4167	T4166	prep	of,dearth
R2565	T4168	T4169	nmod	class,cDNAs
R2566	T4169	T4167	pobj	cDNAs,of
R2567	T4170	T4168	nummod	I,class
R2568	T4171	T4166	prep	in,dearth
R2569	T4172	T4173	poss	our,screen
R257	T579	T568	punct	", ",sequences
R2570	T4173	T4171	pobj	screen,in
R2571	T4174	T4164	acomp	likely,is
R2572	T4175	T4176	aux	to,be
R2573	T4176	T4174	xcomp	be,likely
R2574	T4177	T4176	prep	due,be
R2575	T4178	T4177	pcomp	to,due
R2576	T4179	T4177	pobj	bias,due
R2577	T4180	T4179	prep	in,bias
R2578	T4181	T4182	det	the,libraries
R2579	T4182	T4180	pobj	libraries,in
R258	T580	T581	amod	valuable,resources
R2580	T4183	T4182	cc	and,libraries
R2581	T4184	T4183	punct	/,and
R2582	T4185	T4183	cc	or,and
R2583	T4186	T4187	compound	hybridization,probes
R2584	T4187	T4182	conj	probes,libraries
R2585	T4188	T4131	punct	.,were
R2588	T4620	T4621	det	Some,genes
R2589	T4621	T4624	nsubjpass	genes,expressed
R259	T581	T568	appos	resources,sequences
R2590	T4622	T4623	amod	olfactory,receptor
R2591	T4623	T4621	compound	receptor,genes
R2592	T4625	T4624	auxpass	are,expressed
R2593	T4626	T4624	prep	at,expressed
R2594	T4627	T4628	amod	higher,levels
R2595	T4628	T4626	pobj	levels,at
R2596	T4629	T4628	prep	than,levels
R2597	T4630	T4629	pobj	others,than
R2598	T4632	T4633	poss	Our,screen
R2599	T4633	T4635	nsubj	screen,suggests
R26	T324	T325	amod	olfactory,receptor
R260	T582	T581	prep	for,resources
R2600	T4634	T4633	compound	cDNA,screen
R2601	T4636	T4637	mark	that,expressed
R2602	T4637	T4635	ccomp	expressed,suggests
R2603	T4638	T4639	det	some,genes
R2604	T4639	T4637	nsubjpass	genes,expressed
R2605	T4640	T4641	amod	olfactory,receptor
R2606	T4641	T4639	compound	receptor,genes
R2607	T4642	T4637	auxpass	are,expressed
R2608	T4643	T4637	prep	at,expressed
R2609	T4644	T4645	amod	higher,levels
R261	T583	T584	amod	computational,motif
R2610	T4645	T4643	pobj	levels,at
R2611	T4646	T4645	prep	than,levels
R2612	T4647	T4646	pobj	others,than
R2613	T4648	T4635	punct	.,suggests
R2614	T4650	T4651	mark	If,expressed
R2615	T4651	T4657	advcl	expressed,follow
R2616	T4652	T4653	det	all,genes
R2617	T4653	T4651	nsubjpass	genes,expressed
R2618	T4654	T4655	amod	olfactory,receptor
R2619	T4655	T4653	compound	receptor,genes
R262	T584	T586	compound	motif,searches
R2620	T4656	T4651	auxpass	were,expressed
R2621	T4658	T4651	prep	at,expressed
R2622	T4659	T4660	amod	equal,levels
R2623	T4660	T4658	pobj	levels,at
R2624	T4661	T4651	punct	", ",expressed
R2625	T4662	T4651	cc	and,expressed
R2626	T4663	T4664	poss	our,screen
R2627	T4664	T4665	nsubj	screen,were
R2628	T4665	T4651	conj	were,expressed
R2629	T4666	T4664	cc	and,screen
R263	T585	T584	amod	regulatory,motif
R2630	T4667	T4664	conj	library,screen
R2631	T4668	T4665	acomp	unbiased,were
R2632	T4669	T4665	prep	in,were
R2633	T4670	T4671	poss	their,coverage
R2634	T4671	T4669	pobj	coverage,in
R2635	T4672	T4671	prep	of,coverage
R2636	T4673	T4674	det	the,receptors
R2637	T4674	T4672	pobj	receptors,of
R2638	T4675	T4674	nmod	class,receptors
R2639	T4676	T4675	nummod	II,class
R264	T586	T582	pobj	searches,for
R2640	T4677	T4674	amod	olfactory,receptors
R2641	T4678	T4657	punct	", ",follow
R2642	T4679	T4680	det	the,number
R2643	T4680	T4657	nsubj	number,follow
R2644	T4681	T4680	prep	of,number
R2645	T4682	T4681	pobj	cDNAs,of
R2646	T4683	T4680	acl	detected,number
R2647	T4684	T4683	prep	per,detected
R2648	T4685	T4686	nmod	class,receptor
R2649	T4686	T4684	pobj	receptor,per
R265	T587	T582	cc	and,for
R2650	T4687	T4685	nummod	II,class
R2651	T4688	T4686	amod	olfactory,receptor
R2652	T4689	T4657	aux	should,follow
R2653	T4690	T4691	det	a,distribution
R2654	T4691	T4657	dobj	distribution,follow
R2655	T4692	T4691	compound	Poisson,distribution
R2656	T4693	T4691	punct	", ",distribution
R2657	T4694	T4691	acl	calculated,distribution
R2658	T4695	T4694	prep	based,calculated
R2659	T4696	T4695	prep	on,based
R266	T588	T582	conj	for,for
R2660	T4697	T4698	det	the,assumption
R2661	T4698	T4696	pobj	assumption,on
R2662	T4699	T4700	mark	that,have
R2663	T4700	T4698	advcl	have,assumption
R2664	T4701	T4702	det	all,receptors
R2665	T4702	T4700	nsubj	receptors,have
R2666	T4703	T4702	nummod	983,receptors
R2667	T4704	T4702	amod	intact,receptors
R2668	T4705	T4702	nmod	class,receptors
R2669	T4706	T4705	nummod	II,class
R267	T589	T588	pcomp	designing,for
R2670	T4707	T4702	amod	olfactory,receptors
R2671	T4708	T4709	det	an,chance
R2672	T4709	T4700	dobj	chance,have
R2673	T4710	T4709	amod	equal,chance
R2674	T4711	T4709	prep	of,chance
R2675	T4712	T4713	auxpass	being,represented
R2676	T4713	T4711	pcomp	represented,of
R2677	T4714	T4713	prep	in,represented
R2678	T4715	T4716	det	the,screen
R2679	T4716	T4714	pobj	screen,in
R268	T590	T591	amod	olfactory,receptor
R2680	T4717	T4700	punct	", ",have
R2681	T4718	T4700	cc	but,have
R2682	T4719	T4720	mark	that,found
R2683	T4720	T4700	conj	found,have
R2684	T4721	T4722	nmod	class,receptors
R2685	T4722	T4720	nsubjpass	receptors,found
R2686	T4723	T4721	nummod	I,class
R2687	T4724	T4722	amod	olfactory,receptors
R2688	T4725	T4722	cc	and,receptors
R2689	T4726	T4722	conj	pseudogenes,receptors
R269	T591	T592	compound	receptor,microarrays
R2690	T4727	T4720	aux	can,found
R2691	T4728	T4720	neg	not,found
R2692	T4729	T4720	auxpass	be,found
R2693	T4730	T4731	punct	(,Figure
R2694	T4731	T4720	parataxis	Figure,found
R2695	T4732	T4731	nummod	2,Figure
R2696	T4733	T4731	punct	),Figure
R2697	T4734	T4657	punct	.,follow
R2698	T4736	T4737	nsubj	We,calculate
R2699	T4738	T4739	det	a,probability
R27	T325	T326	npadvmod	receptor,expressed
R270	T592	T589	dobj	microarrays,designing
R2700	T4739	T4737	dobj	probability,calculate
R2701	T4740	T4739	amod	low,probability
R2702	T4741	T4742	punct	(,one
R2703	T4742	T4739	parataxis	one,probability
R2704	T4743	T4742	advmod	approximately,one
R2705	T4744	T4742	prep	in,one
R2706	T4745	T4744	pobj	28,in
R2707	T4746	T4742	punct	),one
R2708	T4747	T4748	mark	that,observe
R2709	T4748	T4739	acl	observe,probability
R271	T593	T592	cc	and,microarrays
R2710	T4749	T4748	nsubj	we,observe
R2711	T4750	T4748	aux	would,observe
R2712	T4751	T4752	det	any,gene
R2713	T4752	T4748	dobj	gene,observe
R2714	T4753	T4752	prep	with,gene
R2715	T4754	T4755	advmod	at,eight
R2716	T4755	T4757	nummod	eight,cDNAs
R2717	T4756	T4755	advmod	least,eight
R2718	T4757	T4753	pobj	cDNAs,with
R2719	T4758	T4757	amod	matching,cDNAs
R272	T594	T595	amod	other,probes
R2720	T4759	T4748	prep	in,observe
R2721	T4760	T4761	det	the,set
R2722	T4761	T4759	pobj	set,in
R2723	T4762	T4761	prep	of,set
R2724	T4763	T4764	nummod	"1,176",cDNAs
R2725	T4764	T4762	pobj	cDNAs,of
R2726	T4765	T4766	nsubj	we,assigned
R2727	T4766	T4764	advcl	assigned,cDNAs
R2728	T4767	T4766	prep	to,assigned
R2729	T4768	T4769	amod	single,sequences
R273	T595	T592	conj	probes,microarrays
R2730	T4769	T4767	pobj	sequences,to
R2731	T4770	T4771	amod	olfactory,receptor
R2732	T4771	T4769	compound	receptor,sequences
R2733	T4772	T4737	punct	.,calculate
R2734	T4774	T4775	advmod	However,found
R2735	T4776	T4775	punct	", ",found
R2736	T4777	T4775	prep	for,found
R2737	T4778	T4779	nummod	17,receptors
R2738	T4779	T4777	pobj	receptors,for
R2739	T4780	T4779	amod	olfactory,receptors
R274	T596	T595	amod	experimental,probes
R2740	T4781	T4775	punct	", ",found
R2741	T4782	T4775	nsubj	we,found
R2742	T4783	T4784	nummod	ten,cDNAs
R2743	T4784	T4775	dobj	cDNAs,found
R2744	T4785	T4783	cc	or,ten
R2745	T4786	T4783	conj	more,ten
R2746	T4787	T4784	compound	matching,cDNAs
R2747	T4788	T4775	punct	", ",found
R2748	T4789	T4775	advcl	suggesting,found
R2749	T4790	T4791	mark	that,expressed
R275	T597	T563	punct	.,provide
R2750	T4791	T4789	ccomp	expressed,suggesting
R2751	T4792	T4791	nsubjpass	they,expressed
R2752	T4793	T4791	aux	might,expressed
R2753	T4794	T4791	auxpass	be,expressed
R2754	T4795	T4791	prep	at,expressed
R2755	T4796	T4797	amod	higher,levels
R2756	T4797	T4795	pobj	levels,at
R2757	T4798	T4797	prep	than,levels
R2758	T4799	T4800	amod	other,genes
R2759	T4800	T4798	pobj	genes,than
R276	T246	T247	compound	Odorant,receptor
R2760	T4801	T4802	amod	olfactory,receptor
R2761	T4802	T4800	compound	receptor,genes
R2762	T4803	T4804	punct	(,Figure
R2763	T4804	T4789	parataxis	Figure,suggesting
R2764	T4805	T4804	nummod	2,Figure
R2765	T4806	T4804	punct	),Figure
R2766	T4807	T4775	punct	.,found
R2767	T4809	T4810	det	The,genes
R2768	T4810	T4812	nsubj	genes,are
R2769	T4811	T4810	nummod	two,genes
R277	T247	T249	npadvmod	receptor,expressed
R2770	T4813	T4814	prep	for,found
R2771	T4814	T4810	relcl	found,genes
R2772	T4815	T4813	pobj	which,for
R2773	T4816	T4814	nsubj	we,found
R2774	T4817	T4818	amod	most,cDNAs
R2775	T4818	T4814	dobj	cDNAs,found
R2776	T4819	T4810	punct	(,genes
R2777	T4820	T4821	compound	AY318726,MOR28
R2778	T4821	T4810	appos	MOR28,genes
R2779	T4822	T4821	punct	/,MOR28
R278	T249	T250	amod	expressed,tags
R2780	T4823	T4821	cc	and,MOR28
R2781	T4824	T4825	compound	AY318727,MOR10
R2782	T4825	T4821	conj	MOR10,MOR28
R2783	T4826	T4825	punct	/,MOR10
R2784	T4827	T4810	punct	),genes
R2785	T4828	T4829	advmod	genomically,adjacent
R2786	T4829	T4812	acomp	adjacent,are
R2787	T4830	T4829	cc	and,adjacent
R2788	T4831	T4829	conj	in,adjacent
R2789	T4832	T4833	det	the,cluster
R279	T250	T252	nsubj	tags,demonstrate
R2790	T4833	T4831	pobj	cluster,in
R2791	T4834	T4835	advmod	well,studied
R2792	T4835	T4833	amod	studied,cluster
R2793	T4836	T4835	punct	-,studied
R2794	T4837	T4838	amod	olfactory,receptor
R2795	T4838	T4833	compound	receptor,cluster
R2796	T4839	T4840	amod	next,to
R2797	T4840	T4833	prep	to,cluster
R2798	T4841	T4842	det	the,locus
R2799	T4842	T4840	pobj	locus,to
R28	T326	T327	amod	expressed,tags
R280	T251	T250	compound	sequence,tags
R2800	T4843	T4844	compound	T,cell
R2801	T4844	T4842	compound	cell,locus
R2802	T4845	T4844	punct	-,cell
R2803	T4846	T4842	compound	receptor,locus
R2804	T4847	T4848	compound	α,δ
R2805	T4848	T4842	compound	δ,locus
R2806	T4849	T4848	punct	/,δ
R2807	T4850	T4851	punct	[,31
R2808	T4851	T4812	parataxis	31,are
R2809	T4852	T4851	nummod	18,31
R281	T253	T254	amod	olfactory,expression
R2810	T4853	T4851	punct	",",31
R2811	T4854	T4851	punct	],31
R2812	T4855	T4812	punct	.,are
R2813	T4857	T4858	amod	Quantitative,PCR
R2814	T4858	T4861	nsubj	PCR,confirms
R2815	T4859	T4858	compound	RT,PCR
R2816	T4860	T4858	punct	-,PCR
R2817	T4862	T4858	prep	of,PCR
R2818	T4863	T4864	nummod	six,receptors
R2819	T4864	T4862	pobj	receptors,of
R282	T254	T252	dobj	expression,demonstrate
R2820	T4865	T4864	amod	olfactory,receptors
R2821	T4866	T4867	mark	that,vary
R2822	T4867	T4861	ccomp	vary,confirms
R2823	T4868	T4869	compound	expression,levels
R2824	T4869	T4867	nsubj	levels,vary
R2825	T4870	T4867	aux	do,vary
R2826	T4871	T4867	advmod	indeed,vary
R2827	T4872	T4867	advmod	considerably,vary
R2828	T4873	T4867	prep	between,vary
R2829	T4874	T4873	pobj	genes,between
R283	T255	T254	prep	of,expression
R2830	T4875	T4861	punct	.,confirms
R2831	T4877	T4878	nsubj	We,used
R2832	T4879	T4880	amod	quantitative,PCR
R2833	T4880	T4878	dobj	PCR,used
R2834	T4881	T4880	punct	(,PCR
R2835	T4882	T4883	amod	real,time
R2836	T4883	T4880	nmod	time,PCR
R2837	T4884	T4883	punct	-,time
R2838	T4885	T4880	punct	),PCR
R2839	T4886	T4887	aux	to,measure
R284	T256	T257	quantmod	over,400
R2840	T4887	T4878	advcl	measure,used
R2841	T4888	T4889	amod	olfactory,epithelium
R2842	T4889	T4890	compound	epithelium,levels
R2843	T4890	T4887	dobj	levels,measure
R2844	T4891	T4890	compound	transcript,levels
R2845	T4892	T4890	prep	of,levels
R2846	T4893	T4894	nummod	six,genes
R2847	T4894	T4892	pobj	genes,of
R2848	T4895	T4896	amod	olfactory,receptor
R2849	T4896	T4894	compound	receptor,genes
R285	T257	T258	nummod	400,genes
R2850	T4897	T4894	cc	and,genes
R2851	T4898	T4899	det	the,gene
R2852	T4899	T4894	conj	gene,genes
R2853	T4900	T4899	amod	ribosomal,gene
R2854	T4901	T4899	compound	S16,gene
R2855	T4902	T4887	prep	in,measure
R2856	T4903	T4904	nummod	three,mice
R2857	T4904	T4902	pobj	mice,in
R2858	T4905	T4904	prep	of,mice
R2859	T4906	T4907	det	the,strain
R286	T258	T255	pobj	genes,of
R2860	T4907	T4905	pobj	strain,of
R2861	T4908	T4907	amod	same,strain
R2862	T4909	T4907	amod	inbred,strain
R2863	T4910	T4911	punct	(,Figure
R2864	T4911	T4887	parataxis	Figure,measure
R2865	T4912	T4911	nummod	3,Figure
R2866	T4913	T4911	punct	),Figure
R2867	T4914	T4878	punct	.,used
R2868	T4916	T4917	det	These,genes
R2869	T4917	T4918	nsubj	genes,include
R287	T259	T254	punct	", ",expression
R2870	T4919	T4920	nummod	two,receptors
R2871	T4920	T4918	dobj	receptors,include
R2872	T4921	T4920	amod	olfactory,receptors
R2873	T4922	T4920	prep	with,receptors
R2874	T4923	T4924	amod	more,20
R2875	T4924	T4926	nummod	20,cDNAs
R2876	T4925	T4924	quantmod	than,20
R2877	T4926	T4922	pobj	cDNAs,with
R2878	T4927	T4926	amod	matching,cDNAs
R2879	T4928	T4920	punct	", ",receptors
R288	T260	T261	amod	extensive,splicing
R2880	T4929	T4920	conj	two,receptors
R2881	T4930	T4929	prep	with,two
R2882	T4931	T4932	nummod	one,cDNAs
R2883	T4932	T4930	pobj	cDNAs,with
R2884	T4933	T4931	cc	or,one
R2885	T4934	T4931	conj	two,one
R2886	T4935	T4932	amod	matching,cDNAs
R2887	T4936	T4929	cc	and,two
R2888	T4937	T4938	nummod	two,receptors
R2889	T4938	T4929	conj	receptors,two
R289	T261	T254	conj	splicing,expression
R2890	T4939	T4938	nmod	class,receptors
R2891	T4940	T4939	nummod	I,class
R2892	T4941	T4938	amod	olfactory,receptors
R2893	T4942	T4938	prep	with,receptors
R2894	T4943	T4944	det	no,cDNAs
R2895	T4944	T4942	pobj	cDNAs,with
R2896	T4945	T4944	amod	matching,cDNAs
R2897	T4946	T4918	punct	.,include
R2898	T4948	T4949	prep	In,measure
R2899	T4950	T4951	det	these,assays
R29	T327	T323	dobj	tags,obtain
R290	T262	T261	amod	alternate,splicing
R2900	T4951	T4948	pobj	assays,In
R2901	T4952	T4949	punct	", ",measure
R2902	T4953	T4949	nsubj	we,measure
R2903	T4954	T4955	compound	transcript,level
R2904	T4955	T4949	dobj	level,measure
R2905	T4956	T4955	prep	per,level
R2906	T4957	T4958	amod	genomic,copy
R2907	T4958	T4956	pobj	copy,per
R2908	T4959	T4958	prep	of,copy
R2909	T4960	T4961	det	the,gene
R291	T263	T261	cc	and,splicing
R2910	T4961	T4959	pobj	gene,of
R2911	T4962	T4949	prep	by,measure
R2912	T4963	T4962	pcomp	comparing,by
R2913	T4964	T4965	advmod	how,well
R2914	T4965	T4966	advmod	well,amplifies
R2915	T4966	T4963	ccomp	amplifies,comparing
R2916	T4967	T4968	det	a,pair
R2917	T4968	T4966	nsubj	pair,amplifies
R2918	T4969	T4970	npadvmod	gene,specific
R2919	T4970	T4968	amod	specific,pair
R292	T264	T265	amod	unequal,levels
R2920	T4971	T4970	punct	-,specific
R2921	T4972	T4968	compound	primer,pair
R2922	T4973	T4974	amod	reverse,transcribed
R2923	T4974	T4976	amod	transcribed,RNA
R2924	T4975	T4974	punct	-,transcribed
R2925	T4976	T4966	dobj	RNA,amplifies
R2926	T4977	T4963	punct	", ",comparing
R2927	T4978	T4963	advcl	relative,comparing
R2928	T4979	T4978	prep	to,relative
R2929	T4980	T4981	det	a,curve
R293	T265	T261	conj	levels,splicing
R2930	T4981	T4979	pobj	curve,to
R2931	T4982	T4981	amod	standard,curve
R2932	T4983	T4981	prep	of,curve
R2933	T4984	T4983	pobj	amplification,of
R2934	T4985	T4984	prep	of,amplification
R2935	T4986	T4987	nmod	mouse,DNA
R2936	T4987	T4985	pobj	DNA,of
R2937	T4988	T4987	amod	genomic,DNA
R2938	T4989	T4949	punct	.,measure
R2939	T4991	T4992	nsubj	We,find
R294	T266	T265	compound	expression,levels
R2940	T4993	T4994	mark	that,vary
R2941	T4994	T4992	ccomp	vary,find
R2942	T4995	T4996	compound	expression,levels
R2943	T4996	T4994	nsubj	levels,vary
R2944	T4997	T4994	aux	can,vary
R2945	T4998	T4994	prep	by,vary
R2946	T4999	T5000	advmod	almost,300-fold
R2947	T5000	T4998	pcomp	300-fold,by
R2948	T5001	T4994	prep	between,vary
R2949	T5002	T5001	pobj	genes,between
R295	T271	T272	det	The,family
R2950	T5003	T5004	punct	(,Figure
R2951	T5004	T5002	parataxis	Figure,genes
R2952	T5005	T5004	prep	for,Figure
R2953	T5006	T5005	pobj	example,for
R2954	T5007	T5004	punct	", ",Figure
R2955	T5008	T5009	compound	genes,A
R2956	T5009	T5004	dep	A,Figure
R2957	T5010	T5009	cc	and,A
R2958	T5011	T5009	conj	D,A
R2959	T5012	T5004	punct	", ",Figure
R296	T272	T276	nsubj	family,is
R2960	T5013	T5004	nummod	3,Figure
R2961	T5014	T5004	punct	),Figure
R2962	T5015	T4992	punct	.,find
R2963	T5017	T5018	advmod	However,are
R2964	T5019	T5018	punct	", ",are
R2965	T5020	T5021	compound	cDNA,numbers
R2966	T5021	T5018	nsubj	numbers,are
R2967	T5022	T5018	neg	not,are
R2968	T5023	T5024	det	a,indicator
R2969	T5024	T5018	attr	indicator,are
R297	T273	T274	amod	olfactory,receptor
R2970	T5025	T5024	amod	good,indicator
R2971	T5026	T5024	prep	of,indicator
R2972	T5027	T5028	compound	expression,level
R2973	T5028	T5026	pobj	level,of
R2974	T5029	T5018	punct	", ",are
R2975	T5030	T5031	det	a,discrepancy
R2976	T5031	T5018	npadvmod	discrepancy,are
R2977	T5032	T5033	dep	that,is
R2978	T5033	T5031	relcl	is,discrepancy
R2979	T5034	T5033	acomp	likely,is
R298	T274	T275	compound	receptor,gene
R2980	T5035	T5036	aux	to,be
R2981	T5036	T5034	xcomp	be,likely
R2982	T5037	T5036	prep	due,be
R2983	T5038	T5037	pcomp	to,due
R2984	T5039	T5037	pobj	bias,due
R2985	T5040	T5039	prep	in,bias
R2986	T5041	T5042	det	the,screen
R2987	T5042	T5040	pobj	screen,in
R2988	T5043	T5044	punct	(,used
R2989	T5044	T5042	parataxis	used,screen
R299	T275	T272	compound	gene,family
R2990	T5045	T5044	nsubj	we,used
R2991	T5046	T5047	amod	degenerate,primers
R2992	T5047	T5044	dobj	primers,used
R2993	T5048	T5049	aux	to,make
R2994	T5049	T5044	advcl	make,used
R2995	T5050	T5051	det	the,probes
R2996	T5051	T5049	dobj	probes,make
R2997	T5052	T5044	punct	", ",used
R2998	T5053	T5054	dep	which,favor
R2999	T5054	T5044	ccomp	favor,used
R30	T328	T327	compound	sequence,tags
R300	T277	T276	attr	one,is
R3000	T5055	T5054	aux	will,favor
R3001	T5056	T5057	det	some,genes
R3002	T5057	T5054	dobj	genes,favor
R3003	T5058	T5054	prep	over,favor
R3004	T5059	T5058	pobj	others,over
R3005	T5060	T5044	punct	),used
R3006	T5061	T5040	cc	and,in
R3007	T5062	T5040	conj	in,in
R3008	T5063	T5064	det	the,libraries
R3009	T5064	T5062	pobj	libraries,in
R301	T278	T277	prep	of,one
R3010	T5065	T5066	punct	(,favor
R3011	T5066	T5064	parataxis	favor,libraries
R3012	T5067	T5068	compound	oligo,dT
R3013	T5068	T5070	compound	dT,priming
R3014	T5069	T5068	punct	-,dT
R3015	T5070	T5066	nsubj	priming,favor
R3016	T5071	T5066	aux	will,favor
R3017	T5072	T5066	dobj	genes,favor
R3018	T5073	T5072	prep	with,genes
R3019	T5074	T5075	amod	shorter,UTRs
R302	T279	T280	det	the,largest
R3020	T5075	T5073	pobj	UTRs,with
R3021	T5076	T5075	nummod	3,UTRs
R3022	T5077	T5076	punct	',3
R3023	T5078	T5066	punct	),favor
R3024	T5079	T5018	punct	.,are
R3025	T5081	T5082	prep	For,observe
R3026	T5083	T5081	pobj	example,For
R3027	T5084	T5082	punct	", ",observe
R3028	T5085	T5082	nsubj	we,observe
R3029	T5086	T5087	amod	large,differences
R303	T280	T278	pobj	largest,of
R3030	T5087	T5082	dobj	differences,observe
R3031	T5088	T5087	compound	expression,differences
R3032	T5089	T5087	prep	in,differences
R3033	T5090	T5091	predet	all,mice
R3034	T5091	T5089	pobj	mice,in
R3035	T5092	T5091	nummod	three,mice
R3036	T5093	T5087	prep	between,differences
R3037	T5094	T5095	nummod	two,genes
R3038	T5095	T5093	pobj	genes,between
R3039	T5096	T5097	prep	for,found
R304	T281	T276	prep	in,is
R3040	T5097	T5095	relcl	found,genes
R3041	T5098	T5096	pobj	which,for
R3042	T5099	T5100	amod	similar,numbers
R3043	T5100	T5097	nsubjpass	numbers,found
R3044	T5101	T5100	prep	of,numbers
R3045	T5102	T5101	pobj	cDNAs,of
R3046	T5103	T5097	auxpass	were,found
R3047	T5104	T5095	punct	(,genes
R3048	T5105	T5106	compound	genes,A
R3049	T5106	T5095	appos	A,genes
R305	T282	T283	det	the,genome
R3050	T5107	T5106	cc	and,A
R3051	T5108	T5106	conj	B,A
R3052	T5109	T5110	punct	", ",Figure
R3053	T5110	T5095	parataxis	Figure,genes
R3054	T5111	T5110	nummod	3,Figure
R3055	T5112	T5110	punct	),Figure
R3056	T5113	T5087	punct	", ",differences
R3057	T5114	T5087	cc	and,differences
R3058	T5115	T5116	advmod	conversely,levels
R3059	T5116	T5087	conj	levels,differences
R306	T283	T281	pobj	genome,in
R3060	T5117	T5116	punct	", ",levels
R3061	T5118	T5116	amod	similar,levels
R3062	T5119	T5116	compound	expression,levels
R3063	T5120	T5116	prep	between,levels
R3064	T5121	T5122	nummod	two,genes
R3065	T5122	T5120	pobj	genes,between
R3066	T5123	T5122	prep	with,genes
R3067	T5124	T5125	det	a,difference
R3068	T5125	T5123	pobj	difference,with
R3069	T5126	T5125	amod	ten-fold,difference
R307	T284	T283	amod	mammalian,genome
R3070	T5127	T5125	prep	in,difference
R3071	T5128	T5127	pobj	number,in
R3072	T5129	T5128	prep	of,number
R3073	T5130	T5129	pobj	cDNAs,of
R3074	T5131	T5128	acl	found,number
R3075	T5132	T5122	punct	(,genes
R3076	T5133	T5134	compound	genes,B
R3077	T5134	T5122	appos	B,genes
R3078	T5135	T5134	cc	and,B
R3079	T5136	T5134	conj	C,B
R308	T285	T276	punct	.,is
R3080	T5137	T5138	punct	", ",Figure
R3081	T5138	T5122	parataxis	Figure,genes
R3082	T5139	T5138	nummod	3,Figure
R3083	T5140	T5138	punct	),Figure
R3084	T5141	T5082	punct	.,observe
R3085	T5143	T5144	compound	Expression,levels
R3086	T5144	T5145	nsubj	levels,are
R3087	T5145	T5146	ccomp	are,find
R3088	T5147	T5148	advmod	mostly,consistent
R3089	T5148	T5145	acomp	consistent,are
R309	T287	T288	amod	Previous,analyses
R3090	T5149	T5145	prep	between,are
R3091	T5150	T5151	amod	different,mice
R3092	T5151	T5149	pobj	mice,between
R3093	T5152	T5146	punct	: ,find
R3094	T5153	T5146	nsubj	we,find
R3095	T5154	T5155	amod	similar,differences
R3096	T5155	T5146	dobj	differences,find
R3097	T5156	T5157	compound	expression,level
R3098	T5157	T5155	compound	level,differences
R3099	T5158	T5157	punct	-,level
R31	T329	T314	punct	", ",screened
R310	T288	T290	nsubj	analyses,identified
R3100	T5159	T5155	prep	between,differences
R3101	T5160	T5161	amod	olfactory,receptor
R3102	T5161	T5162	compound	receptor,genes
R3103	T5162	T5159	pobj	genes,between
R3104	T5163	T5146	prep	in,find
R3105	T5164	T5165	det	all,mice
R3106	T5165	T5163	pobj	mice,in
R3107	T5166	T5165	nummod	three,mice
R3108	T5167	T5165	acl	examined,mice
R3109	T5168	T5169	punct	(,is
R311	T289	T288	amod	computational,analyses
R3110	T5169	T5146	parataxis	is,find
R3111	T5170	T5171	advmod	that,is
R3112	T5171	T5169	advmod	is,is
R3113	T5172	T5169	punct	", ",is
R3114	T5173	T5174	det	the,order
R3115	T5174	T5169	nsubj	order,is
R3116	T5175	T5174	compound	rank,order
R3117	T5176	T5174	prep	of,order
R3118	T5177	T5178	det	the,genes
R3119	T5178	T5176	pobj	genes,of
R312	T291	T290	aux	have,identified
R3120	T5179	T5178	nummod	six,genes
R3121	T5180	T5169	acomp	similar,is
R3122	T5181	T5169	prep	among,is
R3123	T5182	T5183	det	the,mice
R3124	T5183	T5181	pobj	mice,among
R3125	T5184	T5183	nummod	three,mice
R3126	T5185	T5169	punct	),is
R3127	T5186	T5146	punct	", ",find
R3128	T5187	T5188	mark	although,is
R3129	T5188	T5146	advcl	is,find
R313	T292	T293	advmod	approximately,"1,500"
R3130	T5189	T5188	expl	there,is
R3131	T5190	T5188	attr	variation,is
R3132	T5191	T5190	prep	in,variation
R3133	T5192	T5193	compound	expression,level
R3134	T5193	T5191	pobj	level,in
R3135	T5194	T5193	prep	of,level
R3136	T5195	T5196	det	some,genes
R3137	T5196	T5194	pobj	genes,of
R3138	T5197	T5188	prep	between,is
R3139	T5198	T5197	pobj	mice,between
R314	T293	T294	nummod	"1,500",receptors
R3140	T5199	T5200	punct	(,Figure
R3141	T5200	T5188	parataxis	Figure,is
R3142	T5201	T5200	prep	for,Figure
R3143	T5202	T5201	pobj	example,for
R3144	T5203	T5200	punct	", ",Figure
R3145	T5204	T5205	compound	gene,E
R3146	T5205	T5200	dep	E,Figure
R3147	T5206	T5200	punct	", ",Figure
R3148	T5207	T5200	nummod	3,Figure
R3149	T5208	T5200	punct	),Figure
R315	T294	T290	dobj	receptors,identified
R3150	T5209	T5146	punct	.,find
R3151	T5211	T5212	advmod	In,situ
R3152	T5212	T5213	amod	situ,hybridization
R3153	T5213	T5214	nsubj	hybridization,shows
R3154	T5215	T5216	punct	(,Figure
R3155	T5216	T5213	parataxis	Figure,hybridization
R3156	T5217	T5216	nummod	4,Figure
R3157	T5218	T5216	punct	),Figure
R3158	T5219	T5220	mark	that,account
R3159	T5220	T5214	ccomp	account,shows
R316	T295	T294	nmod	mouse,receptors
R3160	T5221	T5222	amod	increased,numbers
R3161	T5222	T5220	nsubj	numbers,account
R3162	T5223	T5222	prep	of,numbers
R3163	T5224	T5225	amod	expressing,cells
R3164	T5225	T5223	pobj	cells,of
R3165	T5226	T5220	prep	for,account
R3166	T5227	T5226	pobj	some,for
R3167	T5228	T5227	punct	", ",some
R3168	T5229	T5227	cc	but,some
R3169	T5230	T5229	neg	not,but
R317	T296	T294	amod	olfactory,receptors
R3170	T5231	T5227	conj	all,some
R3171	T5232	T5227	punct	", ",some
R3172	T5233	T5227	prep	of,some
R3173	T5234	T5235	det	the,difference
R3174	T5235	T5233	pobj	difference,of
R3175	T5236	T5235	prep	in,difference
R3176	T5237	T5238	compound	transcript,levels
R3177	T5238	T5236	pobj	levels,in
R3178	T5239	T5235	prep	between,difference
R3179	T5240	T5239	pobj	two,between
R318	T297	T290	punct	", ",identified
R3180	T5241	T5240	prep	of,two
R3181	T5242	T5243	det	the,genes
R3182	T5243	T5241	pobj	genes,of
R3183	T5244	T5243	acl	tested,genes
R3184	T5245	T5244	prep	by,tested
R3185	T5246	T5247	amod	real,time
R3186	T5247	T5249	compound	time,PCR
R3187	T5248	T5247	punct	-,time
R3188	T5249	T5245	pobj	PCR,by
R3189	T5250	T5240	punct	(,two
R319	T298	T290	cc	but,identified
R3190	T5251	T5252	compound	genes,A
R3191	T5252	T5240	appos	A,two
R3192	T5253	T5252	cc	and,A
R3193	T5254	T5252	conj	D,A
R3194	T5255	T5252	prep	in,A
R3195	T5256	T5255	pobj	Figure,in
R3196	T5257	T5256	nummod	3,Figure
R3197	T5258	T5220	punct	),account
R3198	T5259	T5214	punct	.,shows
R3199	T5261	T5262	nsubj	We,hybridized
R32	T330	T314	advcl	providing,screened
R320	T299	T300	amod	experimental,evidence
R3200	T5263	T5264	amod	alternate,sections
R3201	T5264	T5262	dobj	sections,hybridized
R3202	T5265	T5264	amod	coronal,sections
R3203	T5266	T5264	amod	serial,sections
R3204	T5267	T5264	acl	spanning,sections
R3205	T5268	T5269	det	an,epithelium
R3206	T5269	T5267	dobj	epithelium,spanning
R3207	T5270	T5269	amod	entire,epithelium
R3208	T5271	T5269	amod	olfactory,epithelium
R3209	T5272	T5269	prep	of,epithelium
R321	T300	T301	nsubj	evidence,is
R3210	T5273	T5274	det	a,mouse
R3211	T5274	T5272	pobj	mouse,of
R3212	T5275	T5274	amod	young,mouse
R3213	T5276	T5277	punct	(,P6
R3214	T5277	T5274	parataxis	P6,mouse
R3215	T5278	T5277	punct	),P6
R3216	T5279	T5262	prep	with,hybridized
R3217	T5280	T5279	pobj	probes,with
R3218	T5281	T5280	prep	for,probes
R3219	T5282	T5283	compound	gene,A
R3220	T5283	T5281	pobj	A,for
R3221	T5284	T5283	cc	and,A
R3222	T5285	T5286	compound	gene,D
R3223	T5286	T5283	conj	D,A
R3224	T5287	T5262	punct	.,hybridized
R3225	T5289	T5290	nmod	Southern,blot
R3226	T5290	T5291	nmod	blot,analyses
R3227	T5291	T5294	nsubj	analyses,show
R3228	T5292	T5290	cc	and,blot
R3229	T5293	T5290	conj	BLAST,blot
R3230	T5295	T5296	mark	that,are
R3231	T5296	T5294	ccomp	are,show
R3232	T5297	T5298	det	both,probes
R3233	T5298	T5296	nsubj	probes,are
R3234	T5299	T5296	acomp	likely,are
R3235	T5300	T5301	aux	to,hybridize
R3236	T5301	T5299	xcomp	hybridize,likely
R3237	T5302	T5301	prep	to,hybridize
R3238	T5303	T5304	poss	their,genes
R3239	T5304	T5302	pobj	genes,to
R3240	T5305	T5304	amod	intended,genes
R3241	T5306	T5304	compound	target,genes
R3242	T5307	T5304	cc	and,genes
R3243	T5308	T5309	det	no,others
R3244	T5309	T5304	conj	others,genes
R3245	T5310	T5311	punct	(,shown
R3246	T5311	T5296	parataxis	shown,are
R3247	T5312	T5311	neg	not,shown
R3248	T5313	T5311	punct	),shown
R3249	T5314	T5294	punct	.,show
R3250	T5316	T5317	compound	Gene,A
R3251	T5317	T5318	nsubjpass	A,expressed
R3252	T5319	T5318	auxpass	is,expressed
R3253	T5320	T5318	prep	in,expressed
R3254	T5321	T5320	pobj	zone,in
R3255	T5322	T5321	nummod	4,zone
R3256	T5323	T5321	prep	of,zone
R3257	T5324	T5325	det	the,epithelium
R3258	T5325	T5323	pobj	epithelium,of
R3259	T5326	T5318	prep	according,expressed
R326	T1203	T1204	det	The,interaction
R3260	T5327	T5326	prep	to,according
R3261	T5328	T5329	det	the,nomenclature
R3262	T5329	T5327	pobj	nomenclature,to
R3263	T5330	T5329	prep	of,nomenclature
R3264	T5331	T5330	pobj	Sullivan,of
R3265	T5332	T5333	advmod	et,al.
R3266	T5333	T5331	advmod	al.,Sullivan
R3267	T5334	T5335	punct	[,32
R3268	T5335	T5329	parataxis	32,nomenclature
R3269	T5336	T5335	punct	],32
R327	T1204	T1205	nsubj	interaction,is
R3270	T5337	T5338	punct	(,Figure
R3271	T5338	T5318	parataxis	Figure,expressed
R3272	T5339	T5338	nummod	4a,Figure
R3273	T5340	T5338	punct	),Figure
R3274	T5341	T5318	punct	.,expressed
R3275	T5343	T5344	det	The,pattern
R3276	T5344	T5346	nsubj	pattern,correspond
R3277	T5345	T5344	compound	expression,pattern
R3278	T5346	T5352	ccomp	correspond,found
R3279	T5347	T5344	prep	of,pattern
R328	T1206	T1204	prep	of,interaction
R3280	T5348	T5349	compound	gene,D
R3281	T5349	T5347	pobj	D,of
R3282	T5350	T5346	aux	does,correspond
R3283	T5351	T5346	neg	not,correspond
R3284	T5353	T5346	prep	to,correspond
R3285	T5354	T5353	pobj	any,to
R3286	T5355	T5354	prep	of,any
R3287	T5356	T5357	det	the,zones
R3288	T5357	T5355	pobj	zones,of
R3289	T5358	T5357	nummod	four,zones
R329	T1207	T1208	amod	olfactory,receptors
R3290	T5359	T5357	punct	',zones
R3291	T5360	T5357	amod	classical,zones
R3292	T5361	T5357	punct	',zones
R3293	T5362	T5357	amod	olfactory,zones
R3294	T5363	T5357	amod	epithelial,zones
R3295	T5364	T5365	punct	[,32
R3296	T5365	T5346	parataxis	32,correspond
R3297	T5366	T5365	nummod	14,32
R3298	T5367	T5365	punct	",",32
R3299	T5368	T5365	nummod	15,32
R33	T331	T330	dobj	evidence,providing
R330	T1208	T1206	pobj	receptors,of
R3300	T5369	T5365	punct	",",32
R3301	T5370	T5365	punct	],32
R3302	T5371	T5352	punct	: ,found
R3303	T5372	T5373	amod	positive,cells
R3304	T5373	T5352	nsubjpass	cells,found
R3305	T5374	T5352	auxpass	are,found
R3306	T5375	T5352	prep	in,found
R3307	T5376	T5375	pobj	regions,in
R3308	T5377	T5376	prep	of,regions
R3309	T5378	T5379	nmod	endoturbinates,II
R331	T1209	T1207	punct	(,olfactory
R3310	T5379	T5377	pobj	II,of
R3311	T5380	T5379	cc	and,II
R3312	T5381	T5379	conj	III,II
R3313	T5382	T5379	cc	and,II
R3314	T5383	T5379	conj	ectoturbinate,II
R3315	T5384	T5383	nummod	3,ectoturbinate
R3316	T5385	T5352	punct	", ",found
R3317	T5386	T5352	advcl	resembling,found
R3318	T5387	T5388	det	the,pattern
R3319	T5388	T5386	dobj	pattern,resembling
R332	T1210	T1207	cc	or,olfactory
R3320	T5389	T5388	compound	expression,pattern
R3321	T5390	T5388	acl	seen,pattern
R3322	T5391	T5390	advmod	previously,seen
R3323	T5392	T5390	prep	for,seen
R3324	T5393	T5394	det	the,subfamily
R3325	T5394	T5392	pobj	subfamily,for
R3326	T5395	T5394	compound	OR37,subfamily
R3327	T5396	T5394	cc	and,subfamily
R3328	T5397	T5398	nmod	ORZ6,receptors
R3329	T5398	T5394	conj	receptors,subfamily
R333	T1211	T1207	conj	odorant,olfactory
R3330	T5399	T5398	amod	olfactory,receptors
R3331	T5400	T5401	punct	[,34
R3332	T5401	T5352	parataxis	34,found
R3333	T5402	T5401	nummod	33,34
R3334	T5403	T5401	punct	",",34
R3335	T5404	T5401	punct	],34
R3336	T5405	T5406	punct	(,Figure
R3337	T5406	T5352	parataxis	Figure,found
R3338	T5407	T5406	nummod	4b,Figure
R3339	T5408	T5406	punct	),Figure
R334	T1212	T1208	punct	),receptors
R3340	T5409	T5352	punct	.,found
R3341	T5411	T5412	advcl	Counting,find
R3342	T5413	T5414	det	the,number
R3343	T5414	T5411	dobj	number,Counting
R3344	T5415	T5414	amod	total,number
R3345	T5416	T5414	prep	of,number
R3346	T5417	T5418	amod	positive,cells
R3347	T5418	T5416	pobj	cells,of
R3348	T5419	T5411	prep	in,Counting
R3349	T5420	T5421	amod	alternate,sections
R335	T1213	T1204	prep	with,interaction
R3350	T5421	T5419	pobj	sections,in
R3351	T5422	T5421	prep	across,sections
R3352	T5423	T5424	det	the,epithelium
R3353	T5424	T5422	pobj	epithelium,across
R3354	T5425	T5424	amod	entire,epithelium
R3355	T5426	T5412	punct	", ",find
R3356	T5427	T5412	nsubj	we,find
R3357	T5428	T5429	mark	that,expressed
R3358	T5429	T5412	ccomp	expressed,find
R3359	T5430	T5431	compound	gene,A
R336	T1214	T1215	poss	their,ligands
R3360	T5431	T5429	nsubjpass	A,expressed
R3361	T5432	T5429	auxpass	is,expressed
R3362	T5433	T5429	prep	in,expressed
R3363	T5434	T5435	nummod	"2,905",cells
R3364	T5435	T5433	pobj	cells,in
R3365	T5436	T5429	punct	", ",expressed
R3366	T5437	T5438	advmod	about,12
R3367	T5438	T5439	nummod	12,cells
R3368	T5439	T5429	npadvmod	cells,expressed
R3369	T5440	T5438	quantmod	times,12
R337	T1215	T1213	pobj	ligands,with
R3370	T5441	T5439	amod	more,cells
R3371	T5442	T5439	prep	than,cells
R3372	T5443	T5444	compound	gene,D
R3373	T5444	T5442	pobj	D,than
R3374	T5445	T5444	punct	", ",D
R3375	T5446	T5447	dep	which,expressed
R3376	T5447	T5444	relcl	expressed,D
R3377	T5448	T5447	auxpass	is,expressed
R3378	T5449	T5447	prep	in,expressed
R3379	T5450	T5451	det	a,total
R338	T1216	T1215	compound	odorant,ligands
R3380	T5451	T5449	pobj	total,in
R3381	T5452	T5451	prep	of,total
R3382	T5453	T5454	nummod	249,cells
R3383	T5454	T5452	pobj	cells,of
R3384	T5455	T5412	punct	.,find
R3385	T5457	T5458	det	This,difference
R3386	T5458	T5460	nsubj	difference,account
R3387	T5459	T5458	amod	12-fold,difference
R3388	T5461	T5458	prep	in,difference
R3389	T5462	T5461	pobj	numbers,in
R339	T1217	T1218	det	the,step
R3390	T5463	T5462	prep	of,numbers
R3391	T5464	T5465	amod	expressing,cells
R3392	T5465	T5463	pobj	cells,of
R3393	T5466	T5460	aux	does,account
R3394	T5467	T5460	neg	not,account
R3395	T5468	T5460	prep	for,account
R3396	T5469	T5470	det	the,difference
R3397	T5470	T5468	pobj	difference,for
R3398	T5471	T5472	advmod	almost,300-fold
R3399	T5472	T5470	nummod	300-fold,difference
R34	T332	T331	prep	of,evidence
R340	T1218	T1205	attr	step,is
R3400	T5473	T5470	prep	in,difference
R3401	T5474	T5475	compound	RNA,levels
R3402	T5475	T5473	pobj	levels,in
R3403	T5476	T5470	acl	observed,difference
R3404	T5477	T5476	prep	by,observed
R3405	T5478	T5479	amod	real,time
R3406	T5479	T5481	compound	time,PCR
R3407	T5480	T5479	punct	-,time
R3408	T5481	T5477	pobj	PCR,by
R3409	T5482	T5460	punct	", ",account
R341	T1219	T1218	amod	first,step
R3410	T5483	T5460	advcl	implying,account
R3411	T5484	T5485	mark	that,is
R3412	T5485	T5483	ccomp	is,implying
R3413	T5486	T5487	det	the,level
R3414	T5487	T5485	nsubj	level,is
R3415	T5488	T5487	compound	transcript,level
R3416	T5489	T5487	prep	per,level
R3417	T5490	T5491	amod	expressing,cell
R3418	T5491	T5489	pobj	cell,per
R3419	T5492	T5487	prep	for,level
R342	T1220	T1218	prep	in,step
R3420	T5493	T5494	compound	gene,A
R3421	T5494	T5492	pobj	A,for
R3422	T5495	T5496	quantmod	about,25
R3423	T5496	T5497	npadvmod	25,higher
R3424	T5497	T5485	acomp	higher,is
R3425	T5498	T5496	quantmod	times,25
R3426	T5499	T5497	prep	than,higher
R3427	T5500	T5501	compound	transcript,level
R3428	T5501	T5499	pobj	level,than
R3429	T5502	T5485	prep	in,is
R343	T1221	T1222	det	a,pathway
R3430	T5503	T5504	det	each,cell
R3431	T5504	T5502	pobj	cell,in
R3432	T5505	T5504	amod	expressing,cell
R3433	T5506	T5485	prep	for,is
R3434	T5507	T5508	compound	gene,D
R3435	T5508	T5506	pobj	D,for
R3436	T5509	T5460	punct	.,account
R3437	T5511	T5512	nsubj	We,note
R3438	T5513	T5514	mark	that,appear
R3439	T5514	T5512	ccomp	appear,note
R344	T1222	T1220	pobj	pathway,in
R3440	T5515	T5516	compound	hybridization,intensities
R3441	T5516	T5514	nsubj	intensities,appear
R3442	T5517	T5516	prep	per,intensities
R3443	T5518	T5519	amod	positive,neuron
R3444	T5519	T5517	pobj	neuron,per
R3445	T5520	T5514	oprd	stronger,appear
R3446	T5521	T5520	prep	for,stronger
R3447	T5522	T5523	compound	gene,A
R3448	T5523	T5521	pobj	A,for
R3449	T5524	T5520	prep	than,stronger
R345	T1223	T1224	compound	signal,transduction
R3450	T5525	T5526	compound	gene,D
R3451	T5526	T5524	pobj	D,than
R3452	T5527	T5514	prep	after,appear
R3453	T5528	T5529	amod	comparable,times
R3454	T5529	T5527	pobj	times,after
R3455	T5530	T5529	compound	exposure,times
R3456	T5531	T5514	punct	", ",appear
R3457	T5532	T5514	prep	in,appear
R3458	T5533	T5532	pobj	accordance,in
R3459	T5534	T5533	prep	with,accordance
R346	T1224	T1222	compound	transduction,pathway
R3460	T5535	T5536	det	the,idea
R3461	T5536	T5534	pobj	idea,with
R3462	T5537	T5538	mark	that,are
R3463	T5538	T5536	acl	are,idea
R3464	T5539	T5540	compound	transcript,levels
R3465	T5540	T5538	nsubj	levels,are
R3466	T5541	T5538	acomp	higher,are
R3467	T5542	T5538	prep	per,are
R3468	T5543	T5542	pobj	cell,per
R3469	T5544	T5512	punct	.,note
R347	T1225	T1226	dep	that,results
R3470	T5546	T5547	advmod	Thus,suggest
R3471	T5548	T5547	punct	", ",suggest
R3472	T5549	T5547	nsubj	we,suggest
R3473	T5550	T5551	mark	that,account
R3474	T5551	T5547	ccomp	account,suggest
R3475	T5552	T5551	nsubj	expression,account
R3476	T5553	T5552	prep	in,expression
R3477	T5554	T5555	amod	more,cells
R3478	T5555	T5553	pobj	cells,in
R3479	T5556	T5553	cc	and,in
R348	T1226	T1222	relcl	results,pathway
R3480	T5557	T5553	conj	in,in
R3481	T5558	T5559	amod	higher,levels
R3482	T5559	T5557	pobj	levels,in
R3483	T5560	T5559	prep	per,levels
R3484	T5561	T5560	pobj	cell,per
R3485	T5562	T5551	advmod	together,account
R3486	T5563	T5551	prep	for,account
R3487	T5564	T5565	det	the,levels
R3488	T5565	T5563	pobj	levels,for
R3489	T5566	T5567	advmod	almost,300-fold
R349	T1227	T1226	prep	in,results
R3490	T5567	T5568	amod	300-fold,higher
R3491	T5568	T5565	amod	higher,levels
R3492	T5569	T5570	amod	olfactory,RNA
R3493	T5570	T5565	compound	RNA,levels
R3494	T5571	T5570	amod	epithelial,RNA
R3495	T5572	T5565	prep	of,levels
R3496	T5573	T5574	compound	gene,A
R3497	T5574	T5572	pobj	A,of
R3498	T5575	T5565	amod	relative,levels
R3499	T5576	T5575	prep	to,relative
R35	T333	T334	amod	olfactory,function
R350	T1228	T1229	det	the,perception
R3500	T5577	T5578	compound	gene,D
R3501	T5578	T5576	pobj	D,to
R3502	T5579	T5580	punct	(,Figure
R3503	T5580	T5551	parataxis	Figure,account
R3504	T5581	T5580	nummod	3,Figure
R3505	T5582	T5580	punct	),Figure
R3506	T5583	T5547	punct	.,suggest
R3508	T5714	T5715	advmod	Most,genes
R3509	T5715	T5718	nsubj	genes,have
R351	T1229	T1227	pobj	perception,in
R3510	T5716	T5717	amod	olfactory,receptor
R3511	T5717	T5715	compound	receptor,genes
R3512	T5719	T5720	amod	several,isoforms
R3513	T5720	T5718	dobj	isoforms,have
R3514	T5721	T5720	amod	transcriptional,isoforms
R3515	T5723	T5724	poss	Our,collection
R3516	T5724	T5726	nsubj	collection,reveals
R3517	T5725	T5724	compound	cDNA,collection
R3518	T5727	T5728	mark	that,show
R3519	T5728	T5726	ccomp	show,reveals
R352	T1230	T1229	prep	of,perception
R3520	T5729	T5730	advmod	at,two
R3521	T5730	T5728	nsubj	two,show
R3522	T5731	T5730	advmod	least,two
R3523	T5732	T5730	quantmod	thirds,two
R3524	T5733	T5730	prep	of,two
R3525	T5734	T5735	det	the,receptors
R3526	T5735	T5733	pobj	receptors,of
R3527	T5736	T5735	amod	olfactory,receptors
R3528	T5737	T5735	acl	sampled,receptors
R3529	T5738	T5739	amod	alternative,splicing
R353	T1231	T1230	pobj	smell,of
R3530	T5739	T5728	dobj	splicing,show
R3531	T5740	T5739	prep	of,splicing
R3532	T5741	T5742	poss	their,exons
R3533	T5742	T5740	pobj	exons,of
R3534	T5743	T5742	nummod	5,exons
R3535	T5744	T5743	punct	',5
R3536	T5745	T5742	amod	untranslated,exons
R3537	T5746	T5726	punct	.,reveals
R3538	T5748	T5749	advcl	Using,find
R3539	T5750	T5751	det	a,script
R354	T1232	T1205	punct	.,is
R3540	T5751	T5748	dobj	script,Using
R3541	T5752	T5751	compound	custom,script
R3542	T5753	T5754	aux	to,process
R3543	T5754	T5748	advcl	process,Using
R3544	T5755	T5756	compound	sim4,alignments
R3545	T5756	T5754	dobj	alignments,process
R3546	T5757	T5756	prep	of,alignments
R3547	T5758	T5759	nmod	cDNA,sequences
R3548	T5759	T5757	pobj	sequences,of
R3549	T5760	T5758	cc	and,cDNA
R355	T1234	T1235	det	The,family
R3550	T5761	T5758	conj	genomic,cDNA
R3551	T5762	T5749	punct	", ",find
R3552	T5763	T5749	nsubj	we,find
R3553	T5764	T5765	quantmod	two,eight
R3554	T5765	T5767	nummod	eight,forms
R3555	T5766	T5765	quantmod	to,eight
R3556	T5767	T5749	dobj	forms,find
R3557	T5768	T5767	amod	different,forms
R3558	T5769	T5767	compound	splice,forms
R3559	T5770	T5749	prep	for,find
R356	T1235	T1239	nsubj	family,is
R3560	T5771	T5770	pobj	85,for
R3561	T5772	T5771	punct	(,85
R3562	T5773	T5774	nummod	45,%
R3563	T5774	T5771	appos	%,85
R3564	T5775	T5771	punct	),85
R3565	T5776	T5771	prep	of,85
R3566	T5777	T5778	det	the,genes
R3567	T5778	T5776	pobj	genes,of
R3568	T5779	T5778	nummod	191,genes
R3569	T5780	T5781	prep	for,had
R357	T1236	T1237	amod	olfactory,receptor
R3570	T5781	T5778	relcl	had,genes
R3571	T5782	T5780	pobj	which,for
R3572	T5783	T5781	nsubj	we,had
R3573	T5784	T5781	aux	have,had
R3574	T5785	T5786	det	some,opportunity
R3575	T5786	T5781	dobj	opportunity,had
R3576	T5787	T5788	aux	to,observe
R3577	T5788	T5786	acl	observe,opportunity
R3578	T5789	T5790	amod	alternate,splicing
R3579	T5790	T5788	dobj	splicing,observe
R358	T1237	T1238	compound	receptor,gene
R3580	T5791	T5792	punct	(,minimum
R3581	T5792	T5749	parataxis	minimum,find
R3582	T5793	T5794	advmod	that,is
R3583	T5794	T5792	advmod	is,minimum
R3584	T5795	T5792	punct	", ",minimum
R3585	T5796	T5792	det	a,minimum
R3586	T5797	T5792	prep	of,minimum
R3587	T5798	T5799	nummod	two,cDNAs
R3588	T5799	T5797	pobj	cDNAs,of
R3589	T5800	T5799	punct	", ",cDNAs
R359	T1238	T1235	compound	gene,family
R3590	T5801	T5802	advmod	at,least
R3591	T5802	T5803	advmod	least,one
R3592	T5803	T5804	dep	one,spliced
R3593	T5804	T5799	relcl	spliced,cDNAs
R3594	T5805	T5803	prep	of,one
R3595	T5806	T5805	pobj	which,of
R3596	T5807	T5804	auxpass	is,spliced
R3597	T5808	T5792	punct	),minimum
R3598	T5809	T5792	punct	", ",minimum
R3599	T5810	T5792	cc	and,minimum
R36	T334	T332	pobj	function,of
R360	T1240	T1239	attr	one,is
R3600	T5811	T5792	conj	55,minimum
R3601	T5812	T5813	punct	(,%
R3602	T5813	T5811	parataxis	%,55
R3603	T5814	T5813	nummod	67,%
R3604	T5815	T5813	punct	),%
R3605	T5816	T5811	prep	of,55
R3606	T5817	T5818	det	the,genes
R3607	T5818	T5816	pobj	genes,of
R3608	T5819	T5818	nummod	82,genes
R3609	T5820	T5821	prep	for,have
R361	T1241	T1240	prep	of,one
R3610	T5821	T5818	relcl	have,genes
R3611	T5822	T5820	pobj	which,for
R3612	T5823	T5821	nsubj	we,have
R3613	T5824	T5825	nummod	four,cDNAs
R3614	T5825	T5821	dobj	cDNAs,have
R3615	T5826	T5824	cc	or,four
R3616	T5827	T5824	conj	more,four
R3617	T5828	T5825	punct	(,cDNAs
R3618	T5829	T5825	cc	and,cDNAs
R3619	T5830	T5831	advmod	thus,chance
R362	T1242	T1243	det	the,largest
R3620	T5831	T5825	conj	chance,cDNAs
R3621	T5832	T5831	det	a,chance
R3622	T5833	T5831	amod	higher,chance
R3623	T5834	T5831	prep	of,chance
R3624	T5835	T5834	pcomp	observing,of
R3625	T5836	T5837	det	any,splicing
R3626	T5837	T5835	dobj	splicing,observing
R3627	T5838	T5837	amod	alternate,splicing
R3628	T5839	T5811	punct	),55
R3629	T5840	T5841	punct	(,Figure
R363	T1243	T1241	pobj	largest,of
R3630	T5841	T5749	parataxis	Figure,find
R3631	T5842	T5841	nummod	5,Figure
R3632	T5843	T5841	punct	),Figure
R3633	T5844	T5749	punct	.,find
R3634	T5846	T5847	det	These,events
R3635	T5847	T5850	nsubjpass	events,restricted
R3636	T5848	T5847	amod	alternative,events
R3637	T5849	T5847	compound	splice,events
R3638	T5851	T5850	auxpass	are,restricted
R3639	T5852	T5853	advmod	almost,all
R364	T1244	T1243	prep	in,largest
R3640	T5853	T5850	advmod	all,restricted
R3641	T5854	T5850	prep	to,restricted
R3642	T5855	T5856	det	the,UTR
R3643	T5856	T5854	pobj	UTR,to
R3644	T5857	T5856	nummod	5,UTR
R3645	T5858	T5857	punct	',5
R3646	T5859	T5850	cc	and,restricted
R3647	T5860	T5850	conj	include,restricted
R3648	T5861	T5862	compound	exon,skipping
R3649	T5862	T5860	dobj	skipping,include
R365	T1245	T1246	det	the,genome
R3650	T5863	T5862	cc	and,skipping
R3651	T5864	T5865	amod	alternate,splice
R3652	T5865	T5866	nmod	splice,use
R3653	T5866	T5862	conj	use,skipping
R3654	T5867	T5865	punct	-,splice
R3655	T5868	T5865	appos	donor,splice
R3656	T5869	T5868	cc	and,donor
R3657	T5870	T5871	punct	-,acceptor
R3658	T5871	T5868	conj	acceptor,donor
R3659	T5872	T5850	punct	.,restricted
R366	T1246	T1244	pobj	genome,in
R3660	T5874	T5875	advmod	At,least
R3661	T5875	T5876	advmod	least,half
R3662	T5876	T5877	nsubj	half,utilize
R3663	T5878	T5876	prep	of,half
R3664	T5879	T5880	det	the,receptors
R3665	T5880	T5878	pobj	receptors,of
R3666	T5881	T5880	amod	olfactory,receptors
R3667	T5882	T5880	acl	represented,receptors
R3668	T5883	T5882	prep	in,represented
R3669	T5884	T5885	poss	our,collection
R367	T1247	T1246	amod	mammalian,genome
R3670	T5885	T5883	pobj	collection,in
R3671	T5886	T5885	compound	cDNA,collection
R3672	T5887	T5888	amod	more,one
R3673	T5888	T5890	nummod	one,site
R3674	T5889	T5888	quantmod	than,one
R3675	T5890	T5877	dobj	site,utilize
R3676	T5891	T5890	compound	polyadenylation,site
R3677	T5892	T5877	punct	", ",utilize
R3678	T5893	T5877	advcl	resulting,utilize
R3679	T5894	T5893	prep	in,resulting
R368	T1248	T1239	punct	", ",is
R3680	T5895	T5896	amod	alternative,isoforms
R3681	T5896	T5894	pobj	isoforms,in
R3682	T5897	T5898	nummod	3,UTR
R3683	T5898	T5896	compound	UTR,isoforms
R3684	T5899	T5897	punct	',3
R3685	T5900	T5877	punct	.,utilize
R3686	T5902	T5903	nsubj	We,estimated
R3687	T5904	T5903	aux	have,estimated
R3688	T5905	T5903	advmod	crudely,estimated
R3689	T5906	T5907	nummod	3,UTR
R369	T1249	T1239	advcl	comprising,is
R3690	T5907	T5909	compound	UTR,size
R3691	T5908	T5907	punct	',UTR
R3692	T5909	T5903	dobj	size,estimated
R3693	T5910	T5903	prep	for,estimated
R3694	T5911	T5912	nummod	"1,169",clones
R3695	T5912	T5910	pobj	clones,for
R3696	T5913	T5912	compound	cDNA,clones
R3697	T5914	T5903	prep	by,estimated
R3698	T5915	T5914	pcomp	combining,by
R3699	T5916	T5917	amod	approximate,size
R37	T335	T331	prep	for,evidence
R370	T1250	T1251	quantmod	about,"1,500"
R3700	T5917	T5919	compound	size,information
R3701	T5918	T5917	compound	insert,size
R3702	T5919	T5915	dobj	information,combining
R3703	T5920	T5915	prep	with,combining
R3704	T5921	T5922	nummod	5,sequence
R3705	T5922	T5924	compound	sequence,data
R3706	T5923	T5921	punct	',5
R3707	T5924	T5920	pobj	data,with
R3708	T5925	T5903	punct	.,estimated
R3709	T5927	T5928	amod	More,one
R371	T1251	T1252	nummod	"1,500",members
R3710	T5928	T5930	nummod	one,isoform
R3711	T5929	T5928	quantmod	than,one
R3712	T5930	T5934	nsubjpass	isoform,predicted
R3713	T5931	T5932	nummod	3,UTR
R3714	T5932	T5930	compound	UTR,isoform
R3715	T5933	T5931	punct	',3
R3716	T5935	T5934	auxpass	is,predicted
R3717	T5936	T5934	prep	for,predicted
R3718	T5937	T5938	quantmod	43,77
R3719	T5938	T5941	nummod	77,genes
R372	T1252	T1249	dobj	members,comprising
R3720	T5939	T5938	quantmod	of,77
R3721	T5940	T5938	quantmod	the,77
R3722	T5941	T5936	pobj	genes,for
R3723	T5942	T5943	punct	(,%
R3724	T5943	T5938	parataxis	%,77
R3725	T5944	T5943	nummod	56,%
R3726	T5945	T5943	punct	),%
R3727	T5946	T5947	prep	for,are
R3728	T5947	T5941	relcl	are,genes
R3729	T5948	T5946	pobj	which,for
R373	T1253	T1249	prep	in,comprising
R3730	T5949	T5947	expl	there,are
R3731	T5950	T5951	advmod	at,four
R3732	T5951	T5953	nummod	four,cDNAs
R3733	T5952	T5951	advmod	least,four
R3734	T5953	T5947	attr	cDNAs,are
R3735	T5954	T5953	prep	with,cDNAs
R3736	T5955	T5956	nummod	3,UTR
R3737	T5956	T5958	compound	UTR,size
R3738	T5957	T5955	punct	',3
R3739	T5958	T5959	compound	size,information
R374	T1254	T1255	det	the,genome
R3740	T5959	T5954	pobj	information,with
R3741	T5960	T5934	punct	.,predicted
R3742	T5962	T5963	nsubj	We,confirmed
R3743	T5964	T5965	det	the,isoforms
R3744	T5965	T5963	dobj	isoforms,confirmed
R3745	T5966	T5967	amod	alternative,polyadenylation
R3746	T5967	T5965	compound	polyadenylation,isoforms
R3747	T5968	T5965	prep	of,isoforms
R3748	T5969	T5970	quantmod	four,five
R3749	T5970	T5973	nummod	five,genes
R375	T1255	T1253	pobj	genome,in
R3750	T5971	T5970	quantmod	out,five
R3751	T5972	T5970	quantmod	of,five
R3752	T5973	T5968	pobj	genes,of
R3753	T5974	T5973	amod	selected,genes
R3754	T5975	T5963	prep	by,confirmed
R3755	T5976	T5975	pcomp	sequencing,by
R3756	T5977	T5978	det	the,end
R3757	T5978	T5976	dobj	end,sequencing
R3758	T5979	T5978	nummod	3,end
R3759	T5980	T5979	punct	',3
R376	T1256	T1255	compound	mouse,genome
R3760	T5981	T5978	prep	of,end
R3761	T5982	T5983	nummod	14,clones
R3762	T5983	T5981	pobj	clones,of
R3763	T5984	T5983	compound	cDNA,clones
R3764	T5985	T5963	punct	.,confirmed
R3765	T5987	T5988	det	These,sequences
R3766	T5988	T5990	nsubj	sequences,revealed
R3767	T5989	T5988	nummod	14,sequences
R3768	T5991	T5990	advmod	also,revealed
R3769	T5992	T5993	nummod	one,cDNA
R377	T1257	T1258	punct	[,2
R3770	T5993	T5990	dobj	cDNA,revealed
R3771	T5994	T5995	advmod	where,added
R3772	T5995	T5993	relcl	added,cDNA
R3773	T5996	T5997	det	the,tail
R3774	T5997	T5995	nsubjpass	tail,added
R3775	T5998	T5997	nmod	poly,tail
R3776	T5999	T6000	punct	(,A
R3777	T6000	T5997	parataxis	A,tail
R3778	T6001	T6000	punct	),A
R3779	T6002	T5995	auxpass	was,added
R378	T1258	T1239	parataxis	2,is
R3780	T6003	T6004	nummod	27,bp
R3781	T6004	T6005	npadvmod	bp,before
R3782	T6005	T5995	prep	before,added
R3783	T6006	T6007	det	the,codon
R3784	T6007	T6005	pobj	codon,before
R3785	T6008	T6007	compound	stop,codon
R3786	T6009	T5993	punct	", ",cDNA
R3787	T6010	T5993	cc	and,cDNA
R3788	T6011	T5993	conj	another,cDNA
R3789	T6012	T6013	advmod	where,spliced
R379	T1259	T1258	nummod	1,2
R3790	T6013	T6011	relcl	spliced,another
R3791	T6014	T6015	det	an,intron
R3792	T6015	T6013	nsubjpass	intron,spliced
R3793	T6016	T6013	auxpass	was,spliced
R3794	T6017	T6013	prep	out,spliced
R3795	T6018	T6017	prep	of,out
R3796	T6019	T6020	det	the,UTR
R3797	T6020	T6018	pobj	UTR,of
R3798	T6021	T6020	nummod	3,UTR
R3799	T6022	T6021	punct	',3
R38	T336	T337	amod	many,receptors
R380	T1260	T1258	punct	",",2
R3800	T6023	T5990	punct	", ",revealed
R3801	T6024	T5990	advcl	contrary,revealed
R3802	T6025	T6024	prep	to,contrary
R3803	T6026	T6027	det	the,stereotype
R3804	T6027	T6025	pobj	stereotype,to
R3805	T6028	T6027	amod	conventional,stereotype
R3806	T6029	T6027	prep	of,stereotype
R3807	T6030	T6031	amod	olfactory,receptor
R3808	T6031	T6032	compound	receptor,gene
R3809	T6032	T6033	compound	gene,structure
R381	T1261	T1258	punct	],2
R3810	T6033	T6029	pobj	structure,of
R3811	T6034	T5990	punct	.,revealed
R382	T1262	T1239	punct	.,is
R3822	T6417	T6418	det	A,subset
R3823	T6418	T6419	nsubj	subset,shows
R3824	T6420	T6418	prep	of,subset
R3825	T6421	T6422	amod	olfactory,receptors
R3826	T6422	T6420	pobj	receptors,of
R3827	T6423	T6424	amod	unusual,splicing
R3828	T6424	T6419	dobj	splicing,shows
R3829	T6426	T6427	nsubj	We,identified
R383	T1264	T1265	amod	Olfactory,receptors
R3830	T6428	T6429	nummod	62,cDNAs
R3831	T6429	T6427	dobj	cDNAs,identified
R3832	T6430	T6431	punct	(,%
R3833	T6431	T6429	parataxis	%,cDNAs
R3834	T6432	T6431	nummod	5,%
R3835	T6433	T6431	prep	of,%
R3836	T6434	T6435	det	all,clones
R3837	T6435	T6433	pobj	clones,of
R3838	T6436	T6437	amod	olfactory,receptor
R3839	T6437	T6435	compound	receptor,clones
R384	T1265	T1266	nsubjpass	receptors,identified
R3840	T6438	T6431	punct	),%
R3841	T6439	T6427	prep	from,identified
R3842	T6440	T6441	nummod	38,receptors
R3843	T6441	T6439	pobj	receptors,from
R3844	T6442	T6441	amod	intact,receptors
R3845	T6443	T6441	amod	olfactory,receptors
R3846	T6444	T6441	cc	and,receptors
R3847	T6445	T6446	nummod	one,pseudogene
R3848	T6446	T6441	conj	pseudogene,receptors
R3849	T6447	T6448	amod	olfactory,receptor
R385	T1267	T1266	auxpass	were,identified
R3850	T6448	T6446	compound	receptor,pseudogene
R3851	T6449	T6450	advmod	where,used
R3852	T6450	T6446	relcl	used,pseudogene
R3853	T6451	T6452	det	a,site
R3854	T6452	T6450	nsubjpass	site,used
R3855	T6453	T6452	compound	splice,site
R3856	T6454	T6452	prep	within,site
R3857	T6455	T6456	det	the,region
R3858	T6456	T6454	pobj	region,within
R3859	T6457	T6458	npadvmod	protein,coding
R386	T1268	T1266	advmod	originally,identified
R3860	T6458	T6456	amod	coding,region
R3861	T6459	T6458	punct	-,coding
R3862	T6460	T6450	auxpass	is,used
R3863	T6461	T6427	punct	.,identified
R3864	T6463	T6464	prep	For,appears
R3865	T6465	T6466	nummod	two,genes
R3866	T6466	T6463	pobj	genes,For
R3867	T6467	T6468	punct	(,Figure
R3868	T6468	T6466	parataxis	Figure,genes
R3869	T6469	T6470	amod	top,cDNAs
R387	T1269	T1266	prep	in,identified
R3870	T6470	T6468	dep	cDNAs,Figure
R3871	T6471	T6470	nummod	two,cDNAs
R3872	T6472	T6468	punct	", ",Figure
R3873	T6473	T6468	nummod	6,Figure
R3874	T6474	T6468	punct	),Figure
R3875	T6475	T6464	punct	", ",appears
R3876	T6476	T6477	det	the,protein
R3877	T6477	T6464	nsubj	protein,appears
R3878	T6478	T6477	amod	predicted,protein
R3879	T6479	T6480	aux	to,be
R388	T1270	T1271	det	an,experiment
R3880	T6480	T6464	xcomp	be,appears
R3881	T6481	T6482	det	an,receptor
R3882	T6482	T6480	attr	receptor,be
R3883	T6483	T6482	amod	intact,receptor
R3884	T6484	T6482	amod	olfactory,receptor
R3885	T6485	T6482	prep	with,receptor
R3886	T6486	T6487	nummod	three,acids
R3887	T6487	T6485	pobj	acids,with
R3888	T6488	T6486	cc	or,three
R3889	T6489	T6486	conj	ten,three
R389	T1271	T1269	pobj	experiment,in
R3890	T6490	T6487	compound	amino,acids
R3891	T6491	T6487	punct	", ",acids
R3892	T6492	T6487	prep	including,acids
R3893	T6493	T6494	det	the,methionine
R3894	T6494	T6492	pobj	methionine,including
R3895	T6495	T6494	amod	initiating,methionine
R3896	T6496	T6487	punct	", ",acids
R3897	T6497	T6487	acl	contributed,acids
R3898	T6498	T6497	agent	by,contributed
R3899	T6499	T6500	det	an,exon
R39	T337	T335	pobj	receptors,for
R390	T1272	T1271	amod	elegant,experiment
R3900	T6500	T6498	pobj	exon,by
R3901	T6501	T6500	amod	upstream,exon
R3902	T6502	T6464	punct	.,appears
R3903	T6504	T6505	det	A,structure
R3904	T6505	T6508	nsubjpass	structure,described
R3905	T6506	T6505	amod	similar,structure
R3906	T6507	T6505	compound	gene,structure
R3907	T6509	T6508	auxpass	was,described
R3908	T6510	T6508	advmod	previously,described
R3909	T6511	T6508	prep	for,described
R391	T1273	T1271	prep	based,experiment
R3910	T6512	T6513	det	a,receptor
R3911	T6513	T6511	pobj	receptor,for
R3912	T6514	T6513	amod	human,receptor
R3913	T6515	T6513	amod	olfactory,receptor
R3914	T6516	T6517	punct	[,25
R3915	T6517	T6508	parataxis	25,described
R3916	T6518	T6517	punct	],25
R3917	T6519	T6508	punct	.,described
R3918	T6521	T6522	nsubj	One,has
R3919	T6523	T6521	prep	of,One
R392	T1274	T1273	prep	on,based
R3920	T6524	T6525	det	these,genes
R3921	T6525	T6523	pobj	genes,of
R3922	T6526	T6525	nummod	two,genes
R3923	T6527	T6525	compound	mouse,genes
R3924	T6528	T6529	det	no,codon
R3925	T6529	T6522	dobj	codon,has
R3926	T6530	T6529	compound	start,codon
R3927	T6531	T6529	prep	in,codon
R3928	T6532	T6533	poss	its,exon
R3929	T6533	T6531	pobj	exon,in
R393	T1275	T1276	det	the,hypothesis
R3930	T6534	T6535	advmod	otherwise,intact
R3931	T6535	T6533	amod	intact,exon
R3932	T6536	T6533	amod	main,exon
R3933	T6537	T6533	amod	coding,exon
R3934	T6538	T6522	punct	.,has
R3935	T6540	T6541	det	The,splicing
R3936	T6541	T6543	nsubj	splicing,rescues
R3937	T6542	T6541	amod	unusual,splicing
R3938	T6544	T6543	advmod	thus,rescues
R3939	T6545	T6546	dep	what,be
R394	T1276	T1274	pobj	hypothesis,on
R3940	T6546	T6543	ccomp	be,rescues
R3941	T6547	T6546	aux	would,be
R3942	T6548	T6546	advmod	otherwise,be
R3943	T6549	T6550	det	a,gene
R3944	T6550	T6546	attr	gene,be
R3945	T6551	T6550	amod	dysfunctional,gene
R3946	T6552	T6543	punct	.,rescues
R3947	T6554	T6555	prep	In,encode
R3948	T6556	T6557	amod	most,cases
R3949	T6557	T6554	pobj	cases,In
R395	T1277	T1278	mark	that,be
R3950	T6558	T6557	punct	(,cases
R3951	T6559	T6560	quantmod	60,62
R3952	T6560	T6563	nummod	62,cDNAs
R3953	T6561	T6560	quantmod	out,62
R3954	T6562	T6560	quantmod	of,62
R3955	T6563	T6557	appos	cDNAs,cases
R3956	T6564	T6557	punct	),cases
R3957	T6565	T6555	punct	", ",encode
R3958	T6566	T6567	det	the,transcript
R3959	T6567	T6569	nsubj	transcript,appears
R396	T1278	T1276	acl	be,hypothesis
R3960	T6568	T6567	amod	unusual,transcript
R3961	T6569	T6555	ccomp	appears,encode
R3962	T6570	T6571	aux	to,be
R3963	T6571	T6569	xcomp	be,appears
R3964	T6572	T6573	det	an,form
R3965	T6573	T6571	attr	form,be
R3966	T6574	T6573	amod	aberrant,form
R3967	T6575	T6573	compound	splice,form
R3968	T6576	T6555	punct	-,encode
R3969	T6577	T6578	det	the,transcript
R397	T1279	T1278	nsubj	they,be
R3970	T6578	T6555	nsubj	transcript,encode
R3971	T6579	T6555	aux	would,encode
R3972	T6580	T6555	advmod	probably,encode
R3973	T6581	T6555	neg	not,encode
R3974	T6582	T6583	det	a,protein
R3975	T6583	T6555	dobj	protein,encode
R3976	T6584	T6583	amod	functional,protein
R3977	T6585	T6586	mark	because,introduces
R3978	T6586	T6555	advcl	introduces,encode
R3979	T6587	T6588	det	the,splice
R398	T1280	T1278	aux	would,be
R3980	T6588	T6586	nsubj	splice,introduces
R3981	T6589	T6590	det	a,frameshift
R3982	T6590	T6586	dobj	frameshift,introduces
R3983	T6591	T6586	cc	or,introduces
R3984	T6592	T6586	conj	removes,introduces
R3985	T6593	T6594	amod	conserved,residues
R3986	T6594	T6592	dobj	residues,removes
R3987	T6595	T6594	amod	functional,residues
R3988	T6596	T6597	punct	(,Figure
R3989	T6597	T6555	parataxis	Figure,encode
R399	T1281	T1282	nummod	seven,transmembrane
R3990	T6598	T6597	nummod	6,Figure
R3991	T6599	T6597	punct	),Figure
R3992	T6600	T6555	punct	.,encode
R3993	T6602	T6603	prep	For,is
R3994	T6604	T6605	nummod	two,clones
R3995	T6605	T6602	pobj	clones,For
R3996	T6606	T6607	punct	(,Figure
R3997	T6607	T6605	parataxis	Figure,clones
R3998	T6608	T6609	amod	bottom,cDNAs
R3999	T6609	T6607	dep	cDNAs,Figure
R40	T338	T337	amod	additional,receptors
R400	T1282	T1284	compound	transmembrane,domain
R4000	T6610	T6609	nummod	two,cDNAs
R4001	T6611	T6607	punct	", ",Figure
R4002	T6612	T6607	nummod	6,Figure
R4003	T6613	T6607	punct	),Figure
R4004	T6614	T6603	punct	", ",is
R4005	T6615	T6616	compound	exon,order
R4006	T6616	T6603	nsubj	order,is
R4007	T6617	T6616	prep	in,order
R4008	T6618	T6619	det	the,clone
R4009	T6619	T6617	pobj	clone,in
R401	T1283	T1282	punct	-,transmembrane
R4010	T6620	T6619	compound	cDNA,clone
R4011	T6621	T6603	acomp	inconsistent,is
R4012	T6622	T6621	prep	with,inconsistent
R4013	T6623	T6624	det	the,sequence
R4014	T6624	T6622	pobj	sequence,with
R4015	T6625	T6624	amod	corresponding,sequence
R4016	T6626	T6624	amod	genomic,sequence
R4017	T6627	T6603	punct	.,is
R4018	T6629	T6630	nsubj	It,is
R4019	T6630	T6631	ccomp	is,suggest
R402	T1284	T1286	compound	domain,proteins
R4020	T6632	T6630	acomp	difficult,is
R4021	T6633	T6634	aux	to,imagine
R4022	T6634	T6630	xcomp	imagine,is
R4023	T6635	T6636	det	what,kind
R4024	T6636	T6637	dep	kind,resulted
R4025	T6637	T6634	ccomp	resulted,imagine
R4026	T6638	T6636	prep	of,kind
R4027	T6639	T6640	amod	cloning,artefact
R4028	T6640	T6638	pobj	artefact,of
R4029	T6641	T6637	prep	in,resulted
R403	T1285	T1284	punct	-,domain
R4030	T6642	T6643	det	these,cDNAs
R4031	T6643	T6641	pobj	cDNAs,in
R4032	T6644	T6645	advmod	severely,scrambled
R4033	T6645	T6643	amod	scrambled,cDNAs
R4034	T6646	T6631	punct	: ,suggest
R4035	T6647	T6631	nsubj	we,suggest
R4036	T6648	T6649	mark	that,derive
R4037	T6649	T6631	ccomp	derive,suggest
R4038	T6650	T6649	nsubj	they,derive
R4039	T6651	T6649	prep	from,derive
R404	T1286	T1278	attr	proteins,be
R4040	T6652	T6653	amod	real,transcripts
R4041	T6653	T6651	pobj	transcripts,from
R4042	T6654	T6652	cc	but,real
R4043	T6655	T6652	conj	rare,real
R4044	T6656	T6631	punct	.,suggest
R4045	T6658	T6659	advmod	However,suggests
R4046	T6660	T6659	punct	", ",suggests
R4047	T6661	T6662	poss	their,frequency
R4048	T6662	T6659	nsubj	frequency,suggests
R4049	T6663	T6662	amod	low,frequency
R405	T1287	T1286	acl	encoded,proteins
R4050	T6664	T6662	prep	in,frequency
R4051	T6665	T6666	poss	our,collection
R4052	T6666	T6664	pobj	collection,in
R4053	T6667	T6666	compound	cDNA,collection
R4054	T6668	T6669	mark	that,contribute
R4055	T6669	T6659	ccomp	contribute,suggests
R4056	T6670	T6669	nsubj	splicing,contribute
R4057	T6671	T6670	amod	contrary,splicing
R4058	T6672	T6671	prep	to,contrary
R4059	T6673	T6674	amod	genomic,organization
R406	T1288	T1287	agent	by,encoded
R4060	T6674	T6672	pobj	organization,to
R4061	T6675	T6669	aux	does,contribute
R4062	T6676	T6669	neg	not,contribute
R4063	T6677	T6669	advmod	significantly,contribute
R4064	T6678	T6669	prep	to,contribute
R4065	T6679	T6680	det	the,repertoire
R4066	T6680	T6678	pobj	repertoire,to
R4067	T6681	T6682	amod	olfactory,receptor
R4068	T6682	T6680	compound	receptor,repertoire
R4069	T6683	T6680	compound	transcript,repertoire
R407	T1289	T1290	det	a,family
R4070	T6684	T6659	punct	.,suggests
R4071	T6686	T6687	prep	For,observe
R4072	T6688	T6689	quantmod	21,26
R4073	T6689	T6692	nummod	26,genes
R4074	T6690	T6689	quantmod	of,26
R4075	T6691	T6689	quantmod	the,26
R4076	T6692	T6686	pobj	genes,For
R4077	T6693	T6694	prep	for,found
R4078	T6694	T6692	relcl	found,genes
R4079	T6695	T6693	pobj	which,for
R408	T1290	T1288	pobj	family,by
R4080	T6696	T6697	advmod	unusually,spliced
R4081	T6697	T6698	amod	spliced,cDNAs
R4082	T6698	T6694	nsubjpass	cDNAs,found
R4083	T6699	T6694	auxpass	were,found
R4084	T6700	T6687	punct	", ",observe
R4085	T6701	T6687	nsubj	we,observe
R4086	T6702	T6687	advmod	also,observe
R4087	T6703	T6704	det	an,isoform
R4088	T6704	T6687	dobj	isoform,observe
R4089	T6705	T6704	amod	alternative,isoform
R409	T1291	T1290	amod	large,family
R4090	T6706	T6704	punct	(,isoform
R4091	T6707	T6704	punct	',isoform
R4092	T6708	T6704	amod	normal,isoform
R4093	T6709	T6704	punct	',isoform
R4094	T6710	T6704	punct	),isoform
R4095	T6711	T6712	dep	that,use
R4096	T6712	T6704	relcl	use,isoform
R4097	T6713	T6712	aux	does,use
R4098	T6714	T6712	neg	not,use
R4099	T6715	T6716	compound	splice,sites
R41	T339	T337	amod	olfactory,receptors
R410	T1292	T1290	punct	", ",family
R4100	T6716	T6712	dobj	sites,use
R4101	T6717	T6716	prep	within,sites
R4102	T6718	T6719	det	the,region
R4103	T6719	T6717	pobj	region,within
R4104	T6720	T6719	amod	coding,region
R4105	T6721	T6687	punct	.,observe
R4106	T6723	T6724	punct	(,identified
R4107	T6725	T6724	prep	For,identified
R4108	T6726	T6727	det	the,13
R4109	T6727	T6725	pobj	13,For
R411	T1293	T1290	amod	diverse,family
R4110	T6728	T6727	amod	remaining,13
R4111	T6729	T6727	prep	of,13
R4112	T6730	T6731	det	the,genes
R4113	T6731	T6729	pobj	genes,of
R4114	T6732	T6731	nummod	3,genes
R4115	T6733	T6732	punct	',3
R4116	T6734	T6731	acl	showing,genes
R4117	T6735	T6736	amod	odd,splicing
R4118	T6736	T6734	dobj	splicing,showing
R4119	T6737	T6724	punct	", ",identified
R412	T1294	T1290	compound	gene,family
R4120	T6738	T6724	nsubj	we,identified
R4121	T6739	T6724	aux	have,identified
R4122	T6740	T6741	advmod	only,cDNA
R4123	T6741	T6724	dobj	cDNA,identified
R4124	T6742	T6741	nummod	one,cDNA
R4125	T6743	T6724	cc	so,identified
R4126	T6744	T6745	aux	have,determined
R4127	T6745	T6724	conj	determined,identified
R4128	T6746	T6745	neg	not,determined
R4129	T6747	T6748	mark	whether,are
R413	T1295	T1296	poss	whose,expression
R4130	T6748	T6745	ccomp	are,determined
R4131	T6749	T6750	amod	normal,isoforms
R4132	T6750	T6748	nsubj	isoforms,are
R4133	T6751	T6748	acomp	present,are
R4134	T6752	T6724	punct	.,identified
R4135	T6753	T6724	punct	),identified
R4136	T6755	T6756	nsubjpass	We,intrigued
R4137	T6757	T6756	auxpass	were,intrigued
R4138	T6758	T6759	preconj	both,by
R4139	T6759	T6756	prep	by,intrigued
R414	T1296	T1297	dep	expression,restricted
R4140	T6760	T6761	amod	previous,reports
R4141	T6761	T6759	pobj	reports,by
R4142	T6762	T6761	prep	of,reports
R4143	T6763	T6762	pobj	splicing,of
R4144	T6764	T6763	prep	of,splicing
R4145	T6765	T6766	amod	human,receptors
R4146	T6766	T6764	pobj	receptors,of
R4147	T6767	T6766	amod	olfactory,receptors
R4148	T6768	T6763	prep	near,splicing
R4149	T6769	T6770	det	the,complex
R415	T1297	T1290	relcl	restricted,family
R4150	T6770	T6773	nmod	complex,cluster
R4151	T6771	T6770	amod	major,complex
R4152	T6772	T6770	nmod	histocompatibility,complex
R4153	T6773	T6768	pobj	cluster,near
R4154	T6774	T6770	punct	(,complex
R4155	T6775	T6770	appos	MHC,complex
R4156	T6776	T6773	punct	),cluster
R4157	T6777	T6773	punct	", ",cluster
R4158	T6778	T6779	advmod	where,splice
R4159	T6779	T6773	relcl	splice,cluster
R416	T1298	T1297	auxpass	is,restricted
R4160	T6780	T6781	amod	several,genes
R4161	T6781	T6779	nsubj	genes,splice
R4162	T6782	T6779	prep	over,splice
R4163	T6783	T6784	amod	long,distances
R4164	T6784	T6782	pobj	distances,over
R4165	T6785	T6779	prep	to,splice
R4166	T6786	T6787	det	a,exon
R4167	T6787	T6785	pobj	exon,to
R4168	T6788	T6787	amod	common,exon
R4169	T6789	T6787	amod	upstream,exon
R417	T1299	T1297	prep	to,restricted
R4170	T6790	T6791	punct	[,27
R4171	T6791	T6773	parataxis	27,cluster
R4172	T6792	T6791	nummod	26,27
R4173	T6793	T6791	punct	",",27
R4174	T6794	T6791	punct	],27
R4175	T6795	T6759	cc	and,by
R4176	T6796	T6759	conj	by,by
R4177	T6797	T6798	det	the,idea
R4178	T6798	T6796	pobj	idea,by
R4179	T6799	T6800	mark	that,achieved
R418	T1300	T1301	det	the,epithelium
R4180	T6800	T6798	acl	achieved,idea
R4181	T6801	T6802	amod	olfactory,receptor
R4182	T6802	T6803	nmod	receptor,control
R4183	T6803	T6800	nsubjpass	control,achieved
R4184	T6804	T6803	amod	transcriptional,control
R4185	T6805	T6800	aux	could,achieved
R4186	T6806	T6800	auxpass	be,achieved
R4187	T6807	T6800	prep	by,achieved
R4188	T6808	T6809	compound	DNA,mechanisms
R4189	T6809	T6807	pobj	mechanisms,by
R419	T1301	T1299	pobj	epithelium,to
R4190	T6810	T6809	compound	recombination,mechanisms
R4191	T6811	T6800	punct	", ",achieved
R4192	T6812	T6813	advmod	perhaps,with
R4193	T6813	T6800	prep	with,achieved
R4194	T6814	T6815	det	the,result
R4195	T6815	T6813	pobj	result,with
R4196	T6816	T6817	mark	that,contain
R4197	T6817	T6815	acl	contain,result
R4198	T6818	T6817	nsubj	transcripts,contain
R4199	T6819	T6817	aux	would,contain
R42	T340	T330	punct	", ",providing
R420	T1302	T1301	amod	olfactory,epithelium
R4200	T6820	T6821	det	some,sequence
R4201	T6821	T6817	dobj	sequence,contain
R4202	T6822	T6821	prep	from,sequence
R4203	T6823	T6824	det	another,locus
R4204	T6824	T6822	pobj	locus,from
R4205	T6825	T6756	punct	.,intrigued
R4206	T6827	T6828	nsubj	We,verified
R4207	T6829	T6828	advmod	therefore,verified
R4208	T6830	T6831	mark	that,matches
R4209	T6831	T6828	ccomp	matches,verified
R421	T1303	T1304	punct	[,3
R4210	T6832	T6833	det	the,sequence
R4211	T6833	T6831	nsubj	sequence,matches
R4212	T6834	T6833	amod	entire,sequence
R4213	T6835	T6833	prep	of,sequence
R4214	T6836	T6837	det	each,EST
R4215	T6837	T6835	pobj	EST,of
R4216	T6838	T6839	amod	olfactory,receptor
R4217	T6839	T6837	compound	receptor,EST
R4218	T6840	T6841	det	the,gene
R4219	T6841	T6843	poss	gene,territory
R422	T1304	T1266	parataxis	3,identified
R4220	T6842	T6841	amod	corresponding,gene
R4221	T6843	T6831	dobj	territory,matches
R4222	T6844	T6841	case	's,gene
R4223	T6845	T6843	amod	genomic,territory
R4224	T6846	T6843	punct	',territory
R4225	T6847	T6843	punct	',territory
R4226	T6848	T6843	punct	(,territory
R4227	T6849	T6843	acl	defined,territory
R4228	T6850	T6849	prep	for,defined
R4229	T6851	T6852	det	this,purpose
R423	T1305	T1304	punct	],3
R4230	T6852	T6850	pobj	purpose,for
R4231	T6853	T6849	prep	as,defined
R4232	T6854	T6853	prep	from,as
R4233	T6855	T6856	nummod	1,kb
R4234	T6856	T6857	npadvmod	kb,after
R4235	T6857	T6854	prep	after,from
R4236	T6858	T6859	det	the,gene
R4237	T6859	T6857	pobj	gene,after
R4238	T6860	T6859	amod	preceding,gene
R4239	T6861	T6854	prep	to,from
R424	T1306	T1266	punct	.,identified
R4240	T6862	T6863	nummod	1,kb
R4241	T6863	T6864	npadvmod	kb,after
R4242	T6864	T6861	prep	after,to
R4243	T6865	T6866	det	the,gene
R4244	T6866	T6867	poss	gene,codon
R4245	T6867	T6864	pobj	codon,after
R4246	T6868	T6866	case	's,gene
R4247	T6869	T6867	compound	stop,codon
R4248	T6870	T6828	punct	),verified
R4249	T6871	T6828	punct	.,verified
R425	T1308	T1309	amod	Subsequent,studies
R4250	T6873	T6874	nsubj	We,found
R4251	T6875	T6876	det	no,cDNAs
R4252	T6876	T6874	dobj	cDNAs,found
R4253	T6877	T6878	advmod	where,encompassed
R4254	T6878	T6876	relcl	encompassed,cDNAs
R4255	T6879	T6878	nsubj	introns,encompassed
R4256	T6880	T6881	amod	other,genes
R4257	T6881	T6878	dobj	genes,encompassed
R4258	T6882	T6883	amod	olfactory,receptor
R4259	T6883	T6881	compound	receptor,genes
R426	T1309	T1310	nsubj	studies,shown
R4260	T6884	T6878	punct	", ",encompassed
R4261	T6885	T6886	mark	as,reported
R4262	T6886	T6878	advcl	reported,encompassed
R4263	T6887	T6886	prep	for,reported
R4264	T6888	T6889	amod	olfactory,receptors
R4265	T6889	T6887	pobj	receptors,for
R4266	T6890	T6889	prep	in,receptors
R4267	T6891	T6892	det	the,region
R4268	T6892	T6890	pobj	region,in
R4269	T6893	T6894	amod	human,MHC
R427	T1311	T1310	aux	have,shown
R4270	T6894	T6892	compound	MHC,region
R4271	T6895	T6896	punct	[,27
R4272	T6896	T6886	parataxis	27,reported
R4273	T6897	T6896	nummod	26,27
R4274	T6898	T6896	punct	",",27
R4275	T6899	T6896	punct	],27
R4276	T6900	T6874	punct	.,found
R4277	T6902	T6903	nummod	Six,cDNAs
R4278	T6903	T6904	nsubj	cDNAs,extend
R4279	T6905	T6904	aux	do,extend
R428	T1312	T1313	mark	that,respond
R4280	T6906	T6904	advmod	further,extend
R4281	T6907	T6906	prep	than,further
R4282	T6908	T6909	det	a,gene
R4283	T6909	T6911	poss	gene,territory
R4284	T6910	T6909	amod	single,gene
R4285	T6911	T6907	pobj	territory,than
R4286	T6912	T6909	case	's,gene
R4287	T6913	T6911	punct	',territory
R4288	T6914	T6904	punct	',extend
R4289	T6915	T6904	cc	and,extend
R429	T1313	T1310	ccomp	respond,shown
R4290	T6916	T6904	conj	appear,extend
R4291	T6917	T6918	neg	not,be
R4292	T6918	T6916	xcomp	be,appear
R4293	T6919	T6918	aux	to,be
R4294	T6920	T6918	attr	artifacts,be
R4295	T6921	T6920	prep	of,artifacts
R4296	T6922	T6923	det	the,process
R4297	T6923	T6921	pobj	process,of
R4298	T6924	T6923	nmod	sequencing,process
R4299	T6925	T6924	cc	or,sequencing
R43	T341	T342	advmod	as,as
R430	T1314	T1313	nsubj	some,respond
R4300	T6926	T6924	conj	analysis,sequencing
R4301	T6927	T6904	punct	.,extend
R4302	T6929	T6930	prep	In,use
R4303	T6931	T6929	pobj	each,In
R4304	T6932	T6931	prep	of,each
R4305	T6933	T6934	det	these,cases
R4306	T6934	T6932	pobj	cases,of
R4307	T6935	T6930	punct	", ",use
R4308	T6936	T6937	det	the,clones
R4309	T6937	T6930	nsubj	clones,use
R431	T1315	T1314	prep	of,some
R4310	T6938	T6939	compound	splice,sites
R4311	T6939	T6930	dobj	sites,use
R4312	T6940	T6930	prep	within,use
R4313	T6941	T6942	det	the,UTR
R4314	T6942	T6940	pobj	UTR,within
R4315	T6943	T6942	nummod	3,UTR
R4316	T6944	T6943	punct	',3
R4317	T6945	T6930	cc	and,use
R4318	T6946	T6947	advmod	thus,extend
R4319	T6947	T6930	conj	extend,use
R432	T1316	T1317	det	these,receptors
R4320	T6948	T6949	advmod	further,downstream
R4321	T6949	T6947	advmod	downstream,extend
R4322	T6950	T6948	prep	than,further
R4323	T6951	T6952	det	the,kb
R4324	T6952	T6950	pobj	kb,than
R4325	T6953	T6952	amod	arbitrary,kb
R4326	T6954	T6952	nummod	1,kb
R4327	T6955	T6949	prep	of,downstream
R4328	T6956	T6957	det	the,codon
R4329	T6957	T6955	pobj	codon,of
R433	T1317	T1315	pobj	receptors,of
R4330	T6958	T6957	compound	stop,codon
R4331	T6959	T6930	punct	.,use
R4332	T6961	T6962	nsubj	Five,use
R4333	T6963	T6961	prep	of,Five
R4334	T6964	T6965	det	these,cDNAs
R4335	T6965	T6963	pobj	cDNAs,of
R4336	T6966	T6965	nummod	six,cDNAs
R4337	T6967	T6962	advmod	also,use
R4338	T6968	T6969	compound	splice,donor
R4339	T6969	T6971	compound	donor,sites
R434	T1318	T1313	aux	do,respond
R4340	T6970	T6969	punct	-,donor
R4341	T6971	T6962	dobj	sites,use
R4342	T6972	T6962	prep	within,use
R4343	T6973	T6974	det	the,region
R4344	T6974	T6972	pobj	region,within
R4345	T6975	T6974	amod	coding,region
R4346	T6976	T6962	cc	and,use
R4347	T6977	T6962	conj	encode,use
R4348	T6978	T6979	amod	disrupted,receptors
R4349	T6979	T6977	dobj	receptors,encode
R435	T1319	T1313	advmod	indeed,respond
R4350	T6980	T6979	amod	olfactory,receptors
R4351	T6981	T6982	punct	(,Figure
R4352	T6982	T6977	parataxis	Figure,encode
R4353	T6983	T6982	nummod	6,Figure
R4354	T6984	T6982	punct	),Figure
R4355	T6985	T6962	punct	.,use
R4356	T6987	T6988	prep	In,spliced
R4357	T6989	T6990	det	the,cDNA
R4358	T6990	T6987	pobj	cDNA,In
R4359	T6991	T6990	amod	sixth,cDNA
R436	T1320	T1313	prep	to,respond
R4360	T6992	T6988	punct	", ",spliced
R4361	T6993	T6994	det	a,intron
R4362	T6994	T6988	nsubjpass	intron,spliced
R4363	T6995	T6996	nummod	2.6,kb
R4364	T6996	T6994	compound	kb,intron
R4365	T6997	T6996	punct	-,kb
R4366	T6998	T6988	auxpass	is,spliced
R4367	T6999	T6988	prep	out,spliced
R4368	T7000	T6999	prep	of,out
R4369	T7001	T7002	det	the,UTR
R437	T1321	T1320	pobj	odorants,to
R4370	T7002	T7000	pobj	UTR,of
R4371	T7003	T7002	nummod	3,UTR
R4372	T7004	T7003	punct	',3
R4373	T7005	T6988	punct	", ",spliced
R4374	T7006	T6988	advcl	leaving,spliced
R4375	T7007	T7008	det	the,region
R4376	T7008	T7006	dobj	region,leaving
R4377	T7009	T7008	amod	coding,region
R4378	T7010	T7006	oprd	intact,leaving
R4379	T7011	T6988	punct	.,spliced
R438	T1322	T1313	cc	and,respond
R4380	T7013	T7014	mark	If,achieved
R4381	T7014	T7020	advcl	achieved,derive
R4382	T7015	T7016	amod	olfactory,receptor
R4383	T7016	T7017	nmod	receptor,control
R4384	T7017	T7014	nsubjpass	control,achieved
R4385	T7018	T7017	amod	transcriptional,control
R4386	T7019	T7014	auxpass	is,achieved
R4387	T7021	T7014	prep	by,achieved
R4388	T7022	T7023	compound	DNA,recombination
R4389	T7023	T7021	pobj	recombination,by
R439	T1323	T1324	aux	can,confer
R4390	T7024	T7020	punct	", ",derive
R4391	T7025	T7026	det	the,beginning
R4392	T7026	T7020	nsubj	beginning,derive
R4393	T7027	T7026	prep	of,beginning
R4394	T7028	T7029	det	each,transcript
R4395	T7029	T7027	pobj	transcript,of
R4396	T7030	T7020	aux	might,derive
R4397	T7031	T7020	prep	from,derive
R4398	T7032	T7033	det	a,region
R4399	T7033	T7031	pobj	region,from
R44	T342	T330	cc	as,providing
R440	T1324	T1313	conj	confer,respond
R4400	T7034	T7033	amod	donated,region
R4401	T7035	T7033	compound	promoter,region
R4402	T7036	T7020	punct	", ",derive
R4403	T7037	T7020	prep	with,derive
R4404	T7038	T7039	det	the,rest
R4405	T7039	T7040	nsubj	rest,coming
R4406	T7040	T7037	pcomp	coming,with
R4407	T7041	T7039	prep	of,rest
R4408	T7042	T7043	det	the,transcript
R4409	T7043	T7041	pobj	transcript,of
R441	T1325	T1326	det	that,responsivity
R4410	T7044	T7040	prep	from,coming
R4411	T7045	T7046	det	the,locus
R4412	T7046	T7044	pobj	locus,from
R4413	T7047	T7046	amod	native,locus
R4414	T7048	T7049	npadvmod	ORF,containing
R4415	T7049	T7046	amod	containing,locus
R4416	T7050	T7049	punct	-,containing
R4417	T7051	T7020	punct	.,derive
R4418	T7053	T7054	prep	In,analyzed
R4419	T7055	T7053	pobj	order,In
R442	T1326	T1324	dobj	responsivity,confer
R4420	T7056	T7057	aux	to,examine
R4421	T7057	T7055	acl	examine,order
R4422	T7058	T7059	det	the,hypothesis
R4423	T7059	T7057	dobj	hypothesis,examine
R4424	T7060	T7059	compound	recombination,hypothesis
R4425	T7061	T7054	punct	", ",analyzed
R4426	T7062	T7054	nsubj	we,analyzed
R4427	T7063	T7064	nummod	115,clones
R4428	T7064	T7054	dobj	clones,analyzed
R4429	T7065	T7064	compound	cDNA,clones
R443	T1327	T1328	advmod	when,expressed
R4430	T7066	T7067	prep	for,failed
R4431	T7067	T7064	relcl	failed,clones
R4432	T7068	T7066	pobj	which,for
R4433	T7069	T7067	nsubj	sim4,failed
R4434	T7070	T7071	aux	to,align
R4435	T7071	T7067	xcomp	align,failed
R4436	T7072	T7073	nummod	20,bp
R4437	T7073	T7071	dobj	bp,align
R4438	T7074	T7075	cc	or,more
R4439	T7075	T7073	nummod	more,bp
R444	T1328	T1324	advcl	expressed,confer
R4440	T7076	T7071	prep	to,align
R4441	T7077	T7078	det	the,locus
R4442	T7078	T7076	pobj	locus,to
R4443	T7079	T7078	amod	corresponding,locus
R4444	T7080	T7078	amod	genomic,locus
R4445	T7081	T7054	punct	.,analyzed
R4446	T7083	T7084	prep	In,explained
R4447	T7085	T7086	amod	most,cases
R4448	T7086	T7083	pobj	cases,In
R4449	T7087	T7084	punct	", ",explained
R445	T1329	T1328	prep	in,expressed
R4450	T7088	T7089	det	the,sequence
R4451	T7089	T7084	nsubjpass	sequence,explained
R4452	T7090	T7089	amod	missing,sequence
R4453	T7091	T7084	auxpass	was,explained
R4454	T7092	T7084	prep	by,explained
R4455	T7093	T7092	pobj	gaps,by
R4456	T7094	T7093	prep	in,gaps
R4457	T7095	T7096	det	the,sequence
R4458	T7096	T7094	pobj	sequence,in
R4459	T7097	T7096	amod	genomic,sequence
R446	T1330	T1331	amod	heterologous,types
R4460	T7098	T7092	cc	or,by
R4461	T7099	T7092	conj	by,by
R4462	T7100	T7099	pobj	matches,by
R4463	T7101	T7102	dep	that,fell
R4464	T7102	T7100	relcl	fell,matches
R4465	T7103	T7102	prep	below,fell
R4466	T7104	T7105	poss	our,cutoff
R4467	T7105	T7103	pobj	cutoff,below
R4468	T7106	T7107	compound	percent,identity
R4469	T7107	T7108	npadvmod	identity,based
R447	T1331	T1329	pobj	types,in
R4470	T7108	T7105	amod	based,cutoff
R4471	T7109	T7108	punct	-,based
R4472	T7110	T7105	prep	for,cutoff
R4473	T7111	T7110	pcomp	reporting,for
R4474	T7112	T7111	dobj	matches,reporting
R4475	T7113	T7084	punct	.,explained
R4476	T7115	T7116	prep	For,found
R4477	T7117	T7118	nummod	three,cDNAs
R4478	T7118	T7115	pobj	cDNAs,For
R4479	T7119	T7118	punct	(,cDNAs
R448	T1332	T1331	compound	cell,types
R4480	T7120	T7118	prep	from,cDNAs
R4481	T7121	T7122	nummod	three,receptors
R4482	T7122	T7120	pobj	receptors,from
R4483	T7123	T7122	amod	different,receptors
R4484	T7124	T7122	amod	olfactory,receptors
R4485	T7125	T7116	punct	),found
R4486	T7126	T7116	punct	", ",found
R4487	T7127	T7116	nsubj	we,found
R4488	T7128	T7129	mark	that,matched
R4489	T7129	T7116	ccomp	matched,found
R449	T1333	T1331	punct	(,types
R4490	T7130	T7131	det	the,piece
R4491	T7131	T7129	nsubj	piece,matched
R4492	T7132	T7131	amod	missing,piece
R4493	T7133	T7131	prep	of,piece
R4494	T7134	T7133	pobj	sequence,of
R4495	T7135	T7129	advmod	elsewhere,matched
R4496	T7136	T7135	prep	in,elsewhere
R4497	T7137	T7138	det	the,genome
R4498	T7138	T7136	pobj	genome,in
R4499	T7139	T7116	punct	.,found
R45	T343	T342	advmod	well,as
R450	T1334	T1335	prep	for,4
R4500	T7141	T7142	nsubj	Comparison,confirmed
R4501	T7143	T7141	prep	with,Comparison
R4502	T7144	T7145	det	the,assembly
R4503	T7145	T7143	pobj	assembly,with
R4504	T7146	T7145	amod	public,assembly
R4505	T7147	T7148	compound	mouse,genome
R4506	T7148	T7145	compound	genome,assembly
R4507	T7149	T7150	det	the,matches
R4508	T7150	T7142	dobj	matches,confirmed
R4509	T7151	T7150	amod	distant,matches
R451	T1335	T1331	appos	4,types
R4510	T7152	T7142	punct	.,confirmed
R4511	T7154	T7155	prep	With,conclude
R4512	T7156	T7157	predet	such,number
R4513	T7157	T7160	nsubj	number,exhibiting
R4514	T7158	T7157	det	a,number
R4515	T7159	T7157	amod	small,number
R4516	T7160	T7154	pcomp	exhibiting,With
R4517	T7161	T7157	prep	of,number
R4518	T7162	T7161	pobj	cDNAs,of
R4519	T7163	T7164	det	a,sign
R452	T1336	T1334	pobj	example,for
R4520	T7164	T7160	dobj	sign,exhibiting
R4521	T7165	T7164	amod	possible,sign
R4522	T7166	T7164	prep	of,sign
R4523	T7167	T7168	compound	DNA,recombination
R4524	T7168	T7166	pobj	recombination,of
R4525	T7169	T7164	punct	(,sign
R4526	T7170	T7171	det	a,sign
R4527	T7171	T7164	appos	sign,sign
R4528	T7172	T7173	dep	that,interpreted
R4529	T7173	T7171	relcl	interpreted,sign
R453	T1337	T1335	punct	[,4
R4530	T7174	T7173	aux	could,interpreted
R4531	T7175	T7173	advmod	also,interpreted
R4532	T7176	T7173	auxpass	be,interpreted
R4533	T7177	T7173	prep	as,interpreted
R4534	T7178	T7179	amod	chimeric,clones
R4535	T7179	T7177	pobj	clones,as
R4536	T7180	T7179	compound	cDNA,clones
R4537	T7181	T7155	punct	),conclude
R4538	T7182	T7155	punct	", ",conclude
R4539	T7183	T7155	nsubj	we,conclude
R454	T1338	T1328	punct	],expressed
R4540	T7184	T7185	mark	that,is
R4541	T7185	T7155	ccomp	is,conclude
R4542	T7186	T7187	amod	such,rearrangement
R4543	T7187	T7185	nsubj	rearrangement,is
R4544	T7188	T7185	acomp	unlikely,is
R4545	T7189	T7190	aux	to,occur
R4546	T7190	T7188	xcomp	occur,unlikely
R4547	T7191	T7155	punct	.,conclude
R4548	T7193	T7194	advmod	However,remains
R4549	T7195	T7194	punct	", ",remains
R455	T1339	T1328	punct	),expressed
R4550	T7196	T7197	det	the,possibility
R4551	T7197	T7194	nsubj	possibility,remains
R4552	T7198	T7199	mark	that,is
R4553	T7199	T7194	ccomp	is,remains
R4554	T7200	T7201	compound	DNA,recombination
R4555	T7201	T7199	nsubj	recombination,is
R4556	T7202	T7199	acomp	responsible,is
R4557	T7203	T7202	prep	for,responsible
R4558	T7204	T7205	amod	olfactory,receptor
R4559	T7205	T7206	nmod	receptor,regulation
R456	T1340	T1310	punct	.,shown
R4560	T7206	T7203	pobj	regulation,for
R4561	T7207	T7206	amod	transcriptional,regulation
R4562	T7208	T7199	punct	", ",is
R4563	T7209	T7199	prep	with,is
R4564	T7210	T7211	det	the,region
R4565	T7211	T7213	nsubj	region,contributing
R4566	T7212	T7211	amod	donated,region
R4567	T7213	T7209	pcomp	contributing,with
R4568	T7214	T7215	advmod	only,sequences
R4569	T7215	T7213	dobj	sequences,contributing
R457	T1342	T1343	amod	Recent,investigations
R4570	T7216	T7215	compound	promoter,sequences
R4571	T7217	T7215	cc	but,sequences
R4572	T7218	T7219	det	no,part
R4573	T7219	T7215	conj	part,sequences
R4574	T7220	T7219	prep	of,part
R4575	T7221	T7222	det	the,transcript
R4576	T7222	T7220	pobj	transcript,of
R4577	T7223	T7194	punct	.,remains
R4578	T7399	T7400	preconj	Both,receptors
R4579	T7400	T7403	nsubjpass	receptors,expressed
R458	T1343	T1345	nsubj	investigations,provided
R4580	T7401	T7400	amod	unclustered,receptors
R4581	T7402	T7400	amod	olfactory,receptors
R4582	T7404	T7400	cc	and,receptors
R4583	T7405	T7406	amod	olfactory,receptor
R4584	T7406	T7407	compound	receptor,pseudogenes
R4585	T7407	T7400	conj	pseudogenes,receptors
R4586	T7408	T7403	aux	can,expressed
R4587	T7409	T7403	auxpass	be,expressed
R4588	T7411	T7412	nsubj	We,were
R4589	T7413	T7412	acomp	interested,were
R459	T1344	T1343	amod	computational,investigations
R4590	T7414	T7413	prep	in,interested
R4591	T7415	T7416	mark	whether,need
R4592	T7416	T7414	pobj	need,in
R4593	T7417	T7418	amod	olfactory,receptors
R4594	T7418	T7416	nsubj	receptors,need
R4595	T7419	T7420	aux	to,be
R4596	T7420	T7416	xcomp	be,need
R4597	T7421	T7420	attr	part,be
R4598	T7422	T7421	prep	of,part
R4599	T7423	T7424	det	a,cluster
R46	T344	T330	conj	identifying,providing
R460	T1346	T1345	aux	have,provided
R4600	T7424	T7422	pobj	cluster,of
R4601	T7425	T7424	prep	in,cluster
R4602	T7426	T7427	det	the,genome
R4603	T7427	T7425	pobj	genome,in
R4604	T7428	T7420	prep	in,be
R4605	T7429	T7428	pobj	order,in
R4606	T7430	T7431	aux	to,transcribed
R4607	T7431	T7429	acl	transcribed,order
R4608	T7432	T7431	auxpass	be,transcribed
R4609	T7433	T7416	punct	", ",need
R461	T1347	T1348	det	the,catalogs
R4610	T7434	T7416	cc	or,need
R4611	T7435	T7436	mark	if,is
R4612	T7436	T7416	conj	is,need
R4613	T7437	T7438	det	the,organization
R4614	T7438	T7436	nsubj	organization,is
R4615	T7439	T7438	amod	clustered,organization
R4616	T7440	T7438	amod	genomic,organization
R4617	T7441	T7438	prep	of,organization
R4618	T7442	T7443	amod	olfactory,receptors
R4619	T7443	T7441	pobj	receptors,of
R462	T1348	T1345	dobj	catalogs,provided
R4620	T7444	T7436	advmod	simply,is
R4621	T7445	T7446	det	a,consequence
R4622	T7446	T7436	attr	consequence,is
R4623	T7447	T7446	prep	of,consequence
R4624	T7448	T7449	det	the,fact
R4625	T7449	T7447	pobj	fact,of
R4626	T7450	T7451	mark	that,is
R4627	T7451	T7449	acl	is,fact
R4628	T7452	T7453	amod	local,duplication
R4629	T7453	T7451	nsubj	duplication,is
R463	T1349	T1350	advmod	almost,complete
R4630	T7454	T7455	det	the,mechanism
R4631	T7455	T7451	attr	mechanism,is
R4632	T7456	T7455	amod	major,mechanism
R4633	T7457	T7455	prep	for,mechanism
R4634	T7458	T7457	pcomp	expanding,for
R4635	T7459	T7460	det	the,family
R4636	T7460	T7458	dobj	family,expanding
R4637	T7461	T7460	compound	gene,family
R4638	T7462	T7463	punct	[,1
R4639	T7463	T7412	parataxis	1,were
R464	T1350	T1348	amod	complete,catalogs
R4640	T7464	T7463	punct	],1
R4641	T7465	T7412	punct	.,were
R4642	T7467	T7468	punct	',are
R4643	T7469	T7470	nmod	Singleton,receptors
R4644	T7470	T7468	nsubj	receptors,are
R4645	T7471	T7470	punct	',receptors
R4646	T7472	T7470	amod	olfactory,receptors
R4647	T7473	T7470	punct	(,receptors
R4648	T7474	T7470	acl	defined,receptors
R4649	T7475	T7474	prep	as,defined
R465	T1351	T1352	amod	human,6
R4650	T7476	T7477	amod	full,length
R4651	T7477	T7479	nmod	length,receptors
R4652	T7478	T7477	punct	-,length
R4653	T7479	T7475	pobj	receptors,as
R4654	T7480	T7479	amod	olfactory,receptors
R4655	T7481	T7479	prep	without,receptors
R4656	T7482	T7483	det	another,receptor
R4657	T7483	T7481	pobj	receptor,without
R4658	T7484	T7483	amod	olfactory,receptor
R4659	T7485	T7483	prep	within,receptor
R466	T1352	T1356	parataxis	6,gene
R4660	T7486	T7487	nummod	0.5,Mb
R4661	T7487	T7485	pobj	Mb,within
R4662	T7488	T7468	punct	),are
R4663	T7489	T7490	advmod	more,often
R4664	T7490	T7468	advmod	often,are
R4665	T7491	T7468	attr	pseudogenes,are
R4666	T7492	T7493	mark	than,are
R4667	T7493	T7468	advcl	are,are
R4668	T7494	T7495	amod	olfactory,receptors
R4669	T7495	T7493	nsubj	receptors,are
R467	T1353	T1352	punct	[,6
R4670	T7496	T7495	prep	in,receptors
R4671	T7497	T7496	pobj	clusters,in
R4672	T7498	T7499	punct	(,16
R4673	T7499	T7493	parataxis	16,are
R4674	T7500	T7499	quantmod	8,16
R4675	T7501	T7499	quantmod	out,16
R4676	T7502	T7499	quantmod	of,16
R4677	T7503	T7499	cc	versus,16
R4678	T7504	T7505	quantmod	271,"1,358"
R4679	T7505	T7499	conj	"1,358",16
R468	T1354	T1352	nummod	5,6
R4680	T7506	T7505	quantmod	out,"1,358"
R4681	T7507	T7505	quantmod	of,"1,358"
R4682	T7508	T7499	punct	;,16
R4683	T7509	T7510	nsubj	χ2,8.8
R4684	T7510	T7499	ccomp	8.8,16
R4685	T7511	T7510	punct	=,8.8
R4686	T7512	T7499	punct	", ",16
R4687	T7513	T7514	nsubj	P,0.005
R4688	T7514	T7499	ccomp	0.005,16
R4689	T7515	T7514	punct	<,0.005
R469	T1355	T1352	punct	",",6
R4690	T7516	T7499	punct	),16
R4691	T7517	T7468	punct	.,are
R4692	T7519	T7520	prep	Of,have
R4693	T7521	T7522	det	the,receptors
R4694	T7522	T7519	pobj	receptors,Of
R4695	T7523	T7522	nummod	eight,receptors
R4696	T7524	T7522	amod	intact,receptors
R4697	T7525	T7522	nmod	singleton,receptors
R4698	T7526	T7522	amod	olfactory,receptors
R4699	T7527	T7520	punct	", ",have
R47	T345	T346	compound	gene,structure
R470	T1356	T1348	compound	gene,catalogs
R4700	T7528	T7520	nsubj	two,have
R4701	T7529	T7530	amod	matching,cDNAs
R4702	T7530	T7520	dobj	cDNAs,have
R4703	T7531	T7520	prep	in,have
R4704	T7532	T7533	poss	our,collection
R4705	T7533	T7531	pobj	collection,in
R4706	T7534	T7520	punct	", ",have
R4707	T7535	T7536	det	a,proportion
R4708	T7536	T7520	npadvmod	proportion,have
R4709	T7537	T7536	amod	similar,proportion
R471	T1357	T1352	punct	],6
R4710	T7538	T7539	mark	as,found
R4711	T7539	T7536	advcl	found,proportion
R4712	T7540	T7539	prep	for,found
R4713	T7541	T7542	amod	olfactory,receptors
R4714	T7542	T7540	pobj	receptors,for
R4715	T7543	T7542	prep	in,receptors
R4716	T7544	T7543	pobj	clusters,in
R4717	T7545	T7520	punct	", ",have
R4718	T7546	T7520	advcl	showing,have
R4719	T7547	T7548	mark	that,is
R472	T1358	T1352	cc	and,6
R4720	T7548	T7546	ccomp	is,showing
R4721	T7549	T7548	nsubj	clustering,is
R4722	T7550	T7548	neg	not,is
R4723	T7551	T7552	det	an,requirement
R4724	T7552	T7548	attr	requirement,is
R4725	T7553	T7552	amod	absolute,requirement
R4726	T7554	T7552	prep	for,requirement
R4727	T7555	T7556	amod	olfactory,receptor
R4728	T7556	T7557	compound	receptor,expression
R4729	T7557	T7554	pobj	expression,for
R473	T1359	T1360	dep	mouse,2
R4730	T7558	T7520	punct	.,have
R4731	T7560	T7561	advmod	However,are
R4732	T7562	T7561	punct	", ",are
R4733	T7563	T7564	nsubj	it,is
R4734	T7564	T7561	ccomp	is,are
R4735	T7565	T7564	acomp	possible,is
R4736	T7566	T7567	det	these,genes
R4737	T7567	T7571	nsubj	genes,are
R4738	T7568	T7567	nummod	two,genes
R4739	T7569	T7567	amod	expressed,genes
R474	T1360	T1352	conj	2,6
R4740	T7570	T7567	compound	singleton,genes
R4741	T7571	T7564	advcl	are,is
R4742	T7572	T7571	attr	part,are
R4743	T7573	T7572	prep	of,part
R4744	T7574	T7573	punct	',of
R4745	T7575	T7576	amod	extended,clusters
R4746	T7576	T7573	pobj	clusters,of
R4747	T7577	T7576	punct	',clusters
R4748	T7578	T7579	amod	olfactory,receptor
R4749	T7579	T7576	compound	receptor,clusters
R475	T1361	T1360	punct	[,2
R4750	T7580	T7561	punct	-,are
R4751	T7581	T7582	poss	their,neighbors
R4752	T7582	T7561	nsubj	neighbors,are
R4753	T7583	T7582	amod	nearest,neighbors
R4754	T7584	T7585	amod	olfactory,receptor
R4755	T7585	T7582	compound	receptor,neighbors
R4756	T7586	T7587	nummod	1.7,Mb
R4757	T7587	T7588	npadvmod	Mb,away
R4758	T7588	T7561	advmod	away,are
R4759	T7589	T7587	cc	and,Mb
R476	T1362	T1360	nummod	1,2
R4760	T7590	T7591	nummod	2.6,Mb
R4761	T7591	T7587	conj	Mb,Mb
R4762	T7592	T7561	punct	", ",are
R4763	T7593	T7561	advmod	respectively,are
R4764	T7594	T7561	punct	.,are
R4765	T7596	T7597	nsubj	We,find
R4766	T7598	T7597	advmod	also,find
R4767	T7599	T7600	mark	that,expressed
R4768	T7600	T7597	ccomp	expressed,find
R4769	T7601	T7602	det	some,pseudogenes
R477	T1363	T1360	punct	",",2
R4770	T7602	T7600	nsubjpass	pseudogenes,expressed
R4771	T7603	T7604	amod	olfactory,receptor
R4772	T7604	T7602	compound	receptor,pseudogenes
R4773	T7605	T7600	auxpass	are,expressed
R4774	T7606	T7600	punct	", ",expressed
R4775	T7607	T7600	prep	albeit,expressed
R4776	T7608	T7607	prep	with,albeit
R4777	T7609	T7610	det	a,probability
R4778	T7610	T7608	pobj	probability,with
R4779	T7611	T7610	amod	lower,probability
R478	T1364	T1360	punct	],2
R4780	T7612	T7610	prep	than,probability
R4781	T7613	T7614	amod	intact,receptors
R4782	T7614	T7612	pobj	receptors,than
R4783	T7615	T7614	amod	olfactory,receptors
R4784	T7616	T7597	punct	.,find
R4785	T7618	T7619	advcl	Considering,represented
R4786	T7620	T7621	det	the,sequences
R4787	T7621	T7618	dobj	sequences,Considering
R4788	T7622	T7621	nummod	"1,392",sequences
R4789	T7623	T7624	amod	olfactory,receptor
R479	T1365	T1366	amod	olfactory,receptor
R4790	T7624	T7621	compound	receptor,sequences
R4791	T7625	T7621	compound	gene,sequences
R4792	T7626	T7627	prep	for,are
R4793	T7627	T7621	relcl	are,sequences
R4794	T7628	T7626	pobj	which,for
R4795	T7629	T7630	amod	reliable,data
R4796	T7630	T7627	nsubj	data,are
R4797	T7631	T7632	amod	full,length
R4798	T7632	T7630	amod	length,data
R4799	T7633	T7632	punct	-,length
R48	T346	T344	dobj	structure,identifying
R480	T1366	T1356	compound	receptor,gene
R4800	T7634	T7627	acomp	available,are
R4801	T7635	T7619	punct	", ",represented
R4802	T7636	T7637	quantmod	15,285
R4803	T7637	T7640	nummod	285,pseudogenes
R4804	T7638	T7637	quantmod	out,285
R4805	T7639	T7637	quantmod	of,285
R4806	T7640	T7619	nsubjpass	pseudogenes,represented
R4807	T7641	T7642	punct	(,%
R4808	T7642	T7637	parataxis	%,285
R4809	T7643	T7642	nummod	5,%
R481	T1367	T1356	punct	-,gene
R4810	T7644	T7642	punct	),%
R4811	T7645	T7640	amod	apparent,pseudogenes
R4812	T7646	T7619	auxpass	are,represented
R4813	T7647	T7619	prep	in,represented
R4814	T7648	T7649	poss	our,collection
R4815	T7649	T7647	pobj	collection,in
R4816	T7650	T7649	compound	cDNA,collection
R4817	T7651	T7619	punct	", ",represented
R4818	T7652	T7619	prep	compared,represented
R4819	T7653	T7652	prep	to,compared
R482	T1368	T1345	punct	.,provided
R4820	T7654	T7655	quantmod	393,"1,107"
R4821	T7655	T7658	nummod	"1,107",receptors
R4822	T7656	T7655	quantmod	out,"1,107"
R4823	T7657	T7655	quantmod	of,"1,107"
R4824	T7658	T7653	pobj	receptors,to
R4825	T7659	T7660	punct	(,%
R4826	T7660	T7655	parataxis	%,"1,107"
R4827	T7661	T7660	nummod	36,%
R4828	T7662	T7660	punct	),%
R4829	T7663	T7658	amod	intact,receptors
R483	T1370	T1371	advmod	However,based
R4830	T7664	T7658	amod	olfactory,receptors
R4831	T7665	T7619	punct	.,represented
R4832	T7667	T7668	advmod	However,are
R4833	T7669	T7668	punct	", ",are
R4834	T7670	T7671	quantmod	three,15
R4835	T7671	T7674	nummod	15,pseudogenes
R4836	T7672	T7671	quantmod	of,15
R4837	T7673	T7671	quantmod	these,15
R4838	T7674	T7668	nsubj	pseudogenes,are
R4839	T7675	T7674	punct	',pseudogenes
R484	T1372	T1371	punct	", ",based
R4840	T7676	T7674	amod	expressed,pseudogenes
R4841	T7677	T7668	punct	',are
R4842	T7678	T7679	amod	intact,genes
R4843	T7679	T7668	attr	genes,are
R4844	T7680	T7668	prep	in,are
R4845	T7681	T7682	det	the,sequence
R4846	T7682	T7680	pobj	sequence,in
R4847	T7683	T7682	amod	public,sequence
R4848	T7684	T7685	compound	mouse,genome
R4849	T7685	T7682	compound	genome,sequence
R485	T1373	T1374	det	the,assignment
R4850	T7686	T7668	punct	.,are
R4851	T7688	T7689	det	The,defects
R4852	T7689	T7690	nsubj	defects,be
R4853	T7691	T7689	prep	in,defects
R4854	T7692	T7693	poss	Celera,version
R4855	T7693	T7691	pobj	version,in
R4856	T7694	T7692	case	's,Celera
R4857	T7695	T7693	prep	of,version
R4858	T7696	T7697	det	these,genes
R4859	T7697	T7695	pobj	genes,of
R486	T1374	T1371	nsubjpass	assignment,based
R4860	T7698	T7690	aux	may,be
R4861	T7699	T7690	prep	due,be
R4862	T7700	T7699	pcomp	to,due
R4863	T7701	T7702	amod	sequencing,errors
R4864	T7702	T7699	pobj	errors,due
R4865	T7703	T7702	cc	or,errors
R4866	T7704	T7705	amod	true,polymorphism
R4867	T7705	T7702	conj	polymorphism,errors
R4868	T7706	T7690	punct	.,be
R4869	T7708	T7709	advmod	Publicly,available
R487	T1375	T1374	prep	of,assignment
R4870	T7709	T7710	amod	available,sequence
R4871	T7710	T7712	nsubj	sequence,confirms
R4872	T7711	T7710	compound	mouse,sequence
R4873	T7713	T7714	mark	that,are
R4874	T7714	T7712	ccomp	are,confirms
R4875	T7715	T7716	quantmod	11,12
R4876	T7716	T7719	nummod	12,pseudogenes
R4877	T7717	T7716	quantmod	of,12
R4878	T7718	T7716	quantmod	the,12
R4879	T7719	T7714	nsubj	pseudogenes,are
R488	T1376	T1375	pobj	most,of
R4880	T7720	T7719	amod	remaining,pseudogenes
R4881	T7721	T7719	amod	expressed,pseudogenes
R4882	T7722	T7714	advmod	indeed,are
R4883	T7723	T7714	attr	pseudogenes,are
R4884	T7724	T7712	punct	.,confirms
R4885	T7726	T7727	det	No,sequence
R4886	T7727	T7729	nsubj	sequence,matches
R4887	T7728	T7727	amod	public,sequence
R4888	T7730	T7731	det	the,pseudogene
R4889	T7731	T7729	dobj	pseudogene,matches
R489	T1377	T1376	prep	of,most
R4890	T7732	T7731	amod	12th,pseudogene
R4891	T7733	T7731	punct	',pseudogene
R4892	T7734	T7731	amod	expressed,pseudogene
R4893	T7735	T7729	punct	',matches
R4894	T7736	T7729	prep	with,matches
R4895	T7737	T7738	nummod	99,%
R4896	T7738	T7739	compound	%,identity
R4897	T7739	T7736	pobj	identity,with
R4898	T7740	T7739	cc	or,identity
R4899	T7741	T7739	conj	more,identity
R49	T347	T346	cc	and,structure
R490	T1378	T1379	det	these,genes
R4900	T7742	T7729	punct	.,matches
R4906	T8686	T8687	nsubj	We,identified
R4907	T8688	T8687	aux	have,identified
R4908	T8689	T8687	cc	and,identified
R4909	T8690	T8687	conj	sequenced,identified
R491	T1379	T1377	pobj	genes,of
R4910	T8691	T8692	nummod	"1,264",cDNAs
R4911	T8692	T8690	dobj	cDNAs,sequenced
R4912	T8693	T8694	amod	odorant,receptor
R4913	T8694	T8692	compound	receptor,cDNAs
R4914	T8695	T8690	prep	from,sequenced
R4915	T8696	T8697	nummod	419,genes
R4916	T8697	T8695	pobj	genes,from
R4917	T8698	T8699	amod	olfactory,receptor
R4918	T8699	T8697	compound	receptor,genes
R4919	T8700	T8690	punct	", ",sequenced
R492	T1380	T1374	prep	as,assignment
R4920	T8701	T8690	conj	confirming,sequenced
R4921	T8702	T8703	poss	their,expression
R4922	T8703	T8701	dobj	expression,confirming
R4923	T8704	T8701	prep	in,confirming
R4924	T8705	T8706	det	the,epithelium
R4925	T8706	T8704	pobj	epithelium,in
R4926	T8707	T8706	amod	olfactory,epithelium
R4927	T8708	T8687	punct	.,identified
R4928	T8710	T8711	nsubj	We,validated
R4929	T8712	T8711	aux	have,validated
R493	T1381	T1382	amod	olfactory,receptors
R4930	T8713	T8711	advmod	thus,validated
R4931	T8714	T8715	det	the,prediction
R4932	T8715	T8711	dobj	prediction,validated
R4933	T8716	T8717	npadvmod	similarity,based
R4934	T8717	T8715	amod	based,prediction
R4935	T8718	T8717	punct	-,based
R4936	T8719	T8715	prep	of,prediction
R4937	T8720	T8721	quantmod	over,one
R4938	T8721	T8719	pobj	one,of
R4939	T8722	T8721	punct	-,one
R494	T1382	T1380	pobj	receptors,as
R4940	T8723	T8721	amod	third,one
R4941	T8724	T8721	prep	of,one
R4942	T8725	T8726	det	the,genes
R4943	T8726	T8724	pobj	genes,of
R4944	T8727	T8726	amod	intact,genes
R4945	T8728	T8729	amod	olfactory,receptor
R4946	T8729	T8726	compound	receptor,genes
R4947	T8730	T8726	acl	annotated,genes
R4948	T8731	T8730	prep	in,annotated
R4949	T8732	T8733	det	the,genome
R495	T1383	T1371	auxpass	is,based
R4950	T8733	T8731	pobj	genome,in
R4951	T8734	T8733	compound	mouse,genome
R4952	T8735	T8736	punct	[,2
R4953	T8736	T8715	parataxis	2,prediction
R4954	T8737	T8736	nummod	1,2
R4955	T8738	T8736	punct	",",2
R4956	T8739	T8736	punct	],2
R4957	T8740	T8711	punct	", ",validated
R4958	T8741	T8742	advmod	thereby,increasing
R4959	T8742	T8711	advcl	increasing,validated
R496	T1384	T1371	advmod	solely,based
R4960	T8743	T8742	advmod	vastly,increasing
R4961	T8744	T8745	det	the,proportion
R4962	T8745	T8742	dobj	proportion,increasing
R4963	T8746	T8745	prep	of,proportion
R4964	T8747	T8748	det	the,family
R4965	T8748	T8746	pobj	family,of
R4966	T8749	T8750	prep	for,is
R4967	T8750	T8748	relcl	is,family
R4968	T8751	T8749	pobj	which,for
R4969	T8752	T8753	amod	experimental,evidence
R497	T1385	T1371	prep	on,based
R4970	T8753	T8750	nsubj	evidence,is
R4971	T8754	T8753	prep	of,evidence
R4972	T8755	T8756	amod	olfactory,function
R4973	T8756	T8754	pobj	function,of
R4974	T8757	T8750	acomp	available,is
R4975	T8758	T8711	punct	.,validated
R4976	T8760	T8761	nsubj	We,found
R4977	T8762	T8761	aux	have,found
R4978	T8763	T8761	neg	not,found
R4979	T8764	T8761	dobj	cDNAs,found
R498	T1386	T1385	pobj	similarity,on
R4980	T8765	T8764	prep	for,cDNAs
R4981	T8766	T8767	det	all,genes
R4982	T8767	T8765	pobj	genes,for
R4983	T8768	T8769	amod	olfactory,receptor
R4984	T8769	T8767	compound	receptor,genes
R4985	T8770	T8764	cc	or,cDNAs
R4986	T8771	T8772	det	an,distribution
R4987	T8772	T8764	conj	distribution,cDNAs
R4988	T8773	T8772	advmod	even,distribution
R4989	T8774	T8772	amod	phylogenetic,distribution
R499	T1387	T1386	prep	to,similarity
R4990	T8775	T8772	prep	of,distribution
R4991	T8776	T8775	pobj	cDNAs,of
R4992	T8777	T8761	punct	", ",found
R4993	T8778	T8779	advmod	probably,are
R4994	T8779	T8761	advcl	are,found
R4995	T8780	T8779	mark	because,are
R4996	T8781	T8782	det	the,libraries
R4997	T8782	T8779	nsubj	libraries,are
R4998	T8783	T8782	cc	and,libraries
R4999	T8784	T8783	punct	/,and
R5	T301	T290	conj	is,identified
R50	T348	T349	amod	putative,regions
R500	T1388	T1387	pobj	one,to
R5000	T8785	T8783	cc	or,and
R5001	T8786	T8787	poss	our,screen
R5002	T8787	T8782	conj	screen,libraries
R5003	T8788	T8779	acomp	biased,are
R5004	T8789	T8788	prep	toward,biased
R5005	T8790	T8791	amod	certain,subfamilies
R5006	T8791	T8789	pobj	subfamilies,toward
R5007	T8792	T8793	amod	olfactory,receptor
R5008	T8793	T8791	compound	receptor,subfamilies
R5009	T8794	T8761	punct	.,found
R501	T1389	T1388	prep	of,one
R5010	T8796	T8797	advcl	Using,confirmed
R5011	T8798	T8799	compound	RT,PCR
R5012	T8799	T8796	dobj	PCR,Using
R5013	T8800	T8799	punct	-,PCR
R5014	T8801	T8796	prep	with,Using
R5015	T8802	T8803	preconj	both,degenerate
R5016	T8803	T8804	amod	degenerate,primers
R5017	T8804	T8801	pobj	primers,with
R5018	T8805	T8803	cc	and,degenerate
R5019	T8806	T8803	conj	specific,degenerate
R502	T1390	T1391	det	a,number
R5020	T8807	T8797	punct	", ",confirmed
R5021	T8808	T8797	nsubj	we,confirmed
R5022	T8809	T8797	aux	have,confirmed
R5023	T8810	T8811	amod	olfactory,expression
R5024	T8811	T8797	dobj	expression,confirmed
R5025	T8812	T8811	prep	of,expression
R5026	T8813	T8814	det	a,number
R5027	T8814	T8812	pobj	number,of
R5028	T8815	T8814	prep	of,number
R5029	T8816	T8817	amod	additional,receptors
R503	T1391	T1389	pobj	number,of
R5030	T8817	T8815	pobj	receptors,of
R5031	T8818	T8817	amod	olfactory,receptors
R5032	T8819	T8797	punct	", ",confirmed
R5033	T8820	T8797	advcl	bringing,confirmed
R5034	T8821	T8822	det	the,number
R5035	T8822	T8820	dobj	number,bringing
R5036	T8823	T8822	amod	total,number
R5037	T8824	T8822	prep	of,number
R5038	T8825	T8826	amod	olfactory,receptor
R5039	T8826	T8827	compound	receptor,genes
R504	T1392	T1393	advmod	relatively,small
R5040	T8827	T8824	pobj	genes,of
R5041	T8828	T8827	acl	verified,genes
R5042	T8829	T8828	prep	in,verified
R5043	T8830	T8831	det	this,study
R5044	T8831	T8829	pobj	study,in
R5045	T8832	T8820	prep	to,bringing
R5046	T8833	T8832	pobj	436,to
R5047	T8834	T8820	punct	", ",bringing
R5048	T8835	T8820	cc	and,bringing
R5049	T8836	T8820	conj	ensuring,bringing
R505	T1393	T1391	amod	small,number
R5050	T8837	T8838	mark	that,have
R5051	T8838	T8836	ccomp	have,ensuring
R5052	T8839	T8840	advmod	almost,clades
R5053	T8840	T8838	nsubj	clades,have
R5054	T8841	T8840	det	all,clades
R5055	T8842	T8840	amod	phylogenetic,clades
R5056	T8843	T8844	advmod	at,one
R5057	T8844	T8846	nummod	one,representative
R5058	T8845	T8844	advmod	least,one
R5059	T8846	T8838	dobj	representative,have
R506	T1394	T1391	prep	of,number
R5060	T8847	T8846	prep	with,representative
R5061	T8848	T8847	pobj	evidence,with
R5062	T8849	T8848	prep	of,evidence
R5063	T8850	T8851	amod	olfactory,function
R5064	T8851	T8849	pobj	function,of
R5065	T8852	T8797	punct	.,confirmed
R5066	T8854	T8855	nsubj	Results,suggested
R5067	T8856	T8854	prep	of,Results
R5068	T8857	T8858	poss	our,screen
R5069	T8858	T8856	pobj	screen,of
R507	T1395	T1396	advmod	experimentally,confirmed
R5070	T8859	T8858	compound	cDNA,screen
R5071	T8860	T8858	compound	library,screen
R5072	T8861	T8862	mark	that,expressed
R5073	T8862	T8855	ccomp	expressed,suggested
R5074	T8863	T8864	det	some,receptors
R5075	T8864	T8862	nsubjpass	receptors,expressed
R5076	T8865	T8864	amod	olfactory,receptors
R5077	T8866	T8862	auxpass	are,expressed
R5078	T8867	T8862	prep	at,expressed
R5079	T8868	T8869	advmod	significantly,higher
R508	T1396	T1397	amod	confirmed,receptors
R5080	T8869	T8870	amod	higher,levels
R5081	T8870	T8867	pobj	levels,at
R5082	T8871	T8870	prep	than,levels
R5083	T8872	T8871	pobj	others,than
R5084	T8873	T8855	punct	.,suggested
R5085	T8875	T8876	nsubj	We,used
R5086	T8877	T8878	amod	quantitative,PCR
R5087	T8878	T8876	dobj	PCR,used
R5088	T8879	T8880	aux	to,show
R5089	T8880	T8876	advcl	show,used
R509	T1397	T1394	pobj	receptors,of
R5090	T8881	T8882	mark	that,are
R5091	T8882	T8880	ccomp	are,show
R5092	T8883	T8884	compound	expression,levels
R5093	T8884	T8882	nsubj	levels,are
R5094	T8885	T8882	advmod	indeed,are
R5095	T8886	T8887	advmod	highly,variable
R5096	T8887	T8882	acomp	variable,are
R5097	T8888	T8882	punct	", ",are
R5098	T8889	T8890	mark	with,expressed
R5099	T8890	T8882	advcl	expressed,are
R51	T349	T346	conj	regions,structure
R510	T1398	T1397	nmod	mouse,receptors
R5100	T8891	T8892	nummod	one,receptor
R5101	T8892	T8890	nsubj	receptor,expressed
R5102	T8893	T8892	amod	olfactory,receptor
R5103	T8894	T8890	prep	at,expressed
R5104	T8895	T8896	advmod	almost,300
R5105	T8896	T8897	nummod	300,level
R5106	T8897	T8894	pobj	level,at
R5107	T8898	T8896	quantmod	times,300
R5108	T8899	T8897	det	the,level
R5109	T8900	T8897	prep	of,level
R511	T1399	T1397	amod	olfactory,receptors
R5110	T8901	T8900	pobj	another,of
R5111	T8902	T8876	punct	.,used
R5112	T8904	T8905	amod	Higher,levels
R5113	T8905	T8907	nsubj	levels,be
R5114	T8906	T8905	compound	expression,levels
R5115	T8908	T8907	aux	could,be
R5116	T8909	T8907	prep	due,be
R5117	T8910	T8909	pcomp	to,due
R5118	T8911	T8912	amod	increased,number
R5119	T8912	T8909	pobj	number,due
R512	T1400	T1385	cc	or,on
R5120	T8913	T8912	compound	transcript,number
R5121	T8914	T8912	prep	per,number
R5122	T8915	T8914	pobj	cell,per
R5123	T8916	T8912	cc	and,number
R5124	T8917	T8916	punct	/,and
R5125	T8918	T8916	cc	or,and
R5126	T8919	T8920	det	a,number
R5127	T8920	T8922	nsubj	number,choosing
R5128	T8921	T8920	amod	greater,number
R5129	T8922	T8912	conj	choosing,number
R513	T1401	T1385	punct	", ",on
R5130	T8923	T8920	prep	of,number
R5131	T8924	T8925	amod	olfactory,neurons
R5132	T8925	T8923	pobj	neurons,of
R5133	T8926	T8922	punct	',choosing
R5134	T8927	T8922	punct	',choosing
R5135	T8928	T8929	det	those,genes
R5136	T8929	T8922	dobj	genes,choosing
R5137	T8930	T8907	punct	.,be
R5138	T8932	T8933	prep	For,appear
R5139	T8934	T8935	nummod	one,pair
R514	T1402	T1403	advmod	worse,on
R5140	T8935	T8932	pobj	pair,For
R5141	T8936	T8935	prep	of,pair
R5142	T8937	T8936	pobj	genes,of
R5143	T8938	T8939	nsubj	we,tested
R5144	T8939	T8937	advcl	tested,genes
R5145	T8940	T8933	punct	", ",appear
R5146	T8941	T8942	compound	expression,level
R5147	T8942	T8943	compound	level,differences
R5148	T8943	T8933	nsubj	differences,appear
R5149	T8944	T8945	aux	to,be
R515	T1403	T1385	conj	on,on
R5150	T8945	T8933	xcomp	be,appear
R5151	T8946	T8945	prep	due,be
R5152	T8947	T8946	pcomp	to,due
R5153	T8948	T8949	det	both,factors
R5154	T8949	T8946	pobj	factors,due
R5155	T8950	T8933	punct	.,appear
R5156	T8952	T8953	nsubj	It,be
R5157	T8954	T8953	aux	would,be
R5158	T8955	T8953	acomp	interesting,be
R5159	T8956	T8957	aux	to,collect
R516	T1404	T1403	punct	", ",on
R5160	T8957	T8953	xcomp	collect,be
R5161	T8958	T8957	dobj	data,collect
R5162	T8959	T8957	prep	for,collect
R5163	T8960	T8961	amod	additional,genes
R5164	T8961	T8959	pobj	genes,for
R5165	T8962	T8963	aux	to,determine
R5166	T8963	T8957	advcl	determine,collect
R5167	T8964	T8965	advmod	how,vary
R5168	T8965	T8963	ccomp	vary,determine
R5169	T8966	T8967	det	the,numbers
R517	T1405	T1403	pobj	similarity,on
R5170	T8967	T8965	nsubj	numbers,vary
R5171	T8968	T8967	prep	of,numbers
R5172	T8969	T8970	amod	expressing,cells
R5173	T8970	T8968	pobj	cells,of
R5174	T8971	T8967	cc	and,numbers
R5175	T8972	T8973	compound	transcript,levels
R5176	T8973	T8967	conj	levels,numbers
R5177	T8974	T8973	prep	per,levels
R5178	T8975	T8974	pobj	cell,per
R5179	T8976	T8965	prep	across,vary
R518	T1406	T1405	prep	to,similarity
R5180	T8977	T8978	det	the,family
R5181	T8978	T8976	pobj	family,across
R5182	T8979	T8980	amod	olfactory,receptor
R5183	T8980	T8978	compound	receptor,family
R5184	T8981	T8953	punct	.,be
R5185	T8983	T8984	nsubj	Data,show
R5186	T8985	T8983	prep	from,Data
R5187	T8986	T8987	det	a,number
R5188	T8987	T8985	pobj	number,from
R5189	T8988	T8987	prep	of,number
R519	T1407	T1408	det	a,gene
R5190	T8989	T8990	amod	previous,studies
R5191	T8990	T8988	pobj	studies,of
R5192	T8991	T8984	advmod	also,show
R5193	T8992	T8993	mark	that,expressed
R5194	T8993	T8984	ccomp	expressed,show
R5195	T8994	T8995	amod	different,genes
R5196	T8995	T8993	nsubjpass	genes,expressed
R5197	T8996	T8997	amod	olfactory,receptor
R5198	T8997	T8995	compound	receptor,genes
R5199	T8998	T8995	punct	", ",genes
R52	T350	T349	compound	promoter,regions
R520	T1408	T1406	pobj	gene,to
R5200	T8999	T8995	cc	or,genes
R5201	T9000	T9001	advmod	even,copies
R5202	T9001	T8995	conj	copies,genes
R5203	T9002	T9001	prep	of,copies
R5204	T9003	T9004	det	the,transgene
R5205	T9004	T9002	pobj	transgene,of
R5206	T9005	T9004	amod	same,transgene
R5207	T9006	T9007	amod	olfactory,receptor
R5208	T9007	T9004	compound	receptor,transgene
R5209	T9008	T9001	prep	in,copies
R521	T1409	T1410	dep	that,defined
R5210	T9009	T9010	amod	different,locations
R5211	T9010	T9008	pobj	locations,in
R5212	T9011	T9010	amod	genomic,locations
R5213	T9012	T8993	auxpass	are,expressed
R5214	T9013	T8993	prep	in,expressed
R5215	T9014	T9015	amod	different,numbers
R5216	T9015	T9013	pobj	numbers,in
R5217	T9016	T9015	prep	of,numbers
R5218	T9017	T9016	pobj	cells,of
R5219	T9018	T9019	punct	[,35
R522	T1410	T1408	relcl	defined,gene
R5220	T9019	T8993	parataxis	35,expressed
R5221	T9020	T9019	nummod	14,35
R5222	T9021	T9019	punct	",",35
R5223	T9022	T9019	nummod	18,35
R5224	T9023	T9019	punct	",",35
R5225	T9024	T9019	punct	],35
R5226	T9025	T8993	punct	", ",expressed
R5227	T9026	T8993	cc	but,expressed
R5228	T9027	T9028	aux	do,address
R5229	T9028	T8993	conj	address,expressed
R523	T1411	T1410	prep	in,defined
R5230	T9029	T9028	neg	not,address
R5231	T9030	T9031	det	the,issue
R5232	T9031	T9028	dobj	issue,address
R5233	T9032	T9031	prep	of,issue
R5234	T9033	T9034	compound	transcript,level
R5235	T9034	T9032	pobj	level,of
R5236	T9035	T9034	prep	per,level
R5237	T9036	T9035	pobj	cell,per
R5238	T9037	T8984	punct	.,show
R5239	T9039	T9040	det	The,fact
R524	T1412	T1411	pobj	turn,in
R5240	T9040	T9041	nsubj	fact,is
R5241	T9042	T9043	mark	that,chosen
R5242	T9043	T9040	acl	chosen,fact
R5243	T9044	T9045	det	some,genes
R5244	T9045	T9043	nsubjpass	genes,chosen
R5245	T9046	T9043	auxpass	are,chosen
R5246	T9047	T9048	advmod	more,frequently
R5247	T9048	T9043	advmod	frequently,chosen
R5248	T9049	T9043	punct	", ",chosen
R5249	T9050	T9043	cc	and,chosen
R525	T1413	T1410	auxpass	was,defined
R5250	T9051	T9052	advmod	when,chosen
R5251	T9052	T9053	advcl	chosen,expressed
R5252	T9053	T9043	conj	expressed,chosen
R5253	T9054	T9053	aux	may,expressed
R5254	T9055	T9053	auxpass	be,expressed
R5255	T9056	T9053	prep	at,expressed
R5256	T9057	T9058	amod	higher,levels
R5257	T9058	T9056	pobj	levels,at
R5258	T9059	T9058	prep	per,levels
R5259	T9060	T9059	pobj	cell,per
R526	T1414	T1410	prep	as,defined
R5260	T9061	T9041	punct	", ",is
R5261	T9062	T9041	acomp	intriguing,is
R5262	T9063	T9041	prep	given,is
R5263	T9064	T9065	det	each,neuron
R5264	T9065	T9067	poss	neuron,regime
R5265	T9066	T9065	amod	olfactory,neuron
R5266	T9067	T9063	pobj	regime,given
R5267	T9068	T9065	case	's,neuron
R5268	T9069	T9070	amod	single,allele
R5269	T9070	T9067	compound	allele,regime
R527	T1415	T1416	det	an,receptor
R5270	T9071	T9070	punct	-,allele
R5271	T9072	T9067	compound	expression,regime
R5272	T9073	T9041	punct	.,is
R5273	T9075	T9076	det	The,observation
R5274	T9076	T9077	nsubj	observation,leads
R5275	T9078	T9076	prep	of,observation
R5276	T9079	T9080	amod	unequal,expression
R5277	T9080	T9078	pobj	expression,of
R5278	T9081	T9077	prep	to,leads
R5279	T9082	T9083	det	a,number
R528	T1416	T1414	pobj	receptor,as
R5280	T9083	T9081	pobj	number,to
R5281	T9084	T9083	prep	of,number
R5282	T9085	T9084	pobj	questions,of
R5283	T9086	T9077	punct	.,leads
R5284	T9088	T9089	nsubjpass	It,known
R5285	T9089	T9091	ccomp	known,choose
R5286	T9090	T9089	auxpass	is,known
R5287	T9092	T9093	mark	that,expressed
R5288	T9093	T9089	ccomp	expressed,known
R5289	T9094	T9095	det	each,receptor
R529	T1417	T1416	amod	olfactory,receptor
R5290	T9095	T9093	nsubjpass	receptor,expressed
R5291	T9096	T9095	amod	olfactory,receptor
R5292	T9097	T9093	auxpass	is,expressed
R5293	T9098	T9093	prep	in,expressed
R5294	T9099	T9098	pobj	one,in
R5295	T9100	T9099	prep	of,one
R5296	T9101	T9102	nummod	four,zones
R5297	T9102	T9100	pobj	zones,of
R5298	T9103	T9102	prep	of,zones
R5299	T9104	T9105	det	the,epithelium
R53	T351	T314	punct	.,screened
R530	T1418	T1419	advmod	solely,by
R5300	T9105	T9103	pobj	epithelium,of
R5301	T9106	T9105	amod	olfactory,epithelium
R5302	T9107	T9108	punct	[,15
R5303	T9108	T9093	parataxis	15,expressed
R5304	T9109	T9108	nummod	14,15
R5305	T9110	T9108	punct	",",15
R5306	T9111	T9108	punct	],15
R5307	T9112	T9091	punct	;,choose
R5308	T9113	T9091	aux	do,choose
R5309	T9114	T9115	det	some,zones
R531	T1419	T1410	prep	by,defined
R5310	T9115	T9091	nsubj	zones,choose
R5311	T9116	T9091	prep	from,choose
R5312	T9117	T9118	det	a,sub-repertoire
R5313	T9118	T9116	pobj	sub-repertoire,from
R5314	T9119	T9118	amod	smaller,sub-repertoire
R5315	T9120	T9121	amod	olfactory,receptor
R5316	T9121	T9118	compound	receptor,sub-repertoire
R5317	T9122	T9091	cc	and,choose
R5318	T9123	T9124	advmod	thus,express
R5319	T9124	T9091	conj	express,choose
R532	T1420	T1419	pobj	similarity,by
R5320	T9125	T9126	det	each,receptor
R5321	T9126	T9124	dobj	receptor,express
R5322	T9127	T9126	amod	olfactory,receptor
R5323	T9128	T9124	prep	in,express
R5324	T9129	T9130	det	a,number
R5325	T9130	T9128	pobj	number,in
R5326	T9131	T9130	amod	larger,number
R5327	T9132	T9130	prep	of,number
R5328	T9133	T9132	pobj	cells,of
R5329	T9134	T9091	punct	?,choose
R533	T1421	T1371	punct	.,based
R5330	T9136	T9137	nsubj	We,note
R5331	T9138	T9139	mark	that,expressed
R5332	T9139	T9137	ccomp	expressed,note
R5333	T9140	T9141	amod	several,receptors
R5334	T9141	T9139	nsubjpass	receptors,expressed
R5335	T9142	T9143	advmod	apparently,expressed
R5336	T9143	T9141	amod	expressed,receptors
R5337	T9144	T9143	advmod	highly,expressed
R5338	T9145	T9141	amod	olfactory,receptors
R5339	T9146	T9141	punct	(,receptors
R534	T1423	T1424	mark	While,is
R5340	T9147	T9148	compound	gene,A
R5341	T9148	T9141	appos	A,receptors
R5342	T9149	T9150	punct	", ",study
R5343	T9150	T9148	parataxis	study,A
R5344	T9151	T9150	det	this,study
R5345	T9152	T9150	punct	", ",study
R5346	T9153	T9148	cc	and,A
R5347	T9154	T9148	conj	MOR10,A
R5348	T9155	T9154	cc	and,MOR10
R5349	T9156	T9154	conj	MOR28,MOR10
R535	T1424	T1430	advcl	is,be
R5350	T9157	T9158	punct	[,36
R5351	T9158	T9156	parataxis	36,MOR28
R5352	T9159	T9158	punct	],36
R5353	T9160	T9141	punct	),receptors
R5354	T9161	T9139	auxpass	are,expressed
R5355	T9162	T9139	prep	in,expressed
R5356	T9163	T9162	pobj	zone,in
R5357	T9164	T9163	nummod	4,zone
R5358	T9165	T9163	prep	of,zone
R5359	T9166	T9167	det	the,epithelium
R536	T1425	T1426	npadvmod	similarity,based
R5360	T9167	T9165	pobj	epithelium,of
R5361	T9168	T9167	amod	olfactory,epithelium
R5362	T9169	T9137	punct	.,note
R5363	T9171	T9172	aux	Does,contribute
R5364	T9173	T9174	npadvmod	activity,dependent
R5365	T9174	T9176	amod	dependent,competition
R5366	T9175	T9174	punct	-,dependent
R5367	T9176	T9172	nsubj	competition,contribute
R5368	T9177	T9176	amod	neuronal,competition
R5369	T9178	T9179	punct	[,37
R537	T1426	T1428	amod	based,annotation
R5370	T9179	T9176	parataxis	37,competition
R5371	T9180	T9179	punct	],37
R5372	T9181	T9172	prep	to,contribute
R5373	T9182	T9183	amod	increased,representation
R5374	T9183	T9181	pobj	representation,to
R5375	T9184	T9183	prep	of,representation
R5376	T9185	T9186	det	the,receptors
R5377	T9186	T9184	pobj	receptors,of
R5378	T9187	T9186	amod	olfactory,receptors
R5379	T9188	T9189	dep	that,respond
R538	T1427	T1426	punct	-,based
R5380	T9189	T9186	relcl	respond,receptors
R5381	T9190	T9189	prep	to,respond
R5382	T9191	T9192	amod	common,odorants
R5383	T9192	T9190	pobj	odorants,to
R5384	T9193	T9192	amod	environmental,odorants
R5385	T9194	T9172	punct	?,contribute
R5386	T9196	T9197	aux	Do,have
R5387	T9198	T9199	det	the,receptors
R5388	T9199	T9197	nsubj	receptors,have
R5389	T9200	T9199	amod	favored,receptors
R539	T1428	T1424	nsubj	annotation,is
R5390	T9201	T9199	amod	olfactory,receptors
R5391	T9202	T9203	amod	stronger,sequences
R5392	T9203	T9197	dobj	sequences,have
R5393	T9204	T9203	compound	promoter,sequences
R5394	T9205	T9197	punct	?,have
R5395	T9208	T9209	det	some,mRNAs
R5396	T9209	T9207	nsubj	mRNAs,Are
R5397	T9210	T9211	amod	olfactory,receptor
R5398	T9211	T9209	compound	receptor,mRNAs
R5399	T9212	T9213	advmod	more,stable
R54	T355	T356	nsubj	We,identified
R540	T1429	T1428	compound	genome,annotation
R5400	T9213	T9207	acomp	stable,Are
R5401	T9214	T9213	prep	than,stable
R5402	T9215	T9214	pobj	others,than
R5403	T9216	T9207	punct	", ",Are
R5404	T9217	T9207	advcl	leading,Are
R5405	T9218	T9217	prep	to,leading
R5406	T9219	T9220	amod	higher,levels
R5407	T9220	T9218	pobj	levels,to
R5408	T9221	T9220	compound	transcript,levels
R5409	T9222	T9220	prep	per,levels
R541	T1431	T1432	det	a,method
R5410	T9223	T9224	amod	expressing,cell
R5411	T9224	T9222	pobj	cell,per
R5412	T9225	T9207	punct	?,Are
R5413	T9228	T9229	det	the,receptors
R5414	T9229	T9227	nsubj	receptors,Are
R5415	T9230	T9229	amod	favored,receptors
R5416	T9231	T9229	amod	olfactory,receptors
R5417	T9232	T9227	prep	in,Are
R5418	T9233	T9234	advmod	more,open
R5419	T9234	T9235	amod	open,conformation
R542	T1432	T1424	attr	method,is
R5420	T9235	T9232	pobj	conformation,in
R5421	T9236	T9235	compound	chromatin,conformation
R5422	T9237	T9235	cc	or,conformation
R5423	T9238	T9239	advmod	more,accessible
R5424	T9239	T9240	amod	accessible,locations
R5425	T9240	T9235	conj	locations,conformation
R5426	T9241	T9240	amod	genomic,locations
R5427	T9242	T9227	punct	?,Are
R5428	T9244	T9245	nsubj	Transcription,suggests
R5429	T9246	T9244	prep	of,Transcription
R543	T1433	T1432	amod	good,method
R5430	T9247	T9248	amod	apparent,genes
R5431	T9248	T9246	pobj	genes,of
R5432	T9249	T9248	punct	',genes
R5433	T9250	T9248	nmod	singleton,genes
R5434	T9251	T9248	punct	',genes
R5435	T9252	T9253	amod	olfactory,receptor
R5436	T9253	T9248	compound	receptor,genes
R5437	T9254	T9255	punct	(,Mb
R5438	T9255	T9244	parataxis	Mb,Transcription
R5439	T9256	T9255	nummod	0.5,Mb
R544	T1434	T1432	amod	initial,method
R5440	T9257	T9258	cc	or,more
R5441	T9258	T9255	nummod	more,Mb
R5442	T9259	T9255	prep	from,Mb
R5443	T9260	T9261	det	the,gene
R5444	T9261	T9259	pobj	gene,from
R5445	T9262	T9261	amod	nearest,gene
R5446	T9263	T9261	amod	other,gene
R5447	T9264	T9265	amod	olfactory,receptor
R5448	T9265	T9261	compound	receptor,gene
R5449	T9266	T9255	punct	),Mb
R545	T1435	T1436	aux	to,identify
R5450	T9267	T9268	mark	that,is
R5451	T9268	T9245	ccomp	is,suggests
R5452	T9269	T9268	expl	there,is
R5453	T9270	T9271	det	no,requirement
R5454	T9271	T9268	attr	requirement,is
R5455	T9272	T9271	amod	absolute,requirement
R5456	T9273	T9271	prep	for,requirement
R5457	T9274	T9275	amod	genomic,clustering
R5458	T9275	T9273	pobj	clustering,for
R5459	T9276	T9277	mark	for,transcribed
R546	T1436	T1432	advcl	identify,method
R5460	T9277	T9268	advcl	transcribed,is
R5461	T9278	T9279	det	an,receptor
R5462	T9279	T9277	nsubjpass	receptor,transcribed
R5463	T9280	T9279	amod	olfactory,receptor
R5464	T9281	T9277	aux	to,transcribed
R5465	T9282	T9277	auxpass	be,transcribed
R5466	T9283	T9268	punct	", ",is
R5467	T9284	T9268	advcl	consistent,is
R5468	T9285	T9284	prep	with,consistent
R5469	T9286	T9285	pobj	observations,with
R547	T1437	T1436	dobj	genes,identify
R5470	T9287	T9288	mark	that,expressed
R5471	T9288	T9286	acl	expressed,observations
R5472	T9289	T9290	amod	small,transgenes
R5473	T9290	T9288	nsubjpass	transgenes,expressed
R5474	T9291	T9292	amod	olfactory,receptor
R5475	T9292	T9290	compound	receptor,transgenes
R5476	T9293	T9288	aux	can,expressed
R5477	T9294	T9288	auxpass	be,expressed
R5478	T9295	T9288	advmod	correctly,expressed
R5479	T9296	T9297	advmod	when,integrated
R548	T1438	T1436	cc	and,identify
R5480	T9297	T9288	advcl	integrated,expressed
R5481	T9298	T9297	prep	outside,integrated
R5482	T9299	T9300	amod	native,clusters
R5483	T9300	T9298	pobj	clusters,outside
R5484	T9301	T9302	amod	olfactory,receptor
R5485	T9302	T9300	compound	receptor,clusters
R5486	T9303	T9304	punct	[,35
R5487	T9304	T9245	parataxis	35,suggests
R5488	T9305	T9304	punct	],35
R5489	T9306	T9245	punct	.,suggests
R549	T1439	T1436	conj	predict,identify
R5490	T9308	T9309	advmod	However,suggests
R5491	T9310	T9309	punct	", ",suggests
R5492	T9311	T9312	det	the,count
R5493	T9312	T9309	nsubj	count,suggests
R5494	T9313	T9312	amod	high,count
R5495	T9314	T9312	compound	pseudogene,count
R5496	T9315	T9312	prep	among,count
R5497	T9316	T9317	nmod	singleton,genes
R5498	T9317	T9315	pobj	genes,among
R5499	T9318	T9319	amod	olfactory,receptor
R55	T357	T358	amod	more,"1,200"
R550	T1440	T1441	poss	their,function
R5500	T9319	T9317	compound	receptor,genes
R5501	T9320	T9312	punct	(,count
R5502	T9321	T9322	nummod	50,%
R5503	T9322	T9312	appos	%,count
R5504	T9323	T9322	punct	", ",%
R5505	T9324	T9322	cc	versus,%
R5506	T9325	T9326	nummod	20,%
R5507	T9326	T9322	conj	%,%
R5508	T9327	T9326	prep	for,%
R5509	T9328	T9329	amod	clustered,genes
R551	T1441	T1439	dobj	function,predict
R5510	T9329	T9327	pobj	genes,for
R5511	T9330	T9331	amod	olfactory,receptor
R5512	T9331	T9329	compound	receptor,genes
R5513	T9332	T9312	punct	),count
R5514	T9333	T9334	mark	that,are
R5515	T9334	T9309	ccomp	are,suggests
R5516	T9335	T9336	neg	not,locations
R5517	T9336	T9334	nsubj	locations,are
R5518	T9337	T9336	det	all,locations
R5519	T9338	T9336	amod	genomic,locations
R552	T1442	T1430	punct	", ",be
R5520	T9339	T9334	acomp	favorable,are
R5521	T9340	T9339	prep	for,favorable
R5522	T9341	T9342	amod	olfactory,receptor
R5523	T9342	T9343	compound	receptor,survival
R5524	T9343	T9340	pobj	survival,for
R5525	T9344	T9343	compound	gene,survival
R5526	T9345	T9334	punct	", ",are
R5527	T9346	T9347	advmod	perhaps,due
R5528	T9347	T9334	prep	due,are
R5529	T9348	T9347	pcomp	to,due
R553	T1443	T1430	prep	in,be
R5530	T9349	T9350	amod	transcriptional,constraints
R5531	T9350	T9347	pobj	constraints,due
R5532	T9351	T9309	punct	.,suggests
R5533	T9353	T9354	nsubj	It,is
R5534	T9354	T9355	ccomp	is,rescue
R5535	T9356	T9354	advmod	also,is
R5536	T9357	T9354	acomp	possible,is
R5537	T9358	T9359	mark	that,be
R5538	T9359	T9354	ccomp	be,is
R5539	T9360	T9361	amod	evolutionary,factors
R554	T1444	T1445	det	some,cases
R5540	T9361	T9359	nsubj	factors,be
R5541	T9362	T9359	aux	may,be
R5542	T9363	T9359	acomp	responsible,be
R5543	T9364	T9363	prep	for,responsible
R5544	T9365	T9366	amod	reduced,content
R5545	T9366	T9364	pobj	content,for
R5546	T9367	T9366	compound	pseudogene,content
R5547	T9368	T9366	prep	of,content
R5548	T9369	T9370	amod	clustered,receptors
R5549	T9370	T9368	pobj	receptors,of
R555	T1445	T1443	pobj	cases,in
R5550	T9371	T9370	amod	olfactory,receptors
R5551	T9372	T9355	punct	-,rescue
R5552	T9373	T9374	compound	gene,conversion
R5553	T9374	T9355	nsubj	conversion,rescue
R5554	T9375	T9374	prep	between,conversion
R5555	T9376	T9377	amod	neighboring,receptors
R5556	T9377	T9375	pobj	receptors,between
R5557	T9378	T9377	amod	olfactory,receptors
R5558	T9379	T9355	aux	could,rescue
R5559	T9380	T9381	amod	inactivating,mutations
R556	T1446	T1430	nsubj	it,be
R5560	T9381	T9355	dobj	mutations,rescue
R5561	T9382	T9381	prep	in,mutations
R5562	T9383	T9384	amod	clustered,genes
R5563	T9384	T9382	pobj	genes,in
R5564	T9385	T9381	punct	", ",mutations
R5565	T9386	T9381	cc	but,mutations
R5566	T9387	T9386	neg	not,but
R5567	T9388	T9381	conj	singletons,mutations
R5568	T9389	T9355	punct	.,rescue
R5569	T9391	T9392	mark	Before,answered
R557	T1447	T1430	aux	can,be
R5570	T9392	T9402	advcl	answered,be
R5571	T9393	T9394	det	these,questions
R5572	T9394	T9392	nsubjpass	questions,answered
R5573	T9395	T9394	prep	about,questions
R5574	T9396	T9397	amod	olfactory,receptor
R5575	T9397	T9398	compound	receptor,gene
R5576	T9398	T9399	compound	gene,choice
R5577	T9399	T9395	pobj	choice,about
R5578	T9400	T9392	aux	can,answered
R5579	T9401	T9392	auxpass	be,answered
R558	T1448	T1430	acomp	misleading,be
R5580	T9403	T9402	punct	", ",be
R5581	T9404	T9402	nsubj	it,be
R5582	T9405	T9402	aux	will,be
R5583	T9406	T9402	acomp	important,be
R5584	T9407	T9408	aux	to,measure
R5585	T9408	T9402	xcomp	measure,be
R5586	T9409	T9410	compound	expression,levels
R5587	T9410	T9408	dobj	levels,measure
R5588	T9411	T9410	prep	of,levels
R5589	T9412	T9413	det	a,number
R559	T1449	T1430	punct	", ",be
R5590	T9413	T9411	pobj	number,of
R5591	T9414	T9413	amod	larger,number
R5592	T9415	T9413	prep	of,number
R5593	T9416	T9415	pobj	genes,of
R5594	T9417	T9408	punct	", ",measure
R5595	T9418	T9419	advmod	perhaps,using
R5596	T9419	T9408	advcl	using,measure
R5597	T9420	T9421	det	an,microarray
R5598	T9421	T9419	dobj	microarray,using
R5599	T9422	T9423	amod	olfactory,receptor
R56	T358	T360	nummod	"1,200",cDNAs
R560	T1450	T1451	mark	as,carry
R5600	T9423	T9424	compound	receptor,gene
R5601	T9424	T9421	compound	gene,microarray
R5602	T9425	T9402	punct	.,be
R5603	T9427	T9428	poss	Our,study
R5604	T9428	T9429	nsubj	study,provides
R5605	T9430	T9431	advmod	at,least
R5606	T9431	T9432	advmod	least,partial
R5607	T9432	T9433	amod	partial,data
R5608	T9433	T9429	dobj	data,provides
R5609	T9434	T9433	prep	about,data
R561	T1451	T1430	advcl	carry,be
R5610	T9435	T9436	det	the,structures
R5611	T9436	T9434	pobj	structures,about
R5612	T9437	T9436	amod	upstream,structures
R5613	T9438	T9436	compound	transcript,structures
R5614	T9439	T9436	prep	of,structures
R5615	T9440	T9441	quantmod	over,300
R5616	T9441	T9442	nummod	300,genes
R5617	T9442	T9439	pobj	genes,of
R5618	T9443	T9444	amod	olfactory,receptor
R5619	T9444	T9442	compound	receptor,genes
R562	T1452	T1451	nsubj	genes,carry
R5620	T9445	T9429	punct	.,provides
R5621	T9447	T9448	det	These,data
R5622	T9448	T9449	nsubj	data,provide
R5623	T9450	T9451	amod	tentative,locations
R5624	T9451	T9449	dobj	locations,provide
R5625	T9452	T9451	prep	of,locations
R5626	T9453	T9454	det	a,set
R5627	T9454	T9452	pobj	set,of
R5628	T9455	T9454	amod	large,set
R5629	T9456	T9454	prep	of,set
R563	T1453	T1452	prep	of,genes
R5630	T9457	T9458	compound	promoter,regions
R5631	T9458	T9456	pobj	regions,of
R5632	T9459	T9449	punct	", ",provide
R5633	T9460	T9449	advcl	allowing,provide
R5634	T9461	T9462	amod	computational,searches
R5635	T9462	T9460	dobj	searches,allowing
R5636	T9463	T9462	prep	for,searches
R5637	T9464	T9465	amod	shared,motifs
R5638	T9465	T9463	pobj	motifs,for
R5639	T9466	T9465	compound	sequence,motifs
R564	T1454	T1455	amod	similar,sequence
R5640	T9467	T9468	dep	that,involved
R5641	T9468	T9465	relcl	involved,motifs
R5642	T9469	T9468	aux	might,involved
R5643	T9470	T9468	auxpass	be,involved
R5644	T9471	T9468	prep	in,involved
R5645	T9472	T9473	det	the,regulation
R5646	T9473	T9471	pobj	regulation,in
R5647	T9474	T9473	amod	intriguing,regulation
R5648	T9475	T9473	amod	transcriptional,regulation
R5649	T9476	T9473	prep	of,regulation
R565	T1455	T1453	pobj	sequence,of
R5650	T9477	T9478	amod	olfactory,receptors
R5651	T9478	T9476	pobj	receptors,of
R5652	T9479	T9449	punct	.,provide
R5653	T9481	T9482	advmod	However,be
R5654	T9483	T9482	punct	", ",be
R5655	T9484	T9482	prep	given,be
R5656	T9485	T9486	mark	that,are
R5657	T9486	T9484	pcomp	are,given
R5658	T9487	T9488	neg	not,cDNAs
R5659	T9488	T9486	nsubj	cDNAs,are
R566	T1456	T1451	aux	can,carry
R5660	T9489	T9488	det	all,cDNAs
R5661	T9490	T9491	amod	full,length
R5662	T9491	T9493	compound	length,clones
R5663	T9492	T9491	punct	-,length
R5664	T9493	T9486	attr	clones,are
R5665	T9494	T9482	punct	", ",be
R5666	T9495	T9482	nsubj	some,be
R5667	T9496	T9495	prep	of,some
R5668	T9497	T9498	det	these,candidates
R5669	T9498	T9496	pobj	candidates,of
R567	T1457	T1451	prt	out,carry
R5670	T9499	T9482	aux	will,be
R5671	T9500	T9482	neg	not,be
R5672	T9501	T9502	amod	true,regions
R5673	T9502	T9482	attr	regions,be
R5674	T9503	T9502	compound	promoter,regions
R5675	T9504	T9482	punct	.,be
R5676	T9506	T9507	det	The,sequences
R5677	T9507	T9511	nsubj	sequences,aid
R5678	T9508	T9509	nummod	5,UTR
R5679	T9509	T9507	compound	UTR,sequences
R568	T1458	T1459	amod	different,functions
R5680	T9510	T9508	punct	',5
R5681	T9512	T9513	nsubj	we,obtained
R5682	T9513	T9507	advcl	obtained,sequences
R5683	T9514	T9511	aux	will,aid
R5684	T9515	T9511	advmod	also,aid
R5685	T9516	T9511	prep	in,aid
R5686	T9517	T9518	det	the,design
R5687	T9518	T9516	pobj	design,in
R5688	T9519	T9518	prep	of,design
R5689	T9520	T9521	amod	experimental,probes
R569	T1459	T1451	dobj	functions,carry
R5690	T9521	T9519	pobj	probes,of
R5691	T9522	T9511	punct	", ",aid
R5692	T9523	T9511	prep	for,aid
R5693	T9524	T9523	pobj	example,for
R5694	T9525	T9523	punct	", ",for
R5695	T9526	T9523	conj	for,for
R5696	T9527	T9528	advmod	in,situ
R5697	T9528	T9529	amod	situ,hybridizations
R5698	T9529	T9526	pobj	hybridizations,for
R5699	T9530	T9526	cc	or,for
R57	T359	T358	quantmod	than,"1,200"
R570	T1460	T1451	cc	and,carry
R5700	T9531	T9532	aux	to,immobilize
R5701	T9532	T9526	conj	immobilize,for
R5702	T9533	T9532	prep	on,immobilize
R5703	T9534	T9535	det	an,microarray
R5704	T9535	T9533	pobj	microarray,on
R5705	T9536	T9537	amod	olfactory,receptor
R5706	T9537	T9535	compound	receptor,microarray
R5707	T9538	T9511	punct	.,aid
R5708	T9540	T9541	nsubj	One,be
R5709	T9542	T9540	prep	of,One
R571	T1461	T1462	auxpass	be,expressed
R5710	T9543	T9544	det	the,challenges
R5711	T9544	T9542	pobj	challenges,of
R5712	T9545	T9544	prep	of,challenges
R5713	T9546	T9547	predet	such,array
R5714	T9547	T9545	pobj	array,of
R5715	T9548	T9547	det	an,array
R5716	T9549	T9541	aux	will,be
R5717	T9550	T9551	aux	to,design
R5718	T9551	T9541	xcomp	design,be
R5719	T9552	T9553	amod	unique,probes
R572	T1462	T1451	conj	expressed,carry
R5720	T9553	T9551	dobj	probes,design
R5721	T9554	T9555	prep	with,represent
R5722	T9555	T9553	relcl	represent,probes
R5723	T9556	T9554	pobj	which,with
R5724	T9557	T9555	aux	to,represent
R5725	T9558	T9559	det	each,gene
R5726	T9559	T9555	dobj	gene,represent
R5727	T9560	T9541	punct	.,be
R5728	T9562	T9563	advmod	Often,is
R5729	T9564	T9563	punct	", ",is
R573	T1463	T1462	prep	in,expressed
R5730	T9565	T9566	det	the,region
R5731	T9566	T9563	nsubj	region,is
R5732	T9567	T9566	amod	coding,region
R5733	T9568	T9566	prep	of,region
R5734	T9569	T9570	amod	olfactory,receptors
R5735	T9570	T9568	pobj	receptors,of
R5736	T9571	T9572	advmod	highly,similar
R5737	T9572	T9563	acomp	similar,is
R5738	T9573	T9563	prep	between,is
R5739	T9574	T9575	advmod	recently,duplicated
R574	T1464	T1465	amod	different,tissues
R5740	T9575	T9576	amod	duplicated,genes
R5741	T9576	T9573	pobj	genes,between
R5742	T9577	T9563	punct	.,is
R5743	T9579	T9580	amod	Many,pairs
R5744	T9580	T9581	nsubj	pairs,show
R5745	T9582	T9580	prep	of,pairs
R5746	T9583	T9584	amod	similar,receptors
R5747	T9584	T9582	pobj	receptors,of
R5748	T9585	T9584	amod	olfactory,receptors
R5749	T9586	T9587	amod	more,divergence
R575	T1465	T1463	pobj	tissues,in
R5750	T9587	T9581	dobj	divergence,show
R5751	T9588	T9587	compound	sequence,divergence
R5752	T9589	T9581	prep	in,show
R5753	T9590	T9591	det	the,UTRs
R5754	T9591	T9589	pobj	UTRs,in
R5755	T9592	T9581	prep	than,show
R5756	T9593	T9594	det	the,region
R5757	T9594	T9592	pobj	region,than
R5758	T9595	T9596	npadvmod	protein,coding
R5759	T9596	T9594	amod	coding,region
R576	T1466	T1465	punct	(,tissues
R5760	T9597	T9596	punct	-,coding
R5761	T9598	T9599	punct	(,J.Y.
R5762	T9599	T9581	meta	J.Y.,show
R5763	T9600	T9599	punct	", ",J.Y.
R5764	T9601	T9599	amod	unpublished,J.Y.
R5765	T9602	T9599	nmod	observations,J.Y.
R5766	T9603	T9599	punct	),J.Y.
R5767	T9604	T9581	punct	.,show
R5768	T9606	T9607	det	The,UTRs
R5769	T9607	T9608	nsubj	UTRs,make
R577	T1467	T1468	prep	for,family
R5770	T9609	T9608	aux	would,make
R5771	T9610	T9608	advmod	therefore,make
R5772	T9611	T9612	det	a,choice
R5773	T9612	T9608	dobj	choice,make
R5774	T9613	T9612	amod	better,choice
R5775	T9614	T9612	prep	of,choice
R5776	T9615	T9614	pobj	sequence,of
R5777	T9616	T9617	prep	from,design
R5778	T9617	T9615	relcl	design,sequence
R5779	T9618	T9616	pobj	which,from
R578	T1468	T1465	appos	family,tissues
R5780	T9619	T9617	aux	to,design
R5781	T9620	T9621	amod	unique,oligonucleotides
R5782	T9621	T9617	dobj	oligonucleotides,design
R5783	T9622	T9623	aux	to,distinguish
R5784	T9623	T9617	advcl	distinguish,design
R5785	T9624	T9625	advmod	closely,related
R5786	T9625	T9626	amod	related,genes
R5787	T9626	T9623	dobj	genes,distinguish
R5788	T9627	T9628	amod	olfactory,receptor
R5789	T9628	T9626	compound	receptor,genes
R579	T1469	T1467	pobj	example,for
R5790	T9629	T9608	punct	.,make
R5791	T9631	T9632	nsubj	Locations,are
R5792	T9632	T9639	ccomp	are,provides
R5793	T9633	T9631	prep	of,Locations
R5794	T9634	T9635	det	these,regions
R5795	T9635	T9633	pobj	regions,of
R5796	T9636	T9631	prep	in,Locations
R5797	T9637	T9638	amod	genomic,sequence
R5798	T9638	T9636	pobj	sequence,in
R5799	T9640	T9632	acomp	difficult,are
R58	T360	T356	dobj	cDNAs,identified
R580	T1470	T1468	punct	", ",family
R5800	T9641	T9642	aux	to,predict
R5801	T9642	T9640	advcl	predict,difficult
R5802	T9643	T9639	punct	-,provides
R5803	T9644	T9645	poss	our,study
R5804	T9645	T9639	nsubj	study,provides
R5805	T9646	T9647	nummod	5,UTR
R5806	T9647	T9649	compound	UTR,sequences
R5807	T9648	T9646	punct	',5
R5808	T9649	T9639	dobj	sequences,provides
R5809	T9650	T9649	prep	of,sequences
R581	T1471	T1468	det	the,family
R5810	T9651	T9652	nummod	343,genes
R5811	T9652	T9650	pobj	genes,of
R5812	T9653	T9649	cc	and,sequences
R5813	T9654	T9655	det	the,length
R5814	T9655	T9649	conj	length,sequences
R5815	T9656	T9655	amod	approximate,length
R5816	T9657	T9658	nummod	3,UTR
R5817	T9658	T9655	compound	UTR,length
R5818	T9659	T9657	punct	',3
R5819	T9660	T9655	prep	for,length
R582	T1472	T1473	compound	sugar,transporter
R5820	T9661	T9662	nummod	399,genes
R5821	T9662	T9660	pobj	genes,for
R5822	T9663	T9664	amod	olfactory,receptor
R5823	T9664	T9662	compound	receptor,genes
R5824	T9665	T9639	punct	.,provides
R5825	T9667	T9668	compound	Probe,design
R5826	T9668	T9669	nsubj	design,account
R5827	T9669	T9672	ccomp	account,designed
R5828	T9670	T9669	aux	must,account
R5829	T9671	T9669	advmod	also,account
R583	T1473	T1474	compound	transporter,gene
R5830	T9673	T9669	prep	for,account
R5831	T9674	T9675	det	the,isoforms
R5832	T9675	T9673	pobj	isoforms,for
R5833	T9676	T9675	amod	multiple,isoforms
R5834	T9677	T9675	amod	transcriptional,isoforms
R5835	T9678	T9675	acl	observed,isoforms
R5836	T9679	T9678	prep	for,observed
R5837	T9680	T9681	amod	many,receptors
R5838	T9681	T9679	pobj	receptors,for
R5839	T9682	T9681	amod	olfactory,receptors
R584	T1474	T1468	compound	gene,family
R5840	T9683	T9672	punct	-,designed
R5841	T9684	T9672	prep	depending,designed
R5842	T9685	T9684	prep	on,depending
R5843	T9686	T9687	det	the,question
R5844	T9687	T9685	pobj	question,on
R5845	T9688	T9689	auxpass	being,asked
R5846	T9689	T9687	acl	asked,question
R5847	T9690	T9672	punct	", ",designed
R5848	T9691	T9672	nsubjpass	probes,designed
R5849	T9692	T9672	aux	could,designed
R585	T1475	T1476	punct	[,7
R5850	T9693	T9672	auxpass	be,designed
R5851	T9694	T9672	prep	in,designed
R5852	T9695	T9696	amod	shared,sequence
R5853	T9696	T9694	pobj	sequence,in
R5854	T9697	T9698	aux	to,determine
R5855	T9698	T9672	advcl	determine,designed
R5856	T9699	T9700	det	the,level
R5857	T9700	T9698	dobj	level,determine
R5858	T9701	T9700	amod	total,level
R5859	T9702	T9700	prep	of,level
R586	T1476	T1462	parataxis	7,expressed
R5860	T9703	T9704	det	all,isoforms
R5861	T9704	T9702	pobj	isoforms,of
R5862	T9705	T9672	punct	", ",designed
R5863	T9706	T9672	cc	or,designed
R5864	T9707	T9672	conj	in,designed
R5865	T9708	T9709	amod	unique,exons
R5866	T9709	T9707	pobj	exons,in
R5867	T9710	T9711	aux	to,measure
R5868	T9711	T9707	advcl	measure,in
R5869	T9712	T9713	det	the,level
R587	T1477	T1476	punct	],7
R5870	T9713	T9711	dobj	level,measure
R5871	T9714	T9713	prep	of,level
R5872	T9715	T9716	det	each,isoform
R5873	T9716	T9714	pobj	isoform,of
R5874	T9717	T9711	advmod	separately,measure
R5875	T9718	T9672	punct	.,designed
R5876	T9720	T9721	nsubj	We,find
R5877	T9722	T9723	mark	that,transcribed
R5878	T9723	T9721	ccomp	transcribed,find
R5879	T9724	T9725	det	the,majority
R588	T1478	T1430	punct	),be
R5880	T9725	T9723	nsubjpass	majority,transcribed
R5881	T9726	T9725	prep	of,majority
R5882	T9727	T9728	det	the,receptors
R5883	T9728	T9726	pobj	receptors,of
R5884	T9729	T9728	amod	olfactory,receptors
R5885	T9730	T9723	punct	", ",transcribed
R5886	T9731	T9723	prep	like,transcribed
R5887	T9732	T9733	amod	most,genes
R5888	T9733	T9731	pobj	genes,like
R5889	T9734	T9733	amod	non-olfactory,genes
R589	T1479	T1430	punct	.,be
R5890	T9735	T9733	compound	receptor,genes
R5891	T9842	T9841	punct	", ",stability
R5892	T9843	T9841	conj	localization,stability
R5893	T9844	T9843	cc	or,localization
R5894	T9845	T9843	conj	degradation,localization
R5895	T9846	T9847	punct	[,42
R5896	T9847	T9823	parataxis	42,have
R5897	T9848	T9847	nummod	41,42
R5898	T9849	T9847	punct	",",42
R5899	T9850	T9847	punct	],42
R59	T361	T360	amod	odorant,cDNAs
R590	T1481	T1482	det	A,subset
R5900	T9851	T9823	punct	.,have
R5901	T9853	T9854	poss	Our,study
R5902	T9854	T9855	nsubj	study,shows
R5903	T9856	T9857	mark	that,fit
R5904	T9857	T9855	ccomp	fit,shows
R5905	T9858	T9859	quantmod	about,5
R5906	T9859	T9860	nummod	5,%
R5907	T9860	T9857	nsubj	%,fit
R5908	T9736	T9737	punct	[,39
R5909	T9861	T9860	prep	of,%
R591	T1482	T1484	nsubj	subset,appears
R5910	T9737	T9723	parataxis	39,transcribed
R5911	T9862	T9863	amod	olfactory,receptor
R5912	T9738	T9737	nummod	38,39
R5913	T9739	T9737	punct	",",39
R5914	T9863	T9864	compound	receptor,transcripts
R5915	T9740	T9737	punct	],39
R5916	T9741	T9723	punct	", ",transcribed
R5917	T9864	T9861	pobj	transcripts,of
R5918	T9742	T9723	auxpass	are,transcribed
R5919	T9743	T9723	prep	as,transcribed
R592	T1483	T1482	amod	small,subset
R5920	T9744	T9745	amod	multiple,isoforms
R5921	T9745	T9743	pobj	isoforms,as
R5922	T9865	T9857	aux	do,fit
R5923	T9746	T9723	punct	", ",transcribed
R5924	T9747	T9723	advcl	involving,transcribed
R5925	T9866	T9857	neg	not,fit
R5926	T9748	T9749	amod	alternative,splicing
R5927	T9867	T9868	det	the,notion
R5928	T9868	T9857	dobj	notion,fit
R5929	T9749	T9747	dobj	splicing,involving
R593	T1485	T1482	prep	of,subset
R5930	T9750	T9749	prep	of,splicing
R5931	T9869	T9868	amod	current,notion
R5932	T9751	T9752	nummod	5,exons
R5933	T9752	T9750	pobj	exons,of
R5934	T9753	T9751	punct	',5
R5935	T9870	T9868	prep	of,notion
R5936	T9754	T9752	amod	untranslated,exons
R5937	T9755	T9749	cc	and,splicing
R5938	T9756	T9757	amod	alternate,usage
R5939	T9757	T9749	conj	usage,splicing
R594	T1486	T1487	amod	olfactory,receptors
R5940	T9758	T9759	compound	polyadenylation,site
R5941	T9871	T9872	amod	olfactory,receptor
R5942	T9759	T9757	compound	site,usage
R5943	T9760	T9759	punct	-,site
R5944	T9761	T9721	punct	.,find
R5945	T9872	T9873	compound	receptor,gene
R5946	T9763	T9764	det	The,act
R5947	T9764	T9765	nsubj	act,be
R5948	T9873	T9874	compound	gene,structure
R5949	T9766	T9764	prep	of,act
R595	T1487	T1485	pobj	receptors,of
R5950	T9874	T9870	pobj	structure,of
R5951	T9767	T9766	pobj	splicing,of
R5952	T9875	T9855	punct	.,shows
R5953	T9768	T9764	appos	itself,act
R5954	T9769	T9765	aux	may,be
R5955	T9770	T9765	acomp	important,be
R5956	T9877	T9878	advmod	Occasionally,spliced
R5957	T9771	T9772	prep	for,40
R5958	T9772	T9765	parataxis	40,be
R5959	T9773	T9774	amod	efficient,export
R596	T1488	T1489	aux	to,expressed
R5960	T9774	T9771	pobj	export,for
R5961	T9879	T9878	punct	", ",spliced
R5962	T9775	T9774	compound	mRNA,export
R5963	T9776	T9774	prep	from,export
R5964	T9777	T9778	det	the,nucleus
R5965	T9880	T9881	det	an,intron
R5966	T9778	T9776	pobj	nucleus,from
R5967	T9779	T9772	punct	[,40
R5968	T9780	T9772	punct	],40
R5969	T9881	T9878	nsubjpass	intron,spliced
R597	T1489	T1484	xcomp	expressed,appears
R5970	T9781	T9772	cc	or,40
R5971	T9782	T9783	aux	to,couple
R5972	T9882	T9878	auxpass	is,spliced
R5973	T9783	T9772	conj	couple,40
R5974	T9784	T9785	amod	olfactory,receptor
R5975	T9785	T9786	compound	receptor,regions
R5976	T9883	T9878	prep	out,spliced
R5977	T9786	T9783	dobj	regions,couple
R5978	T9787	T9786	compound	coding,regions
R5979	T9884	T9883	prep	of,out
R598	T1490	T1489	auxpass	be,expressed
R5980	T9788	T9783	prep	with,couple
R5981	T9789	T9790	advmod	genomically,distant
R5982	T9790	T9791	amod	distant,promoters
R5983	T9885	T9886	det	the,region
R5984	T9791	T9788	pobj	promoters,with
R5985	T9792	T9765	punct	.,be
R5986	T9886	T9884	pobj	region,of
R5987	T9794	T9795	det	The,nature
R5988	T9795	T9797	nsubj	nature,be
R5989	T9887	T9886	nummod	3,region
R599	T1491	T1489	prep	in,expressed
R5990	T9796	T9795	amod	exact,nature
R5991	T9798	T9795	prep	of,nature
R5992	T9799	T9800	det	the,transcript
R5993	T9800	T9798	pobj	transcript,of
R5994	T9888	T9887	punct	',3
R5995	T9801	T9800	amod	spliced,transcript
R5996	T9802	T9797	aux	might,be
R5997	T9803	T9797	acomp	unimportant,be
R5998	T9889	T9886	amod	untranslated,region
R5999	T9804	T9797	punct	", ",be
R6	T302	T300	acl	confirming,evidence
R60	T362	T360	compound	receptor,cDNAs
R600	T1492	T1493	amod	non-olfactory,tissues
R6000	T9805	T9806	amod	such,produced
R6001	T9890	T9878	punct	.,spliced
R6002	T9806	T9797	advcl	produced,be
R6003	T9807	T9806	mark	that,produced
R6004	T9808	T9809	amod	several,isoforms
R6005	T9892	T9893	det	A,number
R6006	T9809	T9806	nsubjpass	isoforms,produced
R6007	T9810	T9806	aux	might,produced
R6008	T9811	T9806	auxpass	be,produced
R6009	T9893	T9894	nsubj	number,use
R601	T1493	T1491	pobj	tissues,in
R6010	T9812	T9813	advmod	simply,are
R6011	T9813	T9806	advcl	are,produced
R6012	T9814	T9813	mark	because,are
R6013	T9815	T9816	amod	multiple,sites
R6014	T9895	T9893	prep	of,number
R6015	T9816	T9813	nsubj	sites,are
R6016	T9817	T9816	amod	functional,sites
R6017	T9818	T9816	compound	splice,sites
R6018	T9819	T9813	acomp	available,are
R6019	T9896	T9895	pobj	cDNAs,of
R602	T1494	T1493	punct	", ",tissues
R6020	T9820	T9797	punct	.,be
R6021	T9897	T9898	compound	splice,sites
R6022	T9822	T9823	advmod	Alternatively,have
R6023	T9824	T9823	punct	", ",have
R6024	T9898	T9894	dobj	sites,use
R6025	T9825	T9826	det	the,multiplicity
R6026	T9826	T9823	nsubj	multiplicity,have
R6027	T9827	T9826	prep	of,multiplicity
R6028	T9899	T9898	prep	within,sites
R6029	T9828	T9829	amod	transcriptional,isoforms
R603	T1495	T1496	advmod	principally,testis
R6030	T9829	T9827	pobj	isoforms,of
R6031	T9830	T9823	aux	might,have
R6032	T9900	T9901	det	the,receptor
R6033	T9831	T9832	amod	functional,significance
R6034	T9832	T9823	dobj	significance,have
R6035	T9901	T9903	poss	receptor,ORF
R6036	T9833	T9823	punct	", ",have
R6037	T9834	T9835	mark	as,contain
R6038	T9902	T9901	amod	olfactory,receptor
R6039	T9835	T9823	advcl	contain,have
R604	T1496	T1493	appos	testis,tissues
R6040	T9903	T9899	pobj	ORF,within
R6041	T9836	T9835	nsubj	UTRs,contain
R6042	T9837	T9835	aux	may,contain
R6043	T9838	T9835	dobj	signals,contain
R6044	T9839	T9838	acl	controlling,signals
R6045	T9840	T9841	compound	mRNA,stability
R6046	T9904	T9901	case	's,receptor
R6047	T9841	T9839	dobj	stability,controlling
R6048	T9905	T9894	punct	", ",use
R6049	T9906	T9894	advcl	meaning,use
R605	T1497	T1496	det	the,testis
R6050	T9907	T9908	mark	that,is
R6051	T9908	T9906	ccomp	is,meaning
R6052	T9909	T9910	poss	their,product
R6053	T9910	T9908	nsubj	product,is
R6054	T9948	T9946	compound	amino,acids
R6055	T9949	T9940	agent	by,encoded
R6056	T9911	T9910	compound	protein,product
R6057	T9950	T9951	det	an,exon
R6058	T9951	T9949	pobj	exon,by
R6059	T9912	T9908	acomp	different,is
R606	T1498	T1499	punct	[,8
R6060	T9952	T9951	amod	upstream,exon
R6061	T9953	T9926	punct	", ",encode
R6062	T9954	T9955	mark	as,observed
R6063	T9913	T9912	prep	from,different
R6064	T9955	T9926	advcl	observed,encode
R6065	T9956	T9955	aux	has,observed
R6066	T9914	T9913	pobj	that,from
R6067	T9957	T9955	auxpass	been,observed
R6068	T9958	T9955	advmod	previously,observed
R6069	T9915	T9914	acl	predicted,that
R607	T1499	T1496	parataxis	8,testis
R6070	T9959	T9955	prep	for,observed
R6071	T9960	T9961	det	a,gene
R6072	T9961	T9959	pobj	gene,for
R6073	T9916	T9915	prep	on,predicted
R6074	T9962	T9961	amod	subtelomeric,gene
R6075	T9963	T9961	amod	human,gene
R6076	T9964	T9965	amod	olfactory,receptor
R6077	T9917	T9918	det	the,basis
R6078	T9965	T9961	compound	receptor,gene
R6079	T9966	T9967	punct	[,25
R608	T1500	T1499	punct	],8
R6080	T9918	T9916	pobj	basis,on
R6081	T9967	T9926	parataxis	25,encode
R6082	T9968	T9967	punct	],25
R6083	T9969	T9926	punct	.,encode
R6084	T9919	T9918	prep	of,basis
R6085	T9971	T9972	amod	Such,splicing
R6086	T9972	T9976	nsubj	splicing,increase
R6087	T9920	T9921	amod	genomic,sequence
R6088	T9973	T9972	nmod	within,splicing
R6089	T9974	T9973	punct	-,within
R609	T1501	T1496	punct	", ",testis
R6090	T9975	T9973	pobj	ORF,within
R6091	T9921	T9919	pobj	sequence,of
R6092	T9977	T9976	aux	might,increase
R6093	T9978	T9979	npadvmod	protein,coding
R6094	T9979	T9981	amod	coding,diversity
R6095	T9980	T9979	punct	-,coding
R6096	T9981	T9976	dobj	diversity,increase
R6097	T9982	T9976	punct	", ",increase
R6098	T9922	T9921	advmod	alone,sequence
R6099	T9983	T9984	mark	although,is
R61	T363	T360	acl	representing,cDNAs
R610	T1502	T1496	cc	but,testis
R6100	T9984	T9976	advcl	is,increase
R6101	T9985	T9984	punct	", ",is
R6102	T9923	T9894	punct	.,use
R6103	T9986	T9984	prep	given,is
R6104	T9987	T9988	det	the,number
R6105	T9988	T9986	pobj	number,given
R6106	T9925	T9926	prep	In,encode
R6107	T9989	T9988	amod	small,number
R6108	T9990	T9988	prep	of,number
R6109	T9991	T9990	pobj	genes,of
R611	T1503	T1502	advmod	also,but
R6110	T9992	T9988	acl	involved,number
R6111	T9993	T9984	punct	", ",is
R6112	T9927	T9928	nummod	two,cases
R6113	T9994	T9984	nsubj	splicing,is
R6114	T9995	T9984	acomp	unlikely,is
R6115	T9996	T9997	aux	to,affect
R6116	T9997	T9995	xcomp	affect,unlikely
R6117	T9928	T9925	pobj	cases,In
R6118	T9998	T9997	advmod	significantly,affect
R6119	T9999	T10000	det	the,repertoire
R612	T1504	T1505	compound	taste,tissues
R6120	T10000	T9997	dobj	repertoire,affect
R6121	T10001	T10000	amod	functional,repertoire
R6122	T9929	T9928	amod	such,cases
R6123	T10002	T10000	compound	receptor,repertoire
R6124	T10003	T9976	punct	.,increase
R6125	T9930	T9926	punct	", ",encode
R6126	T10005	T10006	nsubj	Most,appear
R6127	T10007	T10005	prep	of,Most
R6128	T9931	T9932	det	the,transcript
R6129	T10008	T10009	det	the,forms
R613	T1505	T1496	conj	tissues,testis
R6130	T9932	T9926	nsubj	transcript,encode
R6131	T9933	T9926	aux	would,encode
R6132	T10009	T10007	pobj	forms,of
R6133	T9934	T9935	det	a,receptor
R6134	T10010	T10009	amod	atypical,forms
R6135	T10011	T10009	compound	splice,forms
R6136	T10012	T10013	nsubj	we,observe
R6137	T9935	T9926	dobj	receptor,encode
R6138	T10013	T10009	advcl	observe,forms
R6139	T10014	T10015	aux	to,encode
R614	T1506	T1507	punct	[,9
R6140	T10015	T10006	xcomp	encode,appear
R6141	T9936	T9935	amod	functional,receptor
R6142	T10016	T10017	amod	non-functional,transcripts
R6143	T10017	T10015	dobj	transcripts,encode
R6144	T9937	T9935	amod	olfactory,receptor
R6145	T10018	T10017	punct	", ",transcripts
R6146	T10019	T10017	acl	containing,transcripts
R6147	T10020	T10019	dobj	frameshifts,containing
R6148	T10021	T10019	cc	or,containing
R6149	T10022	T10019	conj	lacking,containing
R615	T1507	T1505	parataxis	9,tissues
R6150	T9938	T9926	punct	", ",encode
R6151	T10023	T10024	det	a,codon
R6152	T10024	T10022	dobj	codon,lacking
R6153	T10025	T10024	compound	start,codon
R6154	T9939	T9940	mark	with,encoded
R6155	T10026	T10024	cc	or,codon
R6156	T10027	T10028	amod	other,residues
R6157	T10028	T10024	conj	residues,codon
R6158	T9940	T9926	advcl	encoded,encode
R6159	T10029	T10028	amod	functional,residues
R616	T1508	T1507	punct	],9
R6160	T10030	T10028	acl	conserved,residues
R6161	T10031	T10030	prep	throughout,conserved
R6162	T9941	T9942	det	the,methionine
R6163	T10032	T10033	det	the,family
R6164	T10033	T10031	pobj	family,throughout
R6165	T10034	T10035	amod	olfactory,receptor
R6166	T9942	T9940	nsubj	methionine,encoded
R6167	T10035	T10033	compound	receptor,family
R6168	T10036	T10006	punct	.,appear
R6169	T9943	T9942	amod	initiating,methionine
R617	T1509	T1505	punct	", ",tissues
R6170	T10038	T10039	det	These,transcripts
R6171	T10039	T10041	nsubj	transcripts,are
R6172	T9944	T9942	cc	and,methionine
R6173	T10040	T10039	amod	nonfunctional,transcripts
R6174	T10042	T10041	advmod	probably,are
R6175	T9945	T9946	amod	first,acids
R6176	T10043	T10044	amod	aberrant,products
R6177	T10044	T10041	attr	products,are
R6178	T10045	T10044	compound	by,products
R6179	T9946	T9942	conj	acids,methionine
R618	T1510	T1505	conj	prostate,tissues
R6180	T10046	T10044	punct	-,products
R6181	T10047	T10044	prep	of,products
R6182	T10048	T10049	det	the,system
R6183	T9947	T9946	amod	few,acids
R6184	T10049	T10047	pobj	system,of
R6185	T10050	T10049	compound	splicing,system
R6186	T10051	T10052	punct	[,43
R6187	T10054	T10055	dep	that,degraded
R6188	T10052	T10044	parataxis	43,products
R6189	T10053	T10052	punct	],43
R619	T1511	T1512	punct	[,10
R6190	T10055	T10044	relcl	degraded,products
R6191	T10056	T10055	aux	have,degraded
R6192	T10057	T10055	neg	not,degraded
R6193	T10058	T10055	advmod	yet,degraded
R6194	T10160	T10158	pobj	pseudogenes,for
R6195	T10161	T10160	amod	human,pseudogenes
R6196	T10059	T10055	auxpass	been,degraded
R6197	T10162	T10163	amod	olfactory,receptor
R6198	T10163	T10160	compound	receptor,pseudogenes
R6199	T10164	T10165	punct	[,44
R62	T364	T365	amod	more,400
R620	T1512	T1510	parataxis	10,prostate
R6200	T10165	T10143	parataxis	44,observe
R6201	T10166	T10165	nummod	26,44
R6202	T10060	T10055	agent	by,degraded
R6203	T10167	T10165	punct	",",44
R6204	T10168	T10165	punct	],44
R6205	T10169	T10143	punct	.,observe
R6206	T10061	T10062	compound	RNA,systems
R6207	T10171	T10172	mark	Although,represented
R6208	T10172	T10180	advcl	represented,express
R6209	T10062	T10060	pobj	systems,by
R621	T1513	T1512	punct	],10
R6210	T10173	T10174	amod	many,fewer
R6211	T10174	T10175	amod	fewer,pseudogenes
R6212	T10175	T10172	nsubjpass	pseudogenes,represented
R6213	T10063	T10062	compound	surveillance,systems
R6214	T10176	T10175	prep	than,pseudogenes
R6215	T10177	T10178	amod	intact,genes
R6216	T10064	T10065	punct	[,41
R6217	T10178	T10176	pobj	genes,than
R6218	T10179	T10172	auxpass	were,represented
R6219	T10065	T10055	parataxis	41,degraded
R622	T1514	T1510	punct	", ",prostate
R6220	T10181	T10172	prep	in,represented
R6221	T10066	T10065	nummod	40,41
R6222	T10182	T10183	poss	our,collection
R6223	T10067	T10065	punct	",",41
R6224	T10183	T10181	pobj	collection,in
R6225	T10184	T10183	compound	cDNA,collection
R6226	T10068	T10065	punct	],41
R6227	T10185	T10180	punct	", ",express
R6228	T10186	T10187	det	some,neurons
R6229	T10187	T10180	nsubj	neurons,express
R623	T1515	T1516	amod	erythroid,cells
R6230	T10069	T10041	punct	.,are
R6231	T10188	T10187	prep	in,neurons
R6232	T10189	T10190	det	the,epithelium
R6233	T10071	T10072	det	The,neurons
R6234	T10190	T10188	pobj	epithelium,in
R6235	T10191	T10190	amod	olfactory,epithelium
R6236	T10192	T10180	advmod	evidently,express
R6237	T10193	T10194	amod	disrupted,receptors
R6238	T10072	T10073	nsubj	neurons,make
R6239	T10194	T10180	dobj	receptors,express
R624	T1516	T1510	conj	cells,prostate
R6240	T10195	T10194	amod	olfactory,receptors
R6241	T10196	T10180	cc	and,express
R6242	T10074	T10072	acl	expressing,neurons
R6243	T10197	T10198	advmod	thus,be
R6244	T10198	T10180	conj	be,express
R6245	T10199	T10198	aux	might,be
R6246	T10200	T10198	acomp	unable,be
R6247	T10075	T10076	det	these,transcripts
R6248	T10201	T10202	aux	to,respond
R6249	T10202	T10200	xcomp	respond,unable
R625	T1517	T1518	punct	[,11
R6250	T10076	T10074	dobj	transcripts,expressing
R6251	T10203	T10202	prep	to,respond
R6252	T10204	T10203	pobj	odorants,to
R6253	T10205	T10202	cc	or,respond
R6254	T10206	T10207	aux	to,innervate
R6255	T10207	T10202	conj	innervate,respond
R6256	T10077	T10076	amod	aberrant,transcripts
R6257	T10208	T10207	advmod	correctly,innervate
R6258	T10209	T10210	det	the,bulb
R6259	T10210	T10207	dobj	bulb,innervate
R626	T1518	T1516	parataxis	11,cells
R6260	T10078	T10073	aux	might,make
R6261	T10211	T10210	amod	olfactory,bulb
R6262	T10212	T10180	punct	.,express
R6263	T10079	T10073	advmod	also,make
R6264	T10214	T10215	nsubj	Wang,shown
R6265	T10080	T10081	amod	normal,transcripts
R6266	T10216	T10214	punct	", ",Wang
R6267	T10217	T10214	conj	Axel,Wang
R6268	T10218	T10217	cc	and,Axel
R6269	T10081	T10073	dobj	transcripts,make
R627	T1519	T1518	punct	],11
R6270	T10219	T10217	conj	coworkers,Axel
R6271	T10220	T10215	aux	have,shown
R6272	T10221	T10222	mark	that,support
R6273	T10082	T10073	prep	for,make
R6274	T10222	T10215	advcl	support,shown
R6275	T10223	T10224	det	an,gene
R6276	T10083	T10084	det	the,genes
R6277	T10224	T10222	nsubj	gene,support
R6278	T10225	T10224	amod	artificial,gene
R6279	T10226	T10224	amod	transgenic,gene
R628	T1520	T1516	punct	", ",cells
R6280	T10084	T10082	pobj	genes,for
R6281	T10227	T10228	amod	olfactory,receptor
R6282	T10228	T10224	compound	receptor,gene
R6283	T10229	T10224	acl	containing,gene
R6284	T10085	T10084	amod	same,genes
R6285	T10230	T10231	nummod	two,mutations
R6286	T10231	T10229	dobj	mutations,containing
R6287	T10086	T10073	cc	and,make
R6288	T10232	T10231	amod	nonsense,mutations
R6289	T10233	T10222	aux	can,support
R629	T1521	T1516	conj	notochord,cells
R6290	T10234	T10222	dobj	development,support
R6291	T10087	T10088	advmod	thus,produce
R6292	T10235	T10234	prep	of,development
R6293	T10236	T10237	det	an,neuron
R6294	T10237	T10235	pobj	neuron,of
R6295	T10088	T10073	conj	produce,make
R6296	T10238	T10237	amod	olfactory,neuron
R6297	T10239	T10222	punct	", ",support
R6298	T10240	T10222	cc	but,support
R6299	T10089	T10090	det	a,receptor
R63	T365	T367	nummod	400,genes
R630	T1522	T1523	punct	[,12
R6300	T10241	T10242	mark	that,fail
R6301	T10242	T10222	conj	fail,support
R6302	T10243	T10244	npadvmod	pseudogene,expressing
R6303	T10244	T10246	amod	expressing,neurons
R6304	T10245	T10244	punct	-,expressing
R6305	T10246	T10242	nsubj	neurons,fail
R6306	T10090	T10088	dobj	receptor,produce
R6307	T10247	T10248	aux	to,converge
R6308	T10248	T10242	xcomp	converge,fail
R6309	T10249	T10248	prep	on,converge
R631	T1523	T1521	parataxis	12,notochord
R6310	T10091	T10090	amod	functional,receptor
R6311	T10250	T10251	det	a,glomerulus
R6312	T10251	T10249	pobj	glomerulus,on
R6313	T10252	T10248	prep	in,converge
R6314	T10092	T10090	amod	olfactory,receptor
R6315	T10253	T10254	det	the,bulb
R6316	T10254	T10252	pobj	bulb,in
R6317	T10093	T10073	punct	.,make
R6318	T10255	T10254	amod	olfactory,bulb
R6319	T10256	T10257	punct	[,45
R632	T1524	T1523	punct	],12
R6320	T10095	T10096	advmod	Alternatively,ensure
R6321	T10257	T10215	parataxis	45,shown
R6322	T10258	T10257	punct	],45
R6323	T10259	T10215	punct	.,shown
R6324	T10097	T10096	punct	", ",ensure
R6325	T10261	T10262	prep	By,is
R6326	T10263	T10261	pobj	analogy,By
R6327	T10264	T10263	prep	with,analogy
R6328	T10098	T10099	det	the,regulation
R6329	T10265	T10266	det	an,mutant
R633	T1525	T1521	cc	and,notochord
R6330	T10099	T10096	nsubj	regulation,ensure
R6331	T10100	T10099	amod	unusual,regulation
R6332	T10101	T10099	amod	transcriptional,regulation
R6333	T10102	T10099	prep	of,regulation
R6334	T10103	T10104	amod	olfactory,receptors
R6335	T10104	T10102	pobj	receptors,of
R6336	T10266	T10264	pobj	mutant,with
R6337	T10105	T10096	aux	might,ensure
R6338	T10267	T10268	amod	olfactory,receptor
R6339	T10268	T10269	compound	receptor,deletion
R634	T1526	T1527	advmod	perhaps,tissues
R6340	T10106	T10107	mark	that,expressed
R6341	T10269	T10266	compound	deletion,mutant
R6342	T10270	T10271	punct	[,45
R6343	T10107	T10096	ccomp	expressed,ensure
R6344	T10271	T10263	parataxis	45,analogy
R6345	T10272	T10271	punct	],45
R6346	T10273	T10262	punct	", ",is
R6347	T10108	T10109	advmod	only,isoform
R6348	T10274	T10262	nsubj	it,is
R6349	T10275	T10262	acomp	likely,is
R635	T1527	T1521	conj	tissues,notochord
R6350	T10109	T10107	nsubjpass	isoform,expressed
R6351	T10276	T10277	mark	that,die
R6352	T10277	T10262	ccomp	die,is
R6353	T10278	T10279	amod	most,neurons
R6354	T10110	T10109	nummod	one,isoform
R6355	T10279	T10277	nsubj	neurons,die
R6356	T10280	T10281	npadvmod	pseudogene,expressing
R6357	T10281	T10279	amod	expressing,neurons
R6358	T10282	T10281	punct	-,expressing
R6359	T10283	T10277	cc	or,die
R636	T1528	T1527	amod	other,tissues
R6360	T10111	T10109	compound	splice,isoform
R6361	T10284	T10277	conj	switch,die
R6362	T10285	T10286	aux	to,express
R6363	T10286	T10284	advcl	express,switch
R6364	T10287	T10288	det	a,gene
R6365	T10112	T10107	auxpass	is,expressed
R6366	T10288	T10286	dobj	gene,express
R6367	T10289	T10288	amod	different,gene
R6368	T10113	T10107	prep	per,expressed
R6369	T10290	T10291	amod	olfactory,receptor
R637	T1529	T1484	punct	.,appears
R6370	T10291	T10288	compound	receptor,gene
R6371	T10292	T10277	punct	", ",die
R6372	T10293	T10277	advcl	leaving,die
R6373	T10114	T10113	pobj	cell,per
R6374	T10294	T10295	det	a,number
R6375	T10295	T10293	dobj	number,leaving
R6376	T10115	T10116	punct	(,unlikely
R6377	T10296	T10295	amod	small,number
R6378	T10297	T10295	prep	of,number
R6379	T10298	T10299	npadvmod	pseudogene,expressing
R638	T1531	T1532	nsubj	Expression,led
R6380	T10116	T10107	parataxis	unlikely,expressed
R6381	T10299	T10301	amod	expressing,neurons
R6382	T10300	T10299	punct	-,expressing
R6383	T10117	T10116	punct	", ",unlikely
R6384	T10118	T10116	cc	but,unlikely
R6385	T10119	T10116	conj	possible,unlikely
R6386	T10301	T10297	pobj	neurons,of
R6387	T10120	T10121	mark	if,operates
R6388	T10302	T10293	prep	in,leaving
R6389	T10303	T10304	amod	adult,mice
R639	T1533	T1531	prep	in,Expression
R6390	T10121	T10116	advcl	operates,unlikely
R6391	T10304	T10302	pobj	mice,in
R6392	T10305	T10293	punct	", ",leaving
R6393	T10122	T10123	det	an,mechanism
R6394	T10306	T10307	cc	but,at
R6395	T10307	T10293	prep	at,leaving
R6396	T10308	T10309	advmod	greatly,reduced
R6397	T10123	T10121	nsubj	mechanism,operates
R6398	T10309	T10310	amod	reduced,levels
R6399	T10310	T10307	pobj	levels,at
R64	T366	T365	quantmod	than,400
R640	T1534	T1535	det	the,testis
R6400	T10311	T10310	prep	compared,levels
R6401	T10124	T10125	npadvmod	RNA,based
R6402	T10312	T10311	prep	to,compared
R6403	T10313	T10312	pobj	neurons,to
R6404	T10314	T10313	acl	expressing,neurons
R6405	T10125	T10123	amod	based,mechanism
R6406	T10315	T10316	amod	intact,receptors
R6407	T10316	T10314	dobj	receptors,expressing
R6408	T10317	T10316	amod	olfactory,receptors
R6409	T10318	T10262	punct	.,is
R641	T1535	T1533	pobj	testis,in
R6410	T10126	T10125	punct	-,based
R6411	T10127	T10123	compound	feedback,mechanism
R6412	T10128	T10116	punct	),unlikely
R6413	T10129	T10107	punct	", ",expressed
R6414	T10130	T10131	advmod	thus,condemning
R6415	T10131	T10107	advcl	condemning,expressed
R6416	T10132	T10133	nsubj	cells,be
R6417	T10133	T10131	ccomp	be,condemning
R6418	T10134	T10132	acl	expressing,cells
R6419	T10135	T10136	det	these,isoforms
R642	T1536	T1532	aux	has,led
R6420	T10136	T10134	dobj	isoforms,expressing
R6421	T10137	T10136	amod	aberrant,isoforms
R6422	T10138	T10133	aux	to,be
R6423	T10139	T10133	acomp	dysfunctional,be
R6424	T10140	T10096	punct	.,ensure
R6425	T10142	T10143	nsubj	We,observe
R6426	T10144	T10143	advmod	also,observe
R6427	T10145	T10143	dobj	transcripts,observe
R6428	T10146	T10145	prep	from,transcripts
R6429	T10147	T10148	det	a,number
R643	T1537	T1538	det	some,investigators
R6430	T10148	T10146	pobj	number,from
R6431	T10149	T10148	amod	small,number
R6432	T10150	T10148	prep	of,number
R6433	T10151	T10152	amod	olfactory,receptor
R6434	T10152	T10153	compound	receptor,pseudogenes
R6435	T10153	T10150	pobj	pseudogenes,of
R6436	T10154	T10143	punct	", ",observe
R6437	T10155	T10156	mark	as,described
R6438	T10156	T10143	advcl	described,observe
R6439	T10157	T10156	advmod	previously,described
R644	T1538	T1532	dobj	investigators,led
R6440	T10158	T10156	prep	for,described
R6441	T10159	T10160	nummod	three,pseudogenes
R6442	T10408	T10409	poss	Our,study
R6443	T10409	T10410	nsubj	study,provided
R6444	T10411	T10410	aux	has,provided
R6445	T10412	T10413	det	an,resource
R6446	T10413	T10410	dobj	resource,provided
R6447	T10414	T10415	amod	olfactory,receptor
R6448	T10415	T10416	compound	receptor,cDNA
R6449	T10416	T10413	compound	cDNA,resource
R645	T1539	T1540	aux	to,suggest
R6450	T10417	T10413	acl	representing,resource
R6451	T10418	T10419	quantmod	over,one
R6452	T10419	T10417	dobj	one,representing
R6453	T10420	T10419	punct	-,one
R6454	T10421	T10419	amod	third,one
R6455	T10422	T10419	prep	of,one
R6456	T10423	T10424	det	the,family
R6457	T10424	T10422	pobj	family,of
R6458	T10425	T10426	amod	olfactory,receptor
R6459	T10426	T10427	compound	receptor,gene
R646	T1540	T1532	xcomp	suggest,led
R6460	T10427	T10424	compound	gene,family
R6461	T10428	T10410	punct	.,provided
R6462	T10430	T10431	nsubj	We,established
R6463	T10432	T10431	aux	have,established
R6464	T10433	T10431	advmod	thus,established
R6465	T10434	T10435	quantmod	over,400
R6466	T10435	T10436	nummod	400,genes
R6467	T10436	T10431	dobj	genes,established
R6468	T10437	T10436	amod	annotated,genes
R6469	T10438	T10439	amod	olfactory,receptor
R647	T1541	T1542	mark	that,function
R6470	T10439	T10436	compound	receptor,genes
R6471	T10440	T10431	prep	as,established
R6472	T10441	T10440	pcomp	having,as
R6473	T10442	T10443	amod	olfactory,function
R6474	T10443	T10441	dobj	function,having
R6475	T10444	T10431	punct	.,established
R6476	T10446	T10447	det	The,sequences
R6477	T10447	T10448	nsubj	sequences,demonstrate
R6478	T10449	T10450	nsubj	we,generated
R6479	T10450	T10447	advcl	generated,sequences
R648	T1542	T1540	ccomp	function,suggest
R6480	T10451	T10452	mark	that,has
R6481	T10452	T10448	ccomp	has,demonstrate
R6482	T10453	T10454	det	the,majority
R6483	T10454	T10452	nsubj	majority,has
R6484	T10455	T10454	prep	of,majority
R6485	T10456	T10457	det	the,family
R6486	T10457	T10455	pobj	family,of
R6487	T10458	T10459	amod	olfactory,receptor
R6488	T10459	T10460	compound	receptor,gene
R6489	T10460	T10457	compound	gene,family
R649	T1543	T1544	det	a,subset
R6490	T10461	T10462	amod	multiple,isoforms
R6491	T10462	T10452	dobj	isoforms,has
R6492	T10463	T10462	amod	transcriptional,isoforms
R6493	T10464	T10448	punct	.,demonstrate
R6494	T10466	T10467	advmod	Most,transcripts
R6495	T10467	T10470	nsubj	transcripts,encode
R6496	T10468	T10469	amod	olfactory,receptor
R6497	T10469	T10467	compound	receptor,transcripts
R6498	T10471	T10472	amod	functional,proteins
R6499	T10472	T10470	dobj	proteins,encode
R65	T367	T363	dobj	genes,representing
R650	T1544	T1542	nsubj	subset,function
R6500	T10473	T10472	compound	receptor,proteins
R6501	T10474	T10470	punct	", ",encode
R6502	T10475	T10470	prep	with,encode
R6503	T10476	T10477	amod	rare,exceptions
R6504	T10477	T10475	pobj	exceptions,with
R6505	T10478	T10470	punct	.,encode
R6506	T10480	T10481	nsubj	We,show
R6507	T10482	T10483	mark	that,have
R6508	T10483	T10481	ccomp	have,show
R6509	T10484	T10485	amod	individual,genes
R651	T1545	T1544	prep	of,subset
R6510	T10485	T10483	nsubj	genes,have
R6511	T10486	T10487	amod	olfactory,receptor
R6512	T10487	T10485	compound	receptor,genes
R6513	T10488	T10483	aux	can,have
R6514	T10489	T10490	advmod	vastly,different
R6515	T10490	T10491	amod	different,levels
R6516	T10491	T10483	dobj	levels,have
R6517	T10492	T10491	compound	expression,levels
R6518	T10493	T10481	punct	", ",show
R6519	T10494	T10495	det	an,finding
R652	T1546	T1547	amod	olfactory,receptors
R6520	T10495	T10481	npadvmod	finding,show
R6521	T10496	T10495	amod	intriguing,finding
R6522	T10497	T10495	prep	in,finding
R6523	T10498	T10497	pobj	light,in
R6524	T10499	T10498	prep	of,light
R6525	T10500	T10501	det	the,regime
R6526	T10501	T10499	pobj	regime,of
R6527	T10502	T10501	amod	unusual,regime
R6528	T10503	T10504	nummod	one,neuron
R6529	T10504	T10501	nmod	neuron,regime
R653	T1547	T1545	pobj	receptors,of
R6530	T10505	T10504	punct	-,neuron
R6531	T10506	T10507	nummod	one,gene
R6532	T10507	T10504	appos	gene,neuron
R6533	T10508	T10507	punct	-,gene
R6534	T10509	T10501	amod	transcriptional,regime
R6535	T10510	T10501	prep	of,regime
R6536	T10511	T10512	det	the,epithelium
R6537	T10512	T10510	pobj	epithelium,of
R6538	T10513	T10512	amod	olfactory,epithelium
R6539	T10514	T10481	punct	.,show
R654	T1548	T1542	aux	may,function
R6540	T10516	T10517	poss	Our,results
R6541	T10517	T10518	nsubj	results,facilitate
R6542	T10519	T10517	cc	and,results
R6543	T10520	T10521	det	the,sequences
R6544	T10521	T10517	conj	sequences,results
R6545	T10522	T10523	nsubj	we,provide
R6546	T10523	T10521	advcl	provide,sequences
R6547	T10524	T10518	aux	will,facilitate
R6548	T10525	T10526	amod	future,studies
R6549	T10526	T10518	dobj	studies,facilitate
R655	T1549	T1542	prep	as,function
R6550	T10527	T10526	amod	global,studies
R6551	T10528	T10526	prep	of,studies
R6552	T10529	T10530	det	the,mechanisms
R6553	T10530	T10528	pobj	mechanisms,of
R6554	T10531	T10530	cc	and,mechanisms
R6555	T10532	T10530	conj	dynamics,mechanisms
R6556	T10533	T10530	prep	of,mechanisms
R6557	T10534	T10535	amod	olfactory,receptor
R6558	T10535	T10536	compound	receptor,gene
R6559	T10536	T10537	compound	gene,expression
R656	T1550	T1551	amod	spermatid,chemoreceptors
R6560	T10537	T10533	pobj	expression,of
R6561	T10538	T10518	punct	.,facilitate
R6562	T10658	T10657	prep	of,Identification
R6563	T10659	T10660	amod	olfactory,receptor
R6564	T10660	T10661	compound	receptor,cDNAs
R6565	T10661	T10658	pobj	cDNAs,of
R6566	T10663	T10664	det	An,library
R6567	T10664	T10668	nsubjpass	library,provided
R6568	T10665	T10666	amod	adult,mouse
R6569	T10666	T10667	compound	mouse,cDNA
R657	T1551	T1549	pobj	chemoreceptors,as
R6570	T10667	T10664	compound	cDNA,library
R6571	T10669	T10664	acl	made,library
R6572	T10670	T10669	prep	from,made
R6573	T10671	T10672	det	the,epithelium
R6574	T10672	T10670	pobj	epithelium,from
R6575	T10673	T10672	amod	olfactory,epithelium
R6576	T10674	T10672	prep	of,epithelium
R6577	T10675	T10676	det	a,animal
R6578	T10676	T10674	pobj	animal,of
R6579	T10677	T10676	amod	single,animal
R658	T1552	T1553	punct	[,8
R6580	T10678	T10668	auxpass	was,provided
R6581	T10679	T10668	agent	by,provided
R6582	T10680	T10681	compound	Leslie,Vosshall
R6583	T10681	T10679	pobj	Vosshall,by
R6584	T10682	T10683	punct	(,University
R6585	T10683	T10681	parataxis	University,Vosshall
R6586	T10684	T10683	compound	Rockefeller,University
R6587	T10685	T10683	punct	", ",University
R6588	T10686	T10687	compound	New,York
R6589	T10687	T10683	npadvmod	York,University
R659	T1553	T1540	parataxis	8,suggest
R6590	T10688	T10683	punct	", ",University
R6591	T10689	T10683	npadvmod	NY,University
R6592	T10690	T10683	punct	", ",University
R6593	T10691	T10683	npadvmod	USA,University
R6594	T10692	T10683	punct	),University
R6595	T10693	T10668	punct	", ",provided
R6596	T10694	T10668	cc	and,provided
R6597	T10695	T10696	det	an,library
R6598	T10696	T10698	nsubjpass	library,provided
R6599	T10697	T10696	amod	embryonic,library
R66	T368	T356	punct	.,identified
R660	T1554	T1553	punct	],8
R6600	T10698	T10668	conj	provided,provided
R6601	T10699	T10696	punct	(,library
R6602	T10700	T10696	acl	made,library
R6603	T10701	T10700	prep	from,made
R6604	T10702	T10703	det	the,epithelia
R6605	T10703	T10701	pobj	epithelia,from
R6606	T10704	T10703	amod	olfactory,epithelia
R6607	T10705	T10703	prep	of,epithelia
R6608	T10706	T10707	amod	several,embryos
R6609	T10707	T10705	pobj	embryos,of
R661	T1555	T1532	punct	.,led
R6610	T10708	T10709	compound	E16.5,E18.5
R6611	T10709	T10707	compound	E18.5,embryos
R6612	T10710	T10709	punct	-,E18.5
R6613	T10711	T10698	punct	),provided
R6614	T10712	T10698	auxpass	was,provided
R6615	T10713	T10698	agent	by,provided
R6616	T10714	T10715	compound	Tyler,Cutforth
R6617	T10715	T10713	pobj	Cutforth,by
R6618	T10716	T10717	punct	(,University
R6619	T10717	T10715	parataxis	University,Cutforth
R662	T1557	T1558	amod	Recent,studies
R6620	T10718	T10717	compound	Columbia,University
R6621	T10719	T10717	punct	", ",University
R6622	T10720	T10721	compound	New,York
R6623	T10721	T10717	npadvmod	York,University
R6624	T10722	T10717	punct	", ",University
R6625	T10723	T10717	npadvmod	NY,University
R6626	T10724	T10717	punct	", ",University
R6627	T10725	T10717	npadvmod	USA,University
R6628	T10726	T10717	punct	),University
R6629	T10727	T10698	punct	.,provided
R663	T1558	T1559	nsubj	studies,indicate
R6630	T10729	T10730	det	Both,libraries
R6631	T10730	T10731	nsubjpass	libraries,primed
R6632	T10732	T10731	auxpass	were,primed
R6633	T10733	T10731	dep	oligo,primed
R6634	T10734	T10731	punct	-,primed
R6635	T10735	T10731	dep	dT,primed
R6636	T10736	T10731	cc	and,primed
R6637	T10737	T10738	advmod	directionally,cloned
R6638	T10738	T10731	conj	cloned,primed
R6639	T10739	T10738	prep	into,cloned
R664	T1560	T1558	prep	of,studies
R6640	T10740	T10741	det	the,vector
R6641	T10741	T10739	pobj	vector,into
R6642	T10742	T10743	compound	lambdaZAP,XR
R6643	T10743	T10741	compound	XR,vector
R6644	T10744	T10743	punct	-,XR
R6645	T10745	T10746	punct	(,Stratagene
R6646	T10746	T10741	parataxis	Stratagene,vector
R6647	T10747	T10746	punct	", ",Stratagene
R6648	T10748	T10749	compound	La,Jolla
R6649	T10749	T10746	npadvmod	Jolla,Stratagene
R665	T1561	T1562	nummod	one,receptor
R6650	T10750	T10746	punct	", ",Stratagene
R6651	T10751	T10746	npadvmod	CA,Stratagene
R6652	T10752	T10746	punct	", ",Stratagene
R6653	T10753	T10746	npadvmod	USA,Stratagene
R6654	T10754	T10746	punct	),Stratagene
R6655	T10755	T10731	punct	.,primed
R6656	T10757	T10758	det	The,library
R6657	T10758	T10760	nsubj	library,has
R6658	T10759	T10758	amod	adult,library
R6659	T10761	T10762	det	a,complexity
R666	T1562	T1560	pobj	receptor,of
R6660	T10762	T10760	dobj	complexity,has
R6661	T10763	T10762	prep	of,complexity
R6662	T10764	T10765	quantmod	6.5,106
R6663	T10765	T10767	nummod	106,clones
R6664	T10766	T10765	punct	×,106
R6665	T10767	T10763	pobj	clones,of
R6666	T10768	T10767	amod	primary,clones
R6667	T10769	T10760	punct	", ",has
R6668	T10770	T10760	cc	and,has
R6669	T10771	T10772	det	the,library
R667	T1563	T1562	amod	human,receptor
R6670	T10772	T10774	nsubj	library,has
R6671	T10773	T10772	amod	embryonic,library
R6672	T10774	T10760	conj	has,has
R6673	T10775	T10776	det	a,complexity
R6674	T10776	T10774	dobj	complexity,has
R6675	T10777	T10776	prep	of,complexity
R6676	T10778	T10779	quantmod	1.65,106
R6677	T10779	T10777	pobj	106,of
R6678	T10780	T10779	punct	×,106
R6679	T10781	T10774	punct	.,has
R668	T1564	T1565	npadvmod	testis,expressed
R6680	T10783	T10784	nsubjpass	Libraries,amplified
R6681	T10785	T10784	auxpass	were,amplified
R6682	T10786	T10787	aux	to,give
R6683	T10787	T10784	advcl	give,amplified
R6684	T10788	T10787	dobj	titers,give
R6685	T10789	T10788	prep	of,titers
R6686	T10790	T10791	quantmod	5,109
R6687	T10791	T10793	nummod	109,pfu
R6688	T10792	T10791	punct	×,109
R6689	T10793	T10789	pobj	pfu,of
R669	T1565	T1562	amod	expressed,receptor
R6690	T10794	T10795	punct	/,ml
R6691	T10795	T10793	prep	ml,pfu
R6692	T10796	T10797	punct	(,adult
R6693	T10797	T10793	parataxis	adult,pfu
R6694	T10798	T10797	punct	),adult
R6695	T10799	T10793	cc	or,pfu
R6696	T10800	T10801	quantmod	2,1010
R6697	T10801	T10803	nummod	1010,pfu
R6698	T10802	T10801	punct	×,1010
R6699	T10803	T10793	conj	pfu,pfu
R67	T370	T371	advcl	Using,find
R670	T1566	T1565	punct	-,expressed
R6700	T10804	T10805	punct	/,ml
R6701	T10805	T10803	prep	ml,pfu
R6702	T10806	T10807	punct	(,embryonic
R6703	T10807	T10803	parataxis	embryonic,pfu
R6704	T10808	T10807	punct	),embryonic
R6705	T10809	T10784	punct	.,amplified
R6706	T10811	T10812	compound	Hybridization,probes
R6707	T10812	T10813	nsubjpass	probes,made
R6708	T10814	T10813	auxpass	were,made
R6709	T10815	T10813	prep	by,made
R671	T1567	T1562	amod	olfactory,receptor
R6710	T10816	T10817	amod	degenerate,PCR
R6711	T10817	T10815	pobj	PCR,by
R6712	T10818	T10817	prep	of,PCR
R6713	T10819	T10820	nmod	mouse,DNA
R6714	T10820	T10818	pobj	DNA,of
R6715	T10821	T10820	amod	genomic,DNA
R6716	T10822	T10813	punct	", ",made
R6717	T10823	T10813	prep	in,made
R6718	T10824	T10825	det	a,fashion
R6719	T10825	T10823	pobj	fashion,in
R672	T1568	T1569	mark	that,function
R6720	T10826	T10825	amod	similar,fashion
R6721	T10827	T10825	prep	to,fashion
R6722	T10828	T10827	pobj	those,to
R6723	T10829	T10828	acl	described,those
R6724	T10830	T10829	advmod	previously,described
R6725	T10831	T10832	punct	[,1
R6726	T10832	T10829	parataxis	1,described
R6727	T10833	T10832	punct	],1
R6728	T10834	T10813	punct	", ",made
R6729	T10835	T10836	mark	with,given
R673	T1569	T1559	ccomp	function,indicate
R6730	T10836	T10813	advcl	given,made
R6731	T10837	T10838	compound	primer,pairs
R6732	T10838	T10836	nsubj	pairs,given
R6733	T10839	T10838	cc	and,pairs
R6734	T10840	T10841	amod	annealing,temperatures
R6735	T10841	T10838	conj	temperatures,pairs
R6736	T10842	T10836	prep	in,given
R6737	T10843	T10842	pobj	Table,in
R6738	T10844	T10843	nummod	2,Table
R6739	T10845	T10813	punct	.,made
R674	T1570	T1569	nsubj	it,function
R6740	T10847	T10848	amod	Low,stringency
R6741	T10848	T10850	compound	stringency,conditions
R6742	T10849	T10848	punct	-,stringency
R6743	T10850	T10852	nsubj	conditions,were
R6744	T10851	T10850	compound	hybridization,conditions
R6745	T10853	T10854	mark	as,described
R6746	T10854	T10852	advcl	described,were
R6747	T10855	T10856	punct	[,1
R6748	T10856	T10854	parataxis	1,described
R6749	T10857	T10856	punct	],1
R675	T1571	T1569	aux	does,function
R6750	T10858	T10852	punct	.,were
R6751	T10860	T10861	advmod	Clonally,pure
R6752	T10861	T10863	amod	pure,plaques
R6753	T10862	T10861	punct	-,pure
R6754	T10863	T10864	nsubjpass	plaques,obtained
R6755	T10865	T10864	auxpass	were,obtained
R6756	T10866	T10864	prep	through,obtained
R6757	T10867	T10868	amod	secondary,screens
R6758	T10868	T10866	pobj	screens,through
R6759	T10869	T10864	advcl	using,obtained
R676	T1572	T1569	advmod	indeed,function
R6760	T10870	T10871	det	the,probe
R6761	T10871	T10869	dobj	probe,using
R6762	T10872	T10871	amod	same,probe
R6763	T10873	T10871	prep	as,probe
R6764	T10874	T10875	det	the,screen
R6765	T10875	T10873	pobj	screen,as
R6766	T10876	T10875	amod	corresponding,screen
R6767	T10877	T10875	amod	primary,screen
R6768	T10878	T10864	punct	.,obtained
R6769	T10880	T10881	nsubjpass	PCR,performed
R677	T1573	T1569	prep	in,function
R6770	T10882	T10880	prep	with,PCR
R6771	T10883	T10884	compound	vector,primers
R6772	T10884	T10882	pobj	primers,with
R6773	T10885	T10886	punct	(,R
R6774	T10886	T10880	parataxis	R,PCR
R6775	T10887	T10886	compound	M13F,R
R6776	T10888	T10886	punct	/,R
R6777	T10889	T10886	punct	),R
R6778	T10890	T10881	auxpass	was,performed
R6779	T10891	T10892	aux	to,prepare
R678	T1574	T1575	compound	sperm,chemotaxis
R6780	T10892	T10881	advcl	prepare,performed
R6781	T10893	T10894	compound	sequencing,templates
R6782	T10894	T10892	dobj	templates,prepare
R6783	T10895	T10881	punct	.,performed
R6784	T10897	T10898	compound	cDNA,size
R6785	T10898	T10899	compound	size,estimates
R6786	T10899	T10900	nsubjpass	estimates,obtained
R6787	T10901	T10900	auxpass	were,obtained
R6788	T10902	T10900	prep	by,obtained
R6789	T10903	T10904	compound	agarose,gel
R679	T1575	T1573	pobj	chemotaxis,in
R6790	T10904	T10905	compound	gel,electrophoresis
R6791	T10905	T10902	pobj	electrophoresis,by
R6792	T10906	T10900	punct	", ",obtained
R6793	T10907	T10900	cc	and,obtained
R6794	T10908	T10909	nsubjpass	inserts,sequenced
R6795	T10909	T10900	conj	sequenced,obtained
R6796	T10910	T10909	auxpass	were,sequenced
R6797	T10911	T10909	prep	from,sequenced
R6798	T10912	T10913	det	the,end
R6799	T10913	T10911	pobj	end,from
R68	T372	T373	amod	real,time
R680	T1576	T1577	punct	[,13
R6800	T10914	T10913	nummod	5,end
R6801	T10915	T10914	punct	',5
R6802	T10916	T10909	advcl	using,sequenced
R6803	T10917	T10918	det	the,primer
R6804	T10918	T10916	dobj	primer,using
R6805	T10919	T10918	compound	M13R,primer
R6806	T10920	T10918	cc	and,primer
R6807	T10921	T10922	amod	big,dye
R6808	T10922	T10924	compound	dye,chemistry
R6809	T10923	T10922	punct	-,dye
R681	T1577	T1559	parataxis	13,indicate
R6810	T10924	T10918	conj	chemistry,primer
R6811	T10925	T10924	compound	terminator,chemistry
R6812	T10926	T10916	prep	according,using
R6813	T10927	T10926	prep	to,according
R6814	T10928	T10929	poss	ABI,protocols
R6815	T10929	T10927	pobj	protocols,to
R6816	T10930	T10928	case	's,ABI
R6817	T10931	T10932	punct	(,Biosystems
R6818	T10932	T10929	parataxis	Biosystems,protocols
R6819	T10933	T10932	compound	Applied,Biosystems
R682	T1578	T1577	punct	],13
R6820	T10934	T10932	punct	", ",Biosystems
R6821	T10935	T10936	compound	Foster,City
R6822	T10936	T10932	npadvmod	City,Biosystems
R6823	T10937	T10932	punct	", ",Biosystems
R6824	T10938	T10932	npadvmod	CA,Biosystems
R6825	T10939	T10932	punct	", ",Biosystems
R6826	T10940	T10932	npadvmod	USA,Biosystems
R6827	T10941	T10932	punct	),Biosystems
R6828	T10942	T10909	punct	.,sequenced
R6829	T10944	T10945	prep	In,converted
R683	T1579	T1559	punct	.,indicate
R6830	T10946	T10944	pobj	order,In
R6831	T10947	T10948	aux	to,obtain
R6832	T10948	T10946	acl	obtain,order
R6833	T10949	T10950	nummod	3,sequence
R6834	T10950	T10948	dobj	sequence,obtain
R6835	T10951	T10949	punct	',3
R6836	T10952	T10945	punct	", ",converted
R6837	T10953	T10954	amod	selected,clones
R6838	T10954	T10945	nsubjpass	clones,converted
R6839	T10955	T10954	compound	phage,clones
R684	T1581	T1582	prep	Due,embarked
R6840	T10956	T10945	auxpass	were,converted
R6841	T10957	T10945	prep	to,converted
R6842	T10958	T10959	compound	plasmid,stocks
R6843	T10959	T10957	pobj	stocks,to
R6844	T10960	T10945	prep	following,converted
R6845	T10961	T10962	det	a,version
R6846	T10962	T10960	pobj	version,following
R6847	T10963	T10962	amod	scaled,version
R6848	T10964	T10963	punct	-,scaled
R6849	T10965	T10963	prt	down,scaled
R685	T1583	T1581	pcomp	to,Due
R6850	T10966	T10962	prep	of,version
R6851	T10967	T10968	poss	Stratagene,protocol
R6852	T10968	T10966	pobj	protocol,of
R6853	T10969	T10967	case	's,Stratagene
R6854	T10970	T10971	advmod	in,vivo
R6855	T10971	T10968	amod	vivo,protocol
R6856	T10972	T10968	compound	excision,protocol
R6857	T10973	T10945	punct	.,converted
R6858	T10975	T10976	compound	Plasmid,DNA
R6859	T10976	T10977	nsubj	DNA,gave
R686	T1584	T1585	det	the,paucity
R6860	T10978	T10979	amod	better,sequence
R6861	T10979	T10977	dobj	sequence,gave
R6862	T10980	T10981	nummod	3,end
R6863	T10981	T10979	compound	end,sequence
R6864	T10982	T10980	punct	',3
R6865	T10983	T10981	punct	-,end
R6866	T10984	T10979	prep	than,sequence
R6867	T10985	T10986	compound	PCR,products
R6868	T10986	T10984	pobj	products,than
R6869	T10987	T10986	punct	", ",products
R687	T1585	T1581	pobj	paucity,Due
R6870	T10988	T10989	dep	which,suffered
R6871	T10989	T10986	relcl	suffered,products
R6872	T10990	T10989	advmod	often,suffered
R6873	T10991	T10989	prep	from,suffered
R6874	T10992	T10991	pobj	polymerase,from
R6875	T10993	T10992	amod	stuttering,polymerase
R6876	T10994	T10989	prep	through,suffered
R6877	T10995	T10996	det	the,tract
R6878	T10996	T10994	pobj	tract,through
R6879	T10997	T10996	nmod	poly,tract
R688	T1586	T1585	prep	of,paucity
R6880	T10998	T10997	punct	(,poly
R6881	T10999	T10997	appos	A,poly
R6882	T11000	T10996	punct	),tract
R6883	T11001	T10977	punct	.,gave
R6884	T11003	T11004	compound	cDNA,sequences
R6885	T11004	T11005	nsubj	sequences,are
R6886	T11006	T11004	cc	and,sequences
R6887	T11007	T11008	amod	associated,information
R6888	T11008	T11004	conj	information,sequences
R6889	T11009	T11005	acomp	available,are
R689	T1587	T1588	amod	experimental,evidence
R6890	T11010	T11005	prep	through,are
R6891	T11011	T11010	pobj	dbEST,through
R6892	T11012	T11013	punct	(,CB172832
R6893	T11013	T11011	parataxis	CB172832,dbEST
R6894	T11014	T11013	compound	Genbank,CB172832
R6895	T11015	T11013	compound	accessions,CB172832
R6896	T11016	T11017	punct	-,CB174569
R6897	T11017	T11013	prep	CB174569,CB172832
R6898	T11018	T11013	punct	),CB172832
R6899	T11019	T11011	cc	and,dbEST
R69	T373	T375	compound	time,PCR
R690	T1588	T1586	pobj	evidence,of
R6900	T11020	T11021	poss	our,database
R6901	T11021	T11011	conj	database,dbEST
R6902	T11022	T11023	amod	olfactory,receptor
R6903	T11023	T11021	compound	receptor,database
R6904	T11024	T11025	punct	[,46
R6905	T11025	T11005	parataxis	46,are
R6906	T11026	T11025	punct	],46
R6907	T11027	T11005	punct	.,are
R6908	T11029	T11030	det	The,catalog
R6909	T11030	T11035	nsubj	catalog,is
R691	T1589	T1588	prep	of,evidence
R6910	T11031	T11030	amod	updated,catalog
R6911	T11032	T11033	amod	olfactory,receptor
R6912	T11033	T11034	compound	receptor,gene
R6913	T11034	T11030	compound	gene,catalog
R6914	T11036	T11035	acomp	available,is
R6915	T11037	T11035	prep	through,is
R6916	T11038	T11037	pobj	Genbank,through
R6917	T11039	T11040	punct	(,AY317244
R6918	T11040	T11038	parataxis	AY317244,Genbank
R6919	T11041	T11040	compound	accessions,AY317244
R692	T1590	T1591	det	the,function
R6920	T11042	T11043	punct	-,AY318733
R6921	T11043	T11040	prep	AY318733,AY317244
R6922	T11044	T11040	punct	),AY317244
R6923	T11045	T11035	punct	.,is
R6924	T11047	T11048	prep	Throughout,referred
R6925	T11049	T11050	det	the,manuscript
R6926	T11050	T11047	pobj	manuscript,Throughout
R6927	T11051	T11048	punct	", ",referred
R6928	T11052	T11048	nsubjpass	genes,referred
R6929	T11053	T11048	auxpass	are,referred
R693	T1591	T1589	pobj	function,of
R6930	T11054	T11048	prep	to,referred
R6931	T11055	T11048	prep	by,referred
R6932	T11056	T11057	poss	their,numbers
R6933	T11057	T11055	pobj	numbers,by
R6934	T11058	T11057	compound	Genbank,numbers
R6935	T11059	T11057	compound	accession,numbers
R6936	T11060	T11048	punct	.,referred
R6937	T11276	T11277	compound	Sequence,analysis
R6938	T11279	T11280	compound	cDNA,sequences
R6939	T11280	T11281	nsubj	sequences,called
R694	T1592	T1591	amod	olfactory,function
R6940	T11282	T11281	aux	were,called
R6941	T11283	T11281	dep	base,called
R6942	T11284	T11281	punct	-,called
R6943	T11285	T11281	cc	and,called
R6944	T11286	T11287	dep	quality,trimmed
R6945	T11287	T11281	conj	trimmed,called
R6946	T11288	T11287	punct	-,trimmed
R6947	T11289	T11281	advcl	using,called
R6948	T11290	T11289	dobj	phred,using
R6949	T11291	T11292	punct	(,0.05
R695	T1593	T1591	prep	of,function
R6950	T11292	T11281	parataxis	0.05,called
R6951	T11293	T11292	nsubj	trim_cutoff,0.05
R6952	T11294	T11292	punct	=,0.05
R6953	T11295	T11292	punct	),0.05
R6954	T11296	T11297	punct	[,47
R6955	T11297	T11281	parataxis	47,called
R6956	T11298	T11297	punct	],47
R6957	T11299	T11281	punct	", ",called
R6958	T11300	T11281	cc	and,called
R6959	T11301	T11302	compound	vector,sequences
R696	T1594	T1595	amod	most,genes
R6960	T11302	T11303	nsubjpass	sequences,removed
R6961	T11303	T11281	conj	removed,called
R6962	T11304	T11303	auxpass	were,removed
R6963	T11305	T11303	advcl	using,removed
R6964	T11306	T11305	dobj	cross_match,using
R6965	T11307	T11308	punct	[,48
R6966	T11308	T11303	parataxis	48,removed
R6967	T11309	T11308	punct	],48
R6968	T11310	T11303	punct	.,removed
R6969	T11312	T11313	det	Any,sequences
R697	T1595	T1593	pobj	genes,of
R6970	T11313	T11314	nsubjpass	sequences,discarded
R6971	T11315	T11313	prep	of,sequences
R6972	T11316	T11317	amod	less,50
R6973	T11317	T11319	nummod	50,bp
R6974	T11318	T11317	quantmod	than,50
R6975	T11319	T11315	pobj	bp,of
R6976	T11320	T11319	prep	after,bp
R6977	T11321	T11320	pobj	trimming,after
R6978	T11322	T11314	auxpass	were,discarded
R6979	T11323	T11314	punct	.,discarded
R698	T1596	T1595	prep	in,genes
R6980	T11325	T11326	nummod	3,UTR
R6981	T11326	T11328	compound	UTR,lengths
R6982	T11327	T11325	punct	',3
R6983	T11328	T11329	nsubjpass	lengths,estimated
R6984	T11330	T11329	auxpass	were,estimated
R6985	T11331	T11329	prep	by,estimated
R6986	T11332	T11331	pcomp	combining,by
R6987	T11333	T11334	amod	approximate,insert
R6988	T11334	T11332	dobj	insert,combining
R6989	T11335	T11334	nmod	sizes,insert
R699	T1597	T1598	det	the,family
R6990	T11336	T11334	acl	determined,insert
R6991	T11337	T11336	prep	by,determined
R6992	T11338	T11337	pobj	PCR,by
R6993	T11339	T11338	prep	with,PCR
R6994	T11340	T11341	nummod	5,sequence
R6995	T11341	T11343	compound	sequence,data
R6996	T11342	T11340	punct	',5
R6997	T11343	T11339	pobj	data,with
R6998	T11344	T11345	advmod	where,possible
R6999	T11345	T11336	advcl	possible,determined
R7	T303	T304	amod	olfactory,function
R70	T374	T373	punct	-,time
R700	T1598	T1596	pobj	family,in
R7000	T11346	T11347	punct	(,estimate
R7001	T11347	T11329	parataxis	estimate,estimated
R7002	T11348	T11349	mark	if,extend
R7003	T11349	T11347	advcl	extend,estimate
R7004	T11350	T11351	det	the,sequence
R7005	T11351	T11349	nsubj	sequence,extend
R7006	T11352	T11351	nummod	5,sequence
R7007	T11353	T11352	punct	',5
R7008	T11354	T11349	aux	did,extend
R7009	T11355	T11349	neg	not,extend
R701	T1599	T1585	cc	and,paucity
R7010	T11356	T11349	prep	into,extend
R7011	T11357	T11358	det	the,region
R7012	T11358	T11356	pobj	region,into
R7013	T11359	T11358	amod	coding,region
R7014	T11360	T11347	nsubj	we,estimate
R7015	T11361	T11347	aux	could,estimate
R7016	T11362	T11347	neg	not,estimate
R7017	T11363	T11364	nummod	3,UTR
R7018	T11364	T11366	compound	UTR,size
R7019	T11365	T11363	punct	',3
R702	T1600	T1585	conj	suggestions,paucity
R7020	T11366	T11347	dobj	size,estimate
R7021	T11367	T11347	punct	),estimate
R7022	T11368	T11329	punct	.,estimated
R7023	T11370	T11371	nsubj	We,counted
R7024	T11371	T11372	ccomp	counted,be
R7025	T11373	T11371	dobj	cDNAs,counted
R7026	T11374	T11373	prep	from,cDNAs
R7027	T11375	T11376	det	a,gene
R7028	T11376	T11374	pobj	gene,from
R7029	T11377	T11376	amod	given,gene
R703	T1601	T1600	prep	of,suggestions
R7030	T11378	T11371	prep	as,counted
R7031	T11379	T11378	pcomp	showing,as
R7032	T11380	T11381	amod	alternative,site
R7033	T11381	T11383	compound	site,usage
R7034	T11382	T11381	compound	polyadenylation,site
R7035	T11383	T11379	dobj	usage,showing
R7036	T11384	T11385	mark	if,varied
R7037	T11385	T11371	advcl	varied,counted
R7038	T11386	T11387	nummod	3,UTR
R7039	T11387	T11389	compound	UTR,length
R704	T1602	T1603	amod	extra-olfactory,roles
R7040	T11388	T11386	punct	',3
R7041	T11389	T11390	compound	length,estimates
R7042	T11390	T11385	nsubj	estimates,varied
R7043	T11391	T11385	prep	by,varied
R7044	T11392	T11393	advmod	at,400
R7045	T11393	T11395	nummod	400,bp
R7046	T11394	T11393	advmod	least,400
R7047	T11395	T11391	pobj	bp,by
R7048	T11396	T11372	punct	-,be
R7049	T11397	T11398	amod	smaller,variation
R705	T1603	T1601	pobj	roles,of
R7050	T11398	T11372	nsubj	variation,be
R7051	T11399	T11372	aux	could,be
R7052	T11400	T11372	acomp	real,be
R7053	T11401	T11372	punct	", ",be
R7054	T11402	T11372	cc	but,be
R7055	T11403	T11404	aux	may,be
R7056	T11404	T11372	conj	be,be
R7057	T11405	T11404	neg	not,be
R7058	T11406	T11404	acomp	distinguishable,be
R7059	T11407	T11406	prep	from,distinguishable
R706	T1604	T1582	punct	", ",embarked
R7060	T11408	T11407	pobj	error,from
R7061	T11409	T11404	prep	in,be
R7062	T11410	T11411	poss	our,estimates
R7063	T11411	T11409	pobj	estimates,in
R7064	T11412	T11411	compound	size,estimates
R7065	T11413	T11372	punct	.,be
R7066	T11415	T11416	aux	To,assign
R7067	T11416	T11417	advcl	assign,defined
R7068	T11418	T11416	dobj	cDNAs,assign
R7069	T11419	T11416	prep	to,assign
R707	T1605	T1582	nsubj	we,embarked
R7070	T11420	T11421	poss	their,genes
R7071	T11421	T11419	pobj	genes,to
R7072	T11422	T11421	amod	corresponding,genes
R7073	T11423	T11424	amod	olfactory,receptor
R7074	T11424	T11421	compound	receptor,genes
R7075	T11425	T11417	punct	", ",defined
R7076	T11426	T11417	nsubj	we,defined
R7077	T11427	T11417	advmod	first,defined
R7078	T11428	T11429	det	a,territory
R7079	T11429	T11417	dobj	territory,defined
R708	T1606	T1582	prep	on,embarked
R7080	T11430	T11429	amod	genomic,territory
R7081	T11431	T11429	punct	',territory
R7082	T11432	T11429	punct	',territory
R7083	T11433	T11429	prep	for,territory
R7084	T11434	T11435	det	each,gene
R7085	T11435	T11433	pobj	gene,for
R7086	T11436	T11429	punct	", ",territory
R7087	T11437	T11429	prep	with,territory
R7088	T11438	T11439	det	the,attributes
R7089	T11439	T11437	pobj	attributes,with
R709	T1607	T1608	det	an,project
R7090	T11440	T11439	amod	following,attributes
R7091	T11441	T11439	punct	: ,attributes
R7092	T11442	T11439	appos	strand,attributes
R7093	T11443	T11442	punct	", ",strand
R7094	T11444	T11445	compound	start,position
R7095	T11445	T11442	conj	position,strand
R7096	T11446	T11447	punct	(,is
R7097	T11447	T11445	parataxis	is,position
R7098	T11448	T11449	nummod	100,kb
R7099	T11449	T11450	npadvmod	kb,upstream
R71	T375	T370	dobj	PCR,Using
R710	T1608	T1606	pobj	project,on
R7100	T11450	T11447	advmod	upstream,is
R7101	T11451	T11450	prep	of,upstream
R7102	T11452	T11453	det	the,codon
R7103	T11453	T11451	pobj	codon,of
R7104	T11454	T11453	compound	start,codon
R7105	T11455	T11450	cc	or,upstream
R7106	T11456	T11457	nummod	1,kb
R7107	T11457	T11458	npadvmod	kb,after
R7108	T11458	T11459	prep	after,upstream
R7109	T11459	T11450	conj	upstream,upstream
R711	T1609	T1610	amod	olfactory,receptor
R7110	T11460	T11461	det	the,gene
R7111	T11461	T11458	pobj	gene,after
R7112	T11462	T11461	amod	previous,gene
R7113	T11463	T11459	prep	on,upstream
R7114	T11464	T11465	det	the,strand
R7115	T11465	T11463	pobj	strand,on
R7116	T11466	T11465	amod	same,strand
R7117	T11467	T11447	punct	", ",is
R7118	T11468	T11447	dep	whichever,is
R7119	T11469	T11447	acomp	closer,is
R712	T1610	T1608	nmod	receptor,project
R7120	T11470	T11447	punct	),is
R7121	T11471	T11445	cc	and,position
R7122	T11472	T11473	compound	end,position
R7123	T11473	T11445	conj	position,position
R7124	T11474	T11475	punct	(,downstream
R7125	T11475	T11473	parataxis	downstream,position
R7126	T11476	T11477	nummod	1,kb
R7127	T11477	T11475	npadvmod	kb,downstream
R7128	T11478	T11475	prep	of,downstream
R7129	T11479	T11480	det	the,codon
R713	T1611	T1612	amod	expressed,tag
R7130	T11480	T11478	pobj	codon,of
R7131	T11481	T11480	compound	stop,codon
R7132	T11482	T11475	punct	),downstream
R7133	T11483	T11417	punct	.,defined
R7134	T11485	T11486	amod	Trimmed,sequences
R7135	T11486	T11487	nsubjpass	sequences,compared
R7136	T11488	T11487	auxpass	were,compared
R7137	T11489	T11487	prep	with,compared
R7138	T11490	T11491	amod	genomic,sequences
R7139	T11491	T11489	pobj	sequences,with
R714	T1612	T1610	appos	tag,receptor
R7140	T11492	T11487	advcl	using,compared
R7141	T11493	T11492	dobj	sim4,using
R7142	T11494	T11495	punct	[,30
R7143	T11495	T11487	parataxis	30,compared
R7144	T11496	T11495	punct	],30
R7145	T11497	T11498	punct	(,settings
R7146	T11498	T11487	parataxis	settings,compared
R7147	T11499	T11500	nsubj	P,1
R7148	T11500	T11498	dep	1,settings
R7149	T11501	T11500	punct	=,1
R715	T1613	T1612	compound	sequence,tag
R7150	T11502	T11503	aux	to,remove
R7151	T11503	T11500	advcl	remove,1
R7152	T11504	T11505	compound	polyA,tails
R7153	T11505	T11503	dobj	tails,remove
R7154	T11506	T11500	cc	and,1
R7155	T11507	T11508	nsubj	N,1
R7156	T11508	T11510	ccomp	1,perform
R7157	T11509	T11508	punct	=,1
R7158	T11510	T11500	conj	perform,1
R7159	T11511	T11510	aux	to,perform
R716	T1614	T1612	punct	(,tag
R7160	T11512	T11513	det	an,search
R7161	T11513	T11510	dobj	search,perform
R7162	T11514	T11513	amod	intensive,search
R7163	T11515	T11513	prep	for,search
R7164	T11516	T11517	amod	small,exons
R7165	T11517	T11515	pobj	exons,for
R7166	T11518	T11498	punct	),settings
R7167	T11519	T11487	punct	.,compared
R7168	T11521	T11522	det	The,algorithm
R7169	T11522	T11524	nsubj	algorithm,uses
R717	T1615	T1612	appos	EST,tag
R7170	T11523	T11522	compound	sim4,algorithm
R7171	T11525	T11526	compound	splice,site
R7172	T11526	T11528	compound	site,sequences
R7173	T11527	T11526	punct	-,site
R7174	T11528	T11524	dobj	sequences,uses
R7175	T11529	T11528	compound	consensus,sequences
R7176	T11530	T11531	aux	to,refine
R7177	T11531	T11524	advcl	refine,uses
R7178	T11532	T11531	dep	alignments,refine
R7179	T11533	T11524	punct	.,uses
R718	T1616	T1608	punct	),project
R7180	T11535	T11536	advmod	Only,matches
R7181	T11536	T11537	nsubjpass	matches,considered
R7182	T11538	T11536	prep	of,matches
R7183	T11539	T11540	nummod	96,%
R7184	T11540	T11541	nmod	%,identity
R7185	T11541	T11538	pobj	identity,of
R7186	T11542	T11543	cc	or,greater
R7187	T11543	T11540	nummod	greater,%
R7188	T11544	T11541	compound	nucleotide,identity
R7189	T11545	T11537	auxpass	were,considered
R719	T1617	T1618	aux	to,confirm
R7190	T11546	T11537	punct	.,considered
R7191	T11548	T11549	amod	RepeatMasked,sequences
R7192	T11549	T11550	nsubjpass	sequences,compared
R7193	T11551	T11552	punct	[,49
R7194	T11552	T11549	parataxis	49,sequences
R7195	T11553	T11552	punct	],49
R7196	T11554	T11550	auxpass	were,compared
R7197	T11555	T11550	advmod	also,compared
R7198	T11556	T11550	prep	to,compared
R7199	T11557	T11558	amod	genomic,sequences
R72	T376	T377	aux	to,confirm
R720	T1618	T1582	advcl	confirm,embarked
R7200	T11558	T11556	pobj	sequences,to
R7201	T11559	T11550	punct	;,compared
R7202	T11560	T11561	dep	cDNA,rejected
R7203	T11561	T11550	dep	rejected,compared
R7204	T11562	T11561	punct	:,rejected
R7205	T11563	T11564	amod	genomic,sequence
R7206	T11564	T11565	compound	sequence,pairings
R7207	T11565	T11561	nsubjpass	pairings,rejected
R7208	T11566	T11567	neg	not,found
R7209	T11567	T11565	acl	found,pairings
R721	T1619	T1620	amod	olfactory,expression
R7210	T11568	T11567	prep	in,found
R7211	T11569	T11570	preconj	both,masked
R7212	T11570	T11571	amod	masked,alignments
R7213	T11571	T11568	pobj	alignments,in
R7214	T11572	T11570	cc	and,masked
R7215	T11573	T11570	conj	unmasked,masked
R7216	T11574	T11561	auxpass	were,rejected
R7217	T11575	T11550	punct	.,compared
R7218	T11577	T11578	nsubjpass	Coordinates,used
R7219	T11579	T11577	prep	from,Coordinates
R722	T1620	T1618	dobj	expression,confirm
R7220	T11580	T11581	det	the,alignment
R7221	T11581	T11579	pobj	alignment,from
R7222	T11582	T11581	amod	unmasked,alignment
R7223	T11583	T11578	auxpass	were,used
R7224	T11584	T11578	prep	for,used
R7225	T11585	T11586	amod	further,analysis
R7226	T11586	T11584	pobj	analysis,for
R7227	T11587	T11578	punct	.,used
R7228	T11589	T11590	det	Any,sequence
R7229	T11590	T11592	nsubjpass	sequence,assigned
R723	T1621	T1620	amod	epithelial,expression
R7230	T11591	T11590	compound	cDNA,sequence
R7231	T11593	T11590	acl	matching,sequence
R7232	T11594	T11595	advmod	entirely,within
R7233	T11595	T11593	prep	within,matching
R7234	T11596	T11597	det	a,territory
R7235	T11597	T11595	pobj	territory,within
R7236	T11598	T11592	auxpass	was,assigned
R7237	T11599	T11592	prep	to,assigned
R7238	T11600	T11601	det	that,gene
R7239	T11601	T11599	pobj	gene,to
R724	T1622	T1620	prep	of,expression
R7240	T11602	T11592	punct	.,assigned
R7241	T11604	T11605	mark	If,matched
R7242	T11605	T11608	advcl	matched,chosen
R7243	T11606	T11607	det	a,cDNA
R7244	T11607	T11605	nsubj	cDNA,matched
R7245	T11609	T11610	amod	more,one
R7246	T11610	T11612	nummod	one,territory
R7247	T11611	T11610	quantmod	than,one
R7248	T11612	T11605	dobj	territory,matched
R7249	T11613	T11612	compound	gene,territory
R725	T1623	T1622	pobj	hundreds,of
R7250	T11614	T11608	punct	", ",chosen
R7251	T11615	T11616	det	the,match
R7252	T11616	T11608	nsubjpass	match,chosen
R7253	T11617	T11616	amod	best,match
R7254	T11618	T11608	auxpass	was,chosen
R7255	T11619	T11620	punct	(,one
R7256	T11620	T11608	parataxis	one,chosen
R7257	T11621	T11622	advmod	that,is
R7258	T11622	T11620	advmod	is,one
R7259	T11623	T11620	punct	", ",one
R726	T1624	T1623	prep	of,hundreds
R7260	T11624	T11620	det	the,one
R7261	T11625	T11620	prep	with,one
R7262	T11626	T11627	amod	highest,score
R7263	T11627	T11625	pobj	score,with
R7264	T11628	T11627	punct	',score
R7265	T11629	T11620	punct	',one
R7266	T11630	T11620	punct	", ",one
R7267	T11631	T11632	advmod	where,is
R7268	T11632	T11620	relcl	is,one
R7269	T11633	T11632	nsubj	score,is
R727	T1625	T1626	compound	mouse,genes
R7270	T11634	T11635	det	the,total
R7271	T11635	T11632	attr	total,is
R7272	T11636	T11635	prep	of,total
R7273	T11637	T11638	det	all,exons
R7274	T11638	T11639	poss	exons,lengths
R7275	T11639	T11636	pobj	lengths,of
R7276	T11640	T11638	case	',exons
R7277	T11641	T11635	acl	multiplied,total
R7278	T11642	T11641	prep	by,multiplied
R7279	T11643	T11644	poss	their,identities
R728	T1626	T1624	pobj	genes,of
R7280	T11644	T11642	pobj	identities,by
R7281	T11645	T11644	amod	respective,identities
R7282	T11646	T11644	compound	percent,identities
R7283	T11647	T11620	punct	),one
R7284	T11648	T11608	punct	.,chosen
R7285	T11650	T11651	nsubj	We,flagged
R7286	T11652	T11651	aux	found,flagged
R7287	T11653	T11654	nummod	27,cDNAs
R7288	T11654	T11651	dative	cDNAs,flagged
R7289	T11655	T11656	dep	that,spanned
R729	T1627	T1628	compound	odorant,receptor
R7290	T11656	T11654	relcl	spanned,cDNAs
R7291	T11657	T11658	det	a,range
R7292	T11658	T11656	dobj	range,spanned
R7293	T11659	T11658	amod	larger,range
R7294	T11660	T11658	amod	genomic,range
R7295	T11661	T11658	prep	than,range
R7296	T11662	T11663	nummod	one,territory
R7297	T11663	T11661	pobj	territory,than
R7298	T11664	T11663	compound	gene,territory
R7299	T11665	T11651	cc	and,flagged
R73	T377	T370	advcl	confirm,Using
R730	T1628	T1626	compound	receptor,genes
R7300	T11666	T11651	dobj	them,flagged
R7301	T11667	T11651	prep	for,flagged
R7302	T11668	T11669	advmod	more,careful
R7303	T11669	T11670	amod	careful,analysis
R7304	T11670	T11667	pobj	analysis,for
R7305	T11671	T11651	punct	.,flagged
R7306	T11673	T11674	prep	Of,showed
R7307	T11675	T11673	pobj	these,Of
R7308	T11676	T11674	punct	", ",showed
R7309	T11677	T11678	nummod	six,cDNAs
R731	T1629	T1582	punct	.,embarked
R7310	T11678	T11674	nsubj	cDNAs,showed
R7311	T11679	T11680	amod	unusual,splicing
R7312	T11680	T11674	dobj	splicing,showed
R7313	T11681	T11674	prep	within,showed
R7314	T11682	T11683	det	the,UTR
R7315	T11683	T11681	pobj	UTR,within
R7316	T11684	T11683	nummod	3,UTR
R7317	T11685	T11684	punct	',3
R7318	T11686	T11674	punct	", ",showed
R7319	T11687	T11674	cc	but,showed
R732	T1631	T1632	prep	Within,show
R7320	T11688	T11689	det	the,violators
R7321	T11689	T11693	nsubjpass	violators,found
R7322	T11690	T11689	amod	remaining,violators
R7323	T11691	T11689	punct	',violators
R7324	T11692	T11689	compound	territory,violators
R7325	T11693	T11674	conj	found,showed
R7326	T11694	T11693	punct	',found
R7327	T11695	T11693	auxpass	were,found
R7328	T11696	T11697	aux	to,be
R7329	T11697	T11693	xcomp	be,found
R733	T1633	T1634	det	the,epithelium
R7330	T11698	T11697	attr	artifacts,be
R7331	T11699	T11698	prep	of,artifacts
R7332	T11700	T11701	det	the,process
R7333	T11701	T11699	pobj	process,of
R7334	T11702	T11701	compound	analysis,process
R7335	T11703	T11704	dep	which,fell
R7336	T11704	T11698	relcl	fell,artifacts
R7337	T11705	T11704	prep	into,fell
R7338	T11706	T11707	nummod	three,types
R7339	T11707	T11705	pobj	types,into
R734	T1634	T1631	pobj	epithelium,Within
R7340	T11708	T11674	punct	.,showed
R7341	T11710	T11711	nsubj	These,included
R7342	T11712	T11711	punct	: ,included
R7343	T11713	T11711	dobj	cDNAs,included
R7344	T11714	T11715	advmod	where,appeared
R7345	T11715	T11713	relcl	appeared,cDNAs
R7346	T11716	T11717	det	the,insert
R7347	T11717	T11715	nsubj	insert,appeared
R7348	T11718	T11719	aux	to,cloned
R7349	T11719	T11715	xcomp	cloned,appeared
R735	T1635	T1634	amod	olfactory,epithelium
R7350	T11720	T11719	auxpass	be,cloned
R7351	T11721	T11719	prep	in,cloned
R7352	T11722	T11723	det	the,orientation
R7353	T11723	T11721	pobj	orientation,in
R7354	T11724	T11723	amod	reverse,orientation
R7355	T11725	T11726	punct	(,cDNAs
R7356	T11726	T11713	parataxis	cDNAs,cDNAs
R7357	T11727	T11726	nummod	six,cDNAs
R7358	T11728	T11726	punct	),cDNAs
R7359	T11729	T11713	punct	;,cDNAs
R736	T1636	T1632	punct	", ",show
R7360	T11730	T11713	conj	sequences,cDNAs
R7361	T11731	T11730	prep	from,sequences
R7362	T11732	T11733	advmod	recently,duplicated
R7363	T11733	T11734	amod	duplicated,pairs
R7364	T11734	T11731	pobj	pairs,from
R7365	T11735	T11734	compound	gene,pairs
R7366	T11736	T11730	punct	", ",sequences
R7367	T11737	T11738	advmod	where,assigned
R7368	T11738	T11730	relcl	assigned,sequences
R7369	T11739	T11738	nsubj	sim4,assigned
R737	T1637	T1638	amod	individual,genes
R7370	T11740	T11741	amod	coding,region
R7371	T11741	T11738	dobj	region,assigned
R7372	T11742	T11741	cc	and,region
R7373	T11743	T11744	amod	upstream,exons
R7374	T11744	T11741	conj	exons,region
R7375	T11745	T11738	prep	to,assigned
R7376	T11746	T11747	amod	different,members
R7377	T11747	T11745	pobj	members,to
R7378	T11748	T11747	prep	of,members
R7379	T11749	T11750	det	the,pair
R738	T1638	T1632	nsubj	genes,show
R7380	T11750	T11748	pobj	pair,of
R7381	T11751	T11738	punct	", ",assigned
R7382	T11752	T11753	mark	although,aligned
R7383	T11753	T11738	advcl	aligned,assigned
R7384	T11754	T11753	nsubjpass	exons,aligned
R7385	T11755	T11753	aux	could,aligned
R7386	T11756	T11757	advmod	equally,well
R7387	T11757	T11753	advmod	well,aligned
R7388	T11758	T11753	aux	have,aligned
R7389	T11759	T11753	auxpass	been,aligned
R739	T1639	T1640	amod	olfactory,receptor
R7390	T11760	T11753	advmod	closer,aligned
R7391	T11761	T11760	prep	to,closer
R7392	T11762	T11761	pobj	one,to
R7393	T11763	T11762	det	another,one
R7394	T11764	T11765	punct	(,cDNAs
R7395	T11765	T11730	parataxis	cDNAs,sequences
R7396	T11766	T11765	nummod	six,cDNAs
R7397	T11767	T11765	punct	),cDNAs
R7398	T11768	T11730	punct	;,sequences
R7399	T11769	T11730	cc	and,sequences
R74	T378	T379	compound	expression,level
R740	T1640	T1638	compound	receptor,genes
R7400	T11770	T11730	conj	artifacts,sequences
R7401	T11771	T11770	prep	due,artifacts
R7402	T11772	T11771	pcomp	to,due
R7403	T11773	T11771	pobj	use,due
R7404	T11774	T11773	prep	of,use
R7405	T11775	T11776	poss	sim4,parameter
R7406	T11776	T11774	pobj	parameter,of
R7407	T11777	T11775	case	's,sim4
R7408	T11778	T11779	nsubj	N,1
R7409	T11779	T11776	nummod	1,parameter
R741	T1641	T1642	det	an,pattern
R7410	T11780	T11779	punct	=,1
R7411	T11781	T11782	punct	(,cDNAs
R7412	T11782	T11770	parataxis	cDNAs,artifacts
R7413	T11783	T11782	nummod	nine,cDNAs
R7414	T11784	T11782	punct	),cDNAs
R7415	T11785	T11711	punct	.,included
R7416	T11787	T11788	det	This,parameter
R7417	T11788	T11789	nsubj	parameter,instructs
R7418	T11790	T11791	det	the,program
R7419	T11791	T11789	dobj	program,instructs
R742	T1642	T1632	dobj	pattern,show
R7420	T11792	T11793	aux	to,make
R7421	T11793	T11789	xcomp	make,instructs
R7422	T11794	T11795	amod	extra,effort
R7423	T11795	T11793	dobj	effort,make
R7424	T11796	T11797	aux	to,match
R7425	T11797	T11795	acl	match,effort
R7426	T11798	T11799	amod	small,exons
R7427	T11799	T11797	dobj	exons,match
R7428	T11800	T11799	amod	upstream,exons
R7429	T11801	T11789	punct	", ",instructs
R743	T1643	T1642	amod	intriguing,pattern
R7430	T11802	T11789	advcl	allowing,instructs
R7431	T11803	T11804	det	a,length
R7432	T11804	T11807	nsubjpass	length,matched
R7433	T11805	T11804	amod	greater,length
R7434	T11806	T11804	amod	total,length
R7435	T11807	T11802	ccomp	matched,allowing
R7436	T11808	T11804	prep	of,length
R7437	T11809	T11810	compound	EST,sequence
R7438	T11810	T11808	pobj	sequence,of
R7439	T11811	T11807	aux	to,matched
R744	T1644	T1642	compound	expression,pattern
R7440	T11812	T11807	auxpass	be,matched
R7441	T11813	T11789	punct	.,instructs
R7442	T11815	T11816	advmod	However,caused
R7443	T11817	T11816	punct	", ",caused
R7444	T11818	T11816	advmod	occasionally,caused
R7445	T11819	T11820	det	the,parameter
R7446	T11820	T11816	nsubj	parameter,caused
R7447	T11821	T11822	nsubj	N,1
R7448	T11822	T11820	nummod	1,parameter
R7449	T11823	T11822	punct	=,1
R745	T1645	T1632	punct	.,show
R7450	T11824	T11825	det	the,program
R7451	T11825	T11826	nsubj	program,assign
R7452	T11826	T11816	ccomp	assign,caused
R7453	T11827	T11826	aux	to,assign
R7454	T11828	T11829	advmod	very,small
R7455	T11829	T11830	amod	small,sequences
R7456	T11830	T11826	dobj	sequences,assign
R7457	T11831	T11832	punct	(,bp
R7458	T11832	T11830	parataxis	bp,sequences
R7459	T11833	T11834	compound	1,4
R746	T1647	T1648	det	Each,receptor
R7460	T11834	T11832	nummod	4,bp
R7461	T11835	T11834	punct	-,4
R7462	T11836	T11832	punct	),bp
R7463	T11837	T11826	prep	to,assign
R7464	T11838	T11839	amod	distant,exons
R7465	T11839	T11837	pobj	exons,to
R7466	T11840	T11839	amod	upstream,exons
R7467	T11841	T11826	punct	", ",assign
R7468	T11842	T11843	advmod	when,match
R7469	T11843	T11826	advcl	match,assign
R747	T1648	T1650	nsubjpass	receptor,expressed
R7470	T11844	T11843	nsubj	they,match
R7471	T11845	T11843	advmod	probably,match
R7472	T11846	T11843	advmod	nearer,match
R7473	T11847	T11846	prep	to,nearer
R7474	T11848	T11849	det	the,sequence
R7475	T11849	T11847	pobj	sequence,to
R7476	T11850	T11849	amod	corresponding,sequence
R7477	T11851	T11849	compound	coding,sequence
R7478	T11852	T11816	punct	.,caused
R7479	T11854	T11855	det	The,distribution
R748	T1649	T1648	amod	olfactory,receptor
R7480	T11855	T11857	nsubjpass	distribution,calculated
R7481	T11856	T11855	amod	expected,distribution
R7482	T11858	T11855	acl	shown,distribution
R7483	T11859	T11858	prep	in,shown
R7484	T11860	T11859	pobj	Figure,in
R7485	T11861	T11860	nummod	2,Figure
R7486	T11862	T11857	auxpass	was,calculated
R7487	T11863	T11857	advcl	using,calculated
R7488	T11864	T11865	det	the,equation
R7489	T11865	T11863	dobj	equation,using
R749	T1651	T1650	auxpass	is,expressed
R7490	T11866	T11867	nmod	P,x
R7491	T11867	T11869	nsubj	x,μμx
R7492	T11868	T11867	punct	(,x
R7493	T11869	T11865	appos	μμx,equation
R7494	T11870	T11867	punct	),x
R7495	T11871	T11869	punct	=,μμx
R7496	T11872	T11869	nmod	e,μμx
R7497	T11873	T11869	punct	-,μμx
R7498	T11874	T11875	punct	/,x
R7499	T11875	T11869	prep	x,μμx
R75	T379	T380	compound	level,differences
R750	T1652	T1650	prep	in,expressed
R7500	T11876	T11875	punct	!,x
R7501	T11877	T11865	punct	", ",equation
R7502	T11878	T11879	advmod	where,is
R7503	T11879	T11865	relcl	is,equation
R7504	T11880	T11881	nmod	P,x
R7505	T11881	T11879	nsubj	x,is
R7506	T11882	T11881	punct	(,x
R7507	T11883	T11881	punct	),x
R7508	T11884	T11885	det	the,probability
R7509	T11885	T11879	attr	probability,is
R751	T1653	T1654	det	a,subset
R7510	T11886	T11885	compound	Poisson,probability
R7511	T11887	T11885	prep	of,probability
R7512	T11888	T11887	pcomp	observing,of
R7513	T11889	T11890	compound	x,cDNAs
R7514	T11890	T11888	dobj	cDNAs,observing
R7515	T11891	T11890	prep	per,cDNAs
R7516	T11892	T11891	pobj	gene,per
R7517	T11893	T11879	punct	", ",is
R7518	T11894	T11879	cc	and,is
R7519	T11895	T11896	nsubj	μ,is
R752	T1654	T1652	pobj	subset,in
R7520	T11896	T11879	conj	is,is
R7521	T11897	T11898	det	the,number
R7522	T11898	T11896	attr	number,is
R7523	T11899	T11898	amod	mean,number
R7524	T11900	T11898	prep	of,number
R7525	T11901	T11900	pobj	cDNAs,of
R7526	T11902	T11901	acl	observed,cDNAs
R7527	T11903	T11902	prep	per,observed
R7528	T11904	T11903	pobj	gene,per
R7529	T11905	T11906	punct	(,cDNAs
R753	T1655	T1654	prep	of,subset
R7530	T11906	T11896	parataxis	cDNAs,is
R7531	T11907	T11908	nsubj	μ,1
R7532	T11908	T11906	ccomp	1,cDNAs
R7533	T11909	T11908	punct	=,1
R7534	T11910	T11906	punct	",",cDNAs
R7535	T11911	T11912	nummod	176,983
R7536	T11912	T11906	dep	983,cDNAs
R7537	T11913	T11912	punct	/,983
R7538	T11914	T11906	punct	: ,cDNAs
R7539	T11915	T11906	nummod	"1,176",cDNAs
R754	T1656	T1655	pobj	cells,of
R7540	T11916	T11906	acl	matching,cDNAs
R7541	T11917	T11918	amod	olfactory,receptor
R7542	T11918	T11919	compound	receptor,genes
R7543	T11919	T11916	dobj	genes,matching
R7544	T11920	T11916	prep	in,matching
R7545	T11921	T11922	poss	our,dataset
R7546	T11922	T11920	pobj	dataset,in
R7547	T11923	T11906	cc	and,cDNAs
R7548	T11924	T11925	nummod	983,receptors
R7549	T11925	T11906	conj	receptors,cDNAs
R755	T1657	T1650	prep	in,expressed
R7550	T11926	T11925	amod	intact,receptors
R7551	T11927	T11925	nmod	class,receptors
R7552	T11928	T11927	nummod	II,class
R7553	T11929	T11925	amod	olfactory,receptors
R7554	T11930	T11906	punct	),cDNAs
R7555	T11931	T11857	punct	.,calculated
R7556	T11933	T11934	prep	In,ignored
R7557	T11935	T11936	poss	our,analysis
R7558	T11936	T11933	pobj	analysis,In
R7559	T11937	T11936	prep	of,analysis
R756	T1658	T1657	pobj	one,in
R7560	T11938	T11939	amod	expressed,pseudogenes
R7561	T11939	T11937	pobj	pseudogenes,of
R7562	T11940	T11934	punct	", ",ignored
R7563	T11941	T11934	nsubj	we,ignored
R7564	T11942	T11943	nummod	two,pseudogenes
R7565	T11943	T11934	dobj	pseudogenes,ignored
R7566	T11944	T11945	amod	olfactory,receptor
R7567	T11945	T11943	compound	receptor,pseudogenes
R7568	T11946	T11943	acl	found,pseudogenes
R7569	T11947	T11946	advmod	very,found
R757	T1659	T1658	prep	of,one
R7570	T11948	T11947	advmod	near,very
R7571	T11949	T11950	det	the,ends
R7572	T11950	T11947	npadvmod	ends,very
R7573	T11951	T11950	prep	of,ends
R7574	T11952	T11953	amod	genomic,sequences
R7575	T11953	T11951	pobj	sequences,of
R7576	T11954	T11946	cc	and,found
R7577	T11955	T11956	advmod	thus,likely
R7578	T11956	T11946	conj	likely,found
R7579	T11957	T11958	aux	to,be
R758	T1660	T1661	nummod	four,zones
R7580	T11958	T11956	xcomp	be,likely
R7581	T11959	T11960	npadvmod	error,prone
R7582	T11960	T11958	acomp	prone,be
R7583	T11961	T11960	punct	-,prone
R7584	T11962	T11934	punct	.,ignored
R7585	T11964	T11965	det	A,alignment
R7586	T11965	T11968	nsubjpass	alignment,generated
R7587	T11966	T11967	compound	protein,sequence
R7588	T11967	T11965	compound	sequence,alignment
R7589	T11969	T11965	prep	of,alignment
R759	T1661	T1659	pobj	zones,of
R7590	T11970	T11971	amod	intact,receptors
R7591	T11971	T11969	pobj	receptors,of
R7592	T11972	T11971	nmod	mouse,receptors
R7593	T11973	T11971	amod	olfactory,receptors
R7594	T11974	T11968	auxpass	was,generated
R7595	T11975	T11968	advcl	using,generated
R7596	T11976	T11975	dobj	CLUSTALW,using
R7597	T11977	T11978	punct	[,50
R7598	T11978	T11975	parataxis	50,using
R7599	T11979	T11978	punct	],50
R76	T380	T377	dobj	differences,confirm
R760	T1662	T1661	prep	of,zones
R7600	T11980	T11968	punct	", ",generated
R7601	T11981	T11968	conj	edited,generated
R7602	T11982	T11981	prep	by,edited
R7603	T11983	T11982	pobj	hand,by
R7604	T11984	T11981	punct	", ",edited
R7605	T11985	T11981	cc	and,edited
R7606	T11986	T11981	conj	used,edited
R7607	T11987	T11988	aux	to,produce
R7608	T11988	T11986	advcl	produce,used
R7609	T11989	T11990	det	the,tree
R761	T1663	T1664	det	the,epithelium
R7610	T11990	T11988	dobj	tree,produce
R7611	T11991	T11990	amod	phylogenetic,tree
R7612	T11992	T11990	acl	shown,tree
R7613	T11993	T11992	prep	in,shown
R7614	T11994	T11993	pobj	Figure,in
R7615	T11995	T11994	nummod	1,Figure
R7616	T11996	T11988	advcl	using,produce
R7617	T11997	T11998	poss	PAUP,algorithm
R7618	T11998	T11996	dobj	algorithm,using
R7619	T11999	T11997	case	's,PAUP
R762	T1664	T1662	pobj	epithelium,of
R7620	T12000	T12001	npadvmod	neighbor,joining
R7621	T12001	T11998	amod	joining,algorithm
R7622	T12002	T12001	punct	-,joining
R7623	T12003	T12004	punct	(,Associates
R7624	T12004	T11988	parataxis	Associates,produce
R7625	T12005	T12006	compound	v4.0b6,Version
R7626	T12006	T12004	dep	Version,Associates
R7627	T12007	T12006	nummod	4,Version
R7628	T12008	T12004	punct	", ",Associates
R7629	T12009	T12004	compound	Sinauer,Associates
R763	T1665	T1666	punct	[,15
R7630	T12010	T12004	punct	", ",Associates
R7631	T12011	T12004	npadvmod	Sunderland,Associates
R7632	T12012	T12004	punct	", ",Associates
R7633	T12013	T12004	npadvmod	MA,Associates
R7634	T12014	T12004	punct	),Associates
R7635	T12015	T11968	punct	.,generated
R7636	T12017	T12018	det	The,tree
R7637	T12018	T12019	nsubjpass	tree,colored
R7638	T12020	T12019	auxpass	was,colored
R7639	T12021	T12019	advcl	using,colored
R764	T1666	T1650	parataxis	15,expressed
R7640	T12022	T12023	det	a,script
R7641	T12023	T12021	dobj	script,using
R7642	T12024	T12023	compound	custom,script
R7643	T12025	T12019	punct	.,colored
R7644	T12027	T12028	compound	Information,content
R7645	T12028	T12029	nsubjpass	content,calculated
R7646	T12030	T12028	punct	(,content
R7647	T12031	T12032	det	the,measure
R7648	T12032	T12028	appos	measure,content
R7649	T12033	T12032	prep	of,measure
R765	T1667	T1666	nummod	14,15
R7650	T12034	T12035	compound	sequence,conservation
R7651	T12035	T12033	pobj	conservation,of
R7652	T12036	T12035	acl	shown,conservation
R7653	T12037	T12036	prep	in,shown
R7654	T12038	T12037	pobj	Figure,in
R7655	T12039	T12038	nummod	6,Figure
R7656	T12040	T12029	punct	),calculated
R7657	T12041	T12029	auxpass	was,calculated
R7658	T12042	T12029	prep	for,calculated
R7659	T12043	T12044	det	each,position
R766	T1668	T1666	punct	",",15
R7660	T12044	T12042	pobj	position,for
R7661	T12045	T12044	prep	in,position
R7662	T12046	T12047	det	the,alignment
R7663	T12047	T12045	pobj	alignment,in
R7664	T12048	T12029	advcl	using,calculated
R7665	T12049	T12048	dobj	alpro,using
R7666	T12050	T12051	punct	[,51
R7667	T12051	T12029	parataxis	51,calculated
R7668	T12052	T12051	punct	],51
R7669	T12053	T12029	punct	.,calculated
R767	T1669	T1666	punct	],15
R7670	T12055	T12056	aux	To,determine
R7671	T12056	T12057	advcl	determine,examined
R7672	T12058	T12059	det	the,number
R7673	T12059	T12056	dobj	number,determine
R7674	T12060	T12059	prep	of,number
R7675	T12061	T12062	amod	transcriptional,isoforms
R7676	T12062	T12060	pobj	isoforms,of
R7677	T12063	T12056	prep	for,determine
R7678	T12064	T12065	det	each,gene
R7679	T12065	T12063	pobj	gene,for
R768	T1670	T1650	punct	.,expressed
R7680	T12066	T12057	punct	", ",examined
R7681	T12067	T12057	nsubj	we,examined
R7682	T12068	T12069	det	the,output
R7683	T12069	T12057	dobj	output,examined
R7684	T12070	T12069	compound	sim4,output
R7685	T12071	T12057	prep	for,examined
R7686	T12072	T12073	det	every,cDNA
R7687	T12073	T12071	pobj	cDNA,for
R7688	T12074	T12073	amod	matching,cDNA
R7689	T12075	T12057	prep	in,examined
R769	T1672	T1673	advmod	Furthermore,expresses
R7690	T12076	T12077	amod	decreasing,order
R7691	T12077	T12075	pobj	order,in
R7692	T12078	T12077	prep	of,order
R7693	T12079	T12078	pobj	number,of
R7694	T12080	T12079	prep	of,number
R7695	T12081	T12080	pobj	exons,of
R7696	T12082	T12057	punct	.,examined
R7697	T12084	T12085	det	The,cDNA
R7698	T12085	T12087	nsubjpass	cDNA,counted
R7699	T12086	T12085	amod	first,cDNA
R77	T381	T380	acl	suggested,differences
R770	T1674	T1673	punct	", ",expresses
R7700	T12088	T12087	auxpass	was,counted
R7701	T12089	T12087	prep	as,counted
R7702	T12090	T12091	nummod	one,form
R7703	T12091	T12089	pobj	form,as
R7704	T12092	T12091	compound	splice,form
R7705	T12093	T12087	punct	", ",counted
R7706	T12094	T12087	cc	and,counted
R7707	T12095	T12096	prep	for,determined
R7708	T12096	T12087	conj	determined,counted
R7709	T12097	T12098	det	each,cDNA
R771	T1675	T1676	det	each,neuron
R7710	T12098	T12095	pobj	cDNA,for
R7711	T12099	T12098	amod	subsequent,cDNA
R7712	T12100	T12096	punct	", ",determined
R7713	T12101	T12096	nsubj	we,determined
R7714	T12102	T12103	mark	whether,was
R7715	T12103	T12096	ccomp	was,determined
R7716	T12104	T12105	compound	exon,structure
R7717	T12105	T12103	nsubj	structure,was
R7718	T12106	T12107	advmod	mutually,exclusive
R7719	T12107	T12103	acomp	exclusive,was
R772	T1676	T1673	nsubj	neuron,expresses
R7720	T12108	T12107	prep	to,exclusive
R7721	T12109	T12108	pobj	isoforms,to
R7722	T12110	T12111	advmod	already,counted
R7723	T12111	T12109	acl	counted,isoforms
R7724	T12112	T12096	punct	.,determined
R7725	T12114	T12115	nsubj	We,were
R7726	T12116	T12115	acomp	conservative,were
R7727	T12117	T12115	prep	in,were
R7728	T12118	T12119	poss	our,definition
R7729	T12119	T12117	pobj	definition,in
R773	T1677	T1676	amod	olfactory,neuron
R7730	T12120	T12119	prep	of,definition
R7731	T12121	T12122	advmod	mutually,exclusive
R7732	T12122	T12120	pobj	exclusive,of
R7733	T12123	T12115	punct	", ",were
R7734	T12124	T12115	cc	and,were
R7735	T12125	T12126	advmod	thus,represents
R7736	T12126	T12115	conj	represents,were
R7737	T12127	T12128	poss	our,count
R7738	T12128	T12126	nsubj	count,represents
R7739	T12129	T12130	det	the,number
R774	T1678	T1679	advmod	only,allele
R7740	T12130	T12126	dobj	number,represents
R7741	T12131	T12130	amod	minimum,number
R7742	T12132	T12130	prep	of,number
R7743	T12133	T12132	pobj	isoforms,of
R7744	T12134	T12133	acl	represented,isoforms
R7745	T12135	T12134	prep	in,represented
R7746	T12136	T12137	det	the,collection
R7747	T12137	T12135	pobj	collection,in
R7748	T12138	T12137	compound	cDNA,collection
R7749	T12139	T12126	punct	.,represents
R775	T1679	T1673	dobj	allele,expresses
R7750	T12269	T12270	compound	RT,PCR
R7751	T12271	T12270	punct	-,PCR
R7752	T12273	T12274	det	The,epithelia
R7753	T12274	T12276	nsubjpass	epithelia,dissected
R7754	T12275	T12274	amod	olfactory,epithelia
R7755	T12277	T12276	auxpass	were,dissected
R7756	T12278	T12276	prep	from,dissected
R7757	T12279	T12280	nummod	three,mice
R7758	T12280	T12278	pobj	mice,from
R7759	T12281	T12280	amod	adult,mice
R776	T1680	T1679	nummod	one,allele
R7760	T12282	T12280	amod	female,mice
R7761	T12283	T12280	nmod	C57BL,mice
R7762	T12284	T12283	punct	/,C57BL
R7763	T12285	T12283	nummod	6,C57BL
R7764	T12286	T12276	punct	", ",dissected
R7765	T12287	T12276	prep	including,dissected
R7766	T12288	T12287	pobj	tissues,including
R7767	T12289	T12288	acl	attached,tissues
R7768	T12290	T12289	prep	to,attached
R7769	T12291	T12292	det	the,skull
R777	T1681	T1682	punct	[,16
R7770	T12292	T12290	pobj	skull,to
R7771	T12293	T12292	cc	and,skull
R7772	T12294	T12292	conj	septum,skull
R7773	T12295	T12276	punct	.,dissected
R7774	T12297	T12298	nsubjpass	RNA,isolated
R7775	T12299	T12298	auxpass	was,isolated
R7776	T12300	T12298	advcl	using,isolated
R7777	T12301	T12302	det	the,kit
R7778	T12302	T12300	dobj	kit,using
R7779	T12303	T12302	compound	Qiagen,kit
R778	T1682	T1679	parataxis	16,allele
R7780	T12304	T12302	compound	RNeasy,kit
R7781	T12305	T12302	compound	midi,kit
R7782	T12306	T12307	punct	(,Qiagen
R7783	T12307	T12302	parataxis	Qiagen,kit
R7784	T12308	T12307	punct	", ",Qiagen
R7785	T12309	T12307	npadvmod	Valencia,Qiagen
R7786	T12310	T12307	punct	", ",Qiagen
R7787	T12311	T12307	npadvmod	CA,Qiagen
R7788	T12312	T12307	punct	", ",Qiagen
R7789	T12313	T12307	npadvmod	USA,Qiagen
R779	T1683	T1682	punct	],16
R7790	T12314	T12307	punct	),Qiagen
R7791	T12315	T12300	punct	", ",using
R7792	T12316	T12300	prep	including,using
R7793	T12317	T12318	det	a,step
R7794	T12318	T12316	pobj	step,including
R7795	T12319	T12318	compound	DNase,step
R7796	T12320	T12318	compound	treatment,step
R7797	T12321	T12298	punct	.,isolated
R7798	T12323	T12324	amod	First,strand
R7799	T12324	T12326	compound	strand,cDNA
R78	T382	T381	agent	by,suggested
R780	T1684	T1679	prep	of,allele
R7800	T12325	T12324	punct	-,strand
R7801	T12326	T12327	nsubjpass	cDNA,produced
R7802	T12328	T12327	auxpass	was,produced
R7803	T12329	T12327	prep	from,produced
R7804	T12330	T12331	nummod	2.5,μg
R7805	T12331	T12329	pobj	μg,from
R7806	T12332	T12331	prep	of,μg
R7807	T12333	T12332	pobj	RNA,of
R7808	T12334	T12331	prep	in,μg
R7809	T12335	T12336	det	a,volume
R781	T1685	T1686	det	a,gene
R7810	T12336	T12334	pobj	volume,in
R7811	T12337	T12336	prep	of,volume
R7812	T12338	T12339	nummod	50,μl
R7813	T12339	T12337	pobj	μl,of
R7814	T12340	T12327	advcl	using,produced
R7815	T12341	T12342	amod	random,hexamers
R7816	T12342	T12340	dobj	hexamers,using
R7817	T12343	T12342	cc	and,hexamers
R7818	T12344	T12345	poss	Invitrogen,transcriptase
R7819	T12345	T12342	conj	transcriptase,hexamers
R782	T1686	T1684	pobj	gene,of
R7820	T12346	T12344	case	's,Invitrogen
R7821	T12347	T12345	nmod	Superscript,transcriptase
R7822	T12348	T12347	nummod	II,Superscript
R7823	T12349	T12345	amod	reverse,transcriptase
R7824	T12350	T12351	punct	(,Invitrogen
R7825	T12351	T12345	parataxis	Invitrogen,transcriptase
R7826	T12352	T12351	punct	", ",Invitrogen
R7827	T12353	T12351	npadvmod	Carlsbad,Invitrogen
R7828	T12354	T12351	punct	", ",Invitrogen
R7829	T12355	T12351	npadvmod	CA,Invitrogen
R783	T1687	T1686	amod	single,gene
R7830	T12356	T12351	punct	", ",Invitrogen
R7831	T12357	T12351	npadvmod	USA,Invitrogen
R7832	T12358	T12351	punct	),Invitrogen
R7833	T12359	T12327	punct	", ",produced
R7834	T12360	T12327	prep	according,produced
R7835	T12361	T12360	prep	to,according
R7836	T12362	T12363	det	the,manufacturer
R7837	T12363	T12364	poss	manufacturer,recommendations
R7838	T12364	T12361	pobj	recommendations,to
R7839	T12365	T12363	case	's,manufacturer
R784	T1688	T1689	amod	olfactory,receptor
R7840	T12366	T12327	punct	.,produced
R7841	T12368	T12369	nummod	One,fifth
R7842	T12369	T12373	nsubjpass	fifth,used
R7843	T12370	T12369	punct	-,fifth
R7844	T12371	T12369	nummod	twenty,fifth
R7845	T12372	T12369	punct	-,fifth
R7846	T12374	T12369	prep	of,fifth
R7847	T12375	T12376	det	the,cDNA
R7848	T12376	T12374	pobj	cDNA,of
R7849	T12377	T12376	amod	resulting,cDNA
R785	T1689	T1686	compound	receptor,gene
R7850	T12378	T12373	auxpass	was,used
R7851	T12379	T12373	prep	as,used
R7852	T12380	T12379	pobj	template,as
R7853	T12381	T12373	prep	in,used
R7854	T12382	T12383	amod	subsequent,reactions
R7855	T12383	T12381	pobj	reactions,in
R7856	T12384	T12383	compound	PCR,reactions
R7857	T12385	T12373	punct	.,used
R7858	T12387	T12388	compound	PCR,amplification
R7859	T12388	T12389	nsubjpass	amplification,performed
R786	T1690	T1691	punct	[,18
R7860	T12390	T12388	amod	biased,amplification
R7861	T12391	T12390	prep	towards,biased
R7862	T12392	T12393	nmod	class,receptors
R7863	T12393	T12391	pobj	receptors,towards
R7864	T12394	T12392	nummod	I,class
R7865	T12395	T12393	amod	olfactory,receptors
R7866	T12396	T12389	auxpass	was,performed
R7867	T12397	T12389	advcl	using,performed
R7868	T12398	T12399	amod	degenerate,P26
R7869	T12399	T12397	dobj	P26,using
R787	T1691	T1686	parataxis	18,gene
R7870	T12400	T12399	compound	primers,P26
R7871	T12401	T12402	punct	[,17
R7872	T12402	T12399	parataxis	17,P26
R7873	T12403	T12402	punct	],17
R7874	T12404	T12399	cc	and,P26
R7875	T12405	T12399	conj	classI_R1,P26
R7876	T12406	T12407	punct	(,GGRTTIADIRYIGGNGG
R7877	T12407	T12405	parataxis	GGRTTIADIRYIGGNGG,classI_R1
R7878	T12408	T12407	nummod	5,GGRTTIADIRYIGGNGG
R7879	T12409	T12408	punct	',5
R788	T1692	T1691	nummod	17,18
R7880	T12410	T12407	punct	-,GGRTTIADIRYIGGNGG
R7881	T12411	T12407	punct	-,GGRTTIADIRYIGGNGG
R7882	T12412	T12407	nummod	3,GGRTTIADIRYIGGNGG
R7883	T12413	T12407	punct	',GGRTTIADIRYIGGNGG
R7884	T12414	T12407	punct	),GGRTTIADIRYIGGNGG
R7885	T12415	T12399	prep	with,P26
R7886	T12416	T12417	det	an,temperature
R7887	T12417	T12415	pobj	temperature,with
R7888	T12418	T12417	amod	annealing,temperature
R7889	T12419	T12417	prep	of,temperature
R789	T1693	T1691	punct	",",18
R7890	T12420	T12421	nummod	44,°C
R7891	T12421	T12419	pobj	°C,of
R7892	T12422	T12389	punct	.,performed
R7893	T12424	T12425	det	The,product
R7894	T12425	T12426	nsubjpass	product,cloned
R7895	T12427	T12426	auxpass	was,cloned
R7896	T12428	T12429	punct	(,Invitrogen
R7897	T12429	T12426	parataxis	Invitrogen,cloned
R7898	T12430	T12431	compound	TA,kit
R7899	T12431	T12429	dep	kit,Invitrogen
R79	T383	T384	poss	our,screen
R790	T1694	T1691	punct	],18
R7900	T12432	T12431	compound	cloning,kit
R7901	T12433	T12429	punct	", ",Invitrogen
R7902	T12434	T12429	punct	),Invitrogen
R7903	T12435	T12426	punct	", ",cloned
R7904	T12436	T12426	cc	and,cloned
R7905	T12437	T12438	amod	individual,clones
R7906	T12438	T12439	nsubjpass	clones,sequenced
R7907	T12439	T12426	conj	sequenced,cloned
R7908	T12440	T12439	auxpass	were,sequenced
R7909	T12441	T12439	punct	.,sequenced
R791	T1695	T1673	punct	", ",expresses
R7910	T12443	T12444	amod	Specific,primers
R7911	T12444	T12446	nsubjpass	primers,given
R7912	T12445	T12444	compound	PCR,primers
R7913	T12447	T12444	acl	used,primers
R7914	T12448	T12449	aux	to,confirm
R7915	T12449	T12447	advcl	confirm,used
R7916	T12450	T12449	dobj	expression,confirm
R7917	T12451	T12450	prep	of,expression
R7918	T12452	T12453	amod	individual,genes
R7919	T12453	T12451	pobj	genes,of
R792	T1696	T1673	cc	and,expresses
R7920	T12454	T12455	amod	olfactory,receptor
R7921	T12455	T12453	compound	receptor,genes
R7922	T12456	T12446	auxpass	are,given
R7923	T12457	T12446	prep	in,given
R7924	T12458	T12459	amod	Additional,file
R7925	T12459	T12457	pobj	file,in
R7926	T12460	T12459	compound	data,file
R7927	T12461	T12459	nummod	1,file
R7928	T12462	T12446	punct	.,given
R7929	T12464	T12465	det	Each,product
R793	T1697	T1698	det	the,genes
R7930	T12465	T12467	nsubjpass	product,sequenced
R7931	T12466	T12465	compound	PCR,product
R7932	T12468	T12467	auxpass	was,sequenced
R7933	T12469	T12470	aux	to,confirm
R7934	T12470	T12467	advcl	confirm,sequenced
R7935	T12471	T12472	mark	that,amplified
R7936	T12472	T12470	ccomp	amplified,confirm
R7937	T12473	T12474	det	the,gene
R7938	T12474	T12472	nsubjpass	gene,amplified
R7939	T12475	T12474	amod	expected,gene
R794	T1698	T1702	nsubj	genes,are
R7940	T12476	T12474	cc	and,gene
R7941	T12477	T12478	det	no,others
R7942	T12478	T12474	conj	others,gene
R7943	T12479	T12472	aux	had,amplified
R7944	T12480	T12472	auxpass	been,amplified
R7945	T12481	T12467	punct	.,sequenced
R7946	T12483	T12484	compound	Control,reactions
R7947	T12484	T12485	nsubj	reactions,gave
R7948	T12486	T12484	prep	on,reactions
R7949	T12487	T12488	det	a,template
R795	T1699	T1698	nmod	remaining,genes
R7950	T12488	T12486	pobj	template,on
R7951	T12489	T12488	acl	made,template
R7952	T12490	T12489	prep	by,made
R7953	T12491	T12490	pcomp	omitting,by
R7954	T12492	T12493	amod	reverse,transcriptase
R7955	T12493	T12491	dobj	transcriptase,omitting
R7956	T12494	T12495	det	no,product
R7957	T12495	T12485	dobj	product,gave
R7958	T12496	T12485	punct	", ",gave
R7959	T12497	T12485	advcl	indicating,gave
R796	T1700	T1701	advmod	approximately,"1,499"
R7960	T12498	T12499	mark	that,was
R7961	T12499	T12497	ccomp	was,indicating
R7962	T12500	T12501	det	the,preparation
R7963	T12501	T12499	nsubj	preparation,was
R7964	T12502	T12501	compound	RNA,preparation
R7965	T12503	T12499	acomp	uncontaminated,was
R7966	T12504	T12503	prep	by,uncontaminated
R7967	T12505	T12506	amod	genomic,DNA
R7968	T12506	T12504	pobj	DNA,by
R7969	T12507	T12485	punct	.,gave
R797	T1701	T1698	nummod	"1,499",genes
R7970	T12509	T12510	amod	Relative,levels
R7971	T12510	T12512	nsubjpass	levels,estimated
R7972	T12511	T12510	compound	transcript,levels
R7973	T12513	T12512	auxpass	were,estimated
R7974	T12514	T12512	advcl	using,estimated
R7975	T12515	T12516	amod	real,time
R7976	T12516	T12518	compound	time,PCR
R7977	T12517	T12516	punct	-,time
R7978	T12518	T12514	dobj	PCR,using
R7979	T12519	T12514	prep	according,using
R798	T1702	T1673	conj	are,expresses
R7980	T12520	T12519	prep	to,according
R7981	T12521	T12522	compound	Applied,Biosystems
R7982	T12522	T12523	poss	Biosystems,protocols
R7983	T12523	T12520	pobj	protocols,to
R7984	T12524	T12522	case	',Biosystems
R7985	T12525	T12514	punct	", ",using
R7986	T12526	T12527	mark	with,given
R7987	T12527	T12514	advcl	given,using
R7988	T12528	T12529	compound	magnesium,concentration
R7989	T12529	T12527	nsubj	concentration,given
R799	T1703	T1704	advmod	transcriptionally,inactive
R7990	T12530	T12529	punct	", ",concentration
R7991	T12531	T12532	compound	primer,pair
R7992	T12532	T12529	conj	pair,concentration
R7993	T12533	T12532	cc	and,pair
R7994	T12534	T12535	amod	fluorescent,probe
R7995	T12535	T12532	conj	probe,pair
R7996	T12536	T12527	prep	in,given
R7997	T12537	T12538	amod	Additional,file
R7998	T12538	T12536	pobj	file,in
R7999	T12539	T12538	compound	data,file
R8	T304	T302	dobj	function,confirming
R80	T384	T382	pobj	screen,by
R800	T1704	T1702	acomp	inactive,are
R8000	T12540	T12538	nummod	2,file
R8001	T12541	T12512	punct	.,estimated
R8002	T12543	T12544	det	The,increase
R8003	T12544	T12545	nsubjpass	increase,measured
R8004	T12546	T12544	prep	in,increase
R8005	T12547	T12546	pobj	fluorescence,in
R8006	T12548	T12544	prep	during,increase
R8007	T12549	T12548	pobj	thermocycling,during
R8008	T12550	T12545	auxpass	was,measured
R8009	T12551	T12545	prep	on,measured
R801	T1705	T1673	punct	.,expresses
R8010	T12552	T12553	det	an,7900HT
R8011	T12553	T12551	pobj	7900HT,on
R8012	T12554	T12553	compound	ABI,7900HT
R8013	T12555	T12553	compound	PRISM,7900HT
R8014	T12556	T12545	punct	.,measured
R8015	T12558	T12559	amod	Standard,curves
R8016	T12559	T12560	nsubjpass	curves,constructed
R8017	T12561	T12560	auxpass	were,constructed
R8018	T12562	T12560	prep	for,constructed
R8019	T12563	T12564	det	each,pair
R802	T1707	T1708	mark	While,is
R8020	T12564	T12562	pobj	pair,for
R8021	T12565	T12564	compound	primer,pair
R8022	T12566	T12560	advcl	using,constructed
R8023	T12567	T12568	compound	triplicate,samples
R8024	T12568	T12566	dobj	samples,using
R8025	T12569	T12568	prep	of,samples
R8026	T12570	T12571	nmod	mouse,DNA
R8027	T12571	T12569	pobj	DNA,of
R8028	T12572	T12571	amod	genomic,DNA
R8029	T12573	T12571	prep	of,DNA
R803	T1708	T1714	advcl	is,proposed
R8030	T12574	T12575	nummod	nine,concentrations
R8031	T12575	T12573	pobj	concentrations,of
R8032	T12576	T12575	amod	known,concentrations
R8033	T12577	T12578	punct	(,ng
R8034	T12578	T12575	parataxis	ng,concentrations
R8035	T12579	T12578	dep	range,ng
R8036	T12580	T12581	quantmod	0.01,100
R8037	T12581	T12578	nummod	100,ng
R8038	T12582	T12581	punct	-,100
R8039	T12583	T12578	punct	", ",ng
R804	T1709	T1710	det	the,mechanism
R8040	T12584	T12578	cc	or,ng
R8041	T12585	T12586	advmod	about,"30,000"
R8042	T12586	T12589	nummod	"30,000",copies
R8043	T12587	T12586	quantmod	3,"30,000"
R8044	T12588	T12586	punct	-,"30,000"
R8045	T12589	T12578	conj	copies,ng
R8046	T12590	T12589	prep	of,copies
R8047	T12591	T12592	det	the,genome
R8048	T12592	T12590	pobj	genome,of
R8049	T12593	T12592	compound	haploid,genome
R805	T1710	T1708	nsubj	mechanism,is
R8050	T12594	T12578	punct	),ng
R8051	T12595	T12560	punct	.,constructed
R8052	T12597	T12598	amod	Relative,level
R8053	T12598	T12600	nsubjpass	level,determined
R8054	T12599	T12598	compound	expression,level
R8055	T12601	T12598	prep	of,level
R8056	T12602	T12603	det	each,gene
R8057	T12603	T12601	pobj	gene,of
R8058	T12604	T12600	auxpass	was,determined
R8059	T12605	T12600	prep	by,determined
R806	T1711	T1710	acl	ensuring,mechanism
R8060	T12606	T12605	pcomp	comparing,by
R8061	T12607	T12608	det	the,Ct
R8062	T12608	T12606	dobj	Ct,comparing
R8063	T12609	T12608	amod	mean,Ct
R8064	T12610	T12608	punct	(,Ct
R8065	T12611	T12608	appos	cycle,Ct
R8066	T12612	T12613	advmod	where,reaches
R8067	T12613	T12611	relcl	reaches,cycle
R8068	T12614	T12613	nsubj	fluorescence,reaches
R8069	T12615	T12616	det	an,value
R807	T1712	T1713	amod	singular,expression
R8070	T12616	T12613	dobj	value,reaches
R8071	T12617	T12616	amod	arbitrary,value
R8072	T12618	T12616	compound	threshold,value
R8073	T12619	T12608	punct	),Ct
R8074	T12620	T12608	acl	obtained,Ct
R8075	T12621	T12620	prep	with,obtained
R8076	T12622	T12623	nummod	six,samples
R8077	T12623	T12621	pobj	samples,with
R8078	T12624	T12623	compound	replicate,samples
R8079	T12625	T12623	prep	of,samples
R808	T1713	T1711	dobj	expression,ensuring
R8080	T12626	T12627	amod	reverse,transcribed
R8081	T12627	T12629	amod	transcribed,RNA
R8082	T12628	T12627	punct	-,transcribed
R8083	T12629	T12625	pobj	RNA,of
R8084	T12630	T12606	prep	to,comparing
R8085	T12631	T12632	det	the,curve
R8086	T12632	T12630	pobj	curve,to
R8087	T12633	T12632	amod	standard,curve
R8088	T12634	T12632	prep	for,curve
R8089	T12635	T12636	det	the,primers
R809	T1715	T1708	acomp	unknown,is
R8090	T12636	T12634	pobj	primers,for
R8091	T12637	T12636	amod	corresponding,primers
R8092	T12638	T12600	punct	.,determined
R8093	T12640	T12641	amod	Relative,levels
R8094	T12641	T12643	nsubjpass	levels,measured
R8095	T12642	T12641	compound	RNA,levels
R8096	T12644	T12641	prep	of,levels
R8097	T12645	T12646	det	a,gene
R8098	T12646	T12644	pobj	gene,of
R8099	T12647	T12646	compound	housekeeping,gene
R81	T385	T371	punct	", ",find
R810	T1716	T1714	punct	", ",proposed
R8100	T12648	T12646	punct	", ",gene
R8101	T12649	T12650	amod	ribosomal,S16
R8102	T12650	T12646	appos	S16,gene
R8103	T12651	T12643	punct	", ",measured
R8104	T12652	T12643	auxpass	were,measured
R8105	T12653	T12654	mark	as,described
R8106	T12654	T12643	advcl	described,measured
R8107	T12655	T12654	advmod	previously,described
R8108	T12656	T12657	punct	[,52
R8109	T12657	T12654	parataxis	52,described
R811	T1717	T1718	amod	many,hypotheses
R8110	T12658	T12657	punct	],52
R8111	T12659	T12643	punct	.,measured
R8112	T12661	T12662	compound	Control,reactions
R8113	T12662	T12663	nsubj	reactions,amplified
R8114	T12664	T12662	prep	on,reactions
R8115	T12665	T12664	pobj	template,on
R8116	T12666	T12665	acl	prepared,template
R8117	T12667	T12666	prep	by,prepared
R8118	T12668	T12667	pcomp	omitting,by
R8119	T12669	T12670	amod	reverse,transcriptase
R812	T1718	T1714	nsubjpass	hypotheses,proposed
R8120	T12670	T12668	dobj	transcriptase,omitting
R8121	T12671	T12663	prep	at,amplified
R8122	T12672	T12673	det	a,level
R8123	T12673	T12671	pobj	level,at
R8124	T12674	T12673	amod	relative,level
R8125	T12675	T12673	prep	of,level
R8126	T12676	T12677	quantmod	0.03,0.01
R8127	T12677	T12679	nummod	0.01,ng
R8128	T12678	T12677	punct	±,0.01
R8129	T12679	T12675	pobj	ng,of
R813	T1719	T1714	aux	have,proposed
R8130	T12680	T12679	cc	or,ng
R8131	T12681	T12679	conj	less,ng
R8132	T12682	T12663	prep	in,amplified
R8133	T12683	T12684	det	each,case
R8134	T12684	T12682	pobj	case,in
R8135	T12685	T12663	punct	.,amplified
R8136	T12687	T12688	compound	Expression,measurements
R8137	T12688	T12689	nsubjpass	measurements,normalized
R8138	T12690	T12688	prep	of,measurements
R8139	T12691	T12692	det	the,genes
R814	T1720	T1714	auxpass	been,proposed
R8140	T12692	T12690	pobj	genes,of
R8141	T12693	T12692	nummod	seven,genes
R8142	T12694	T12689	auxpass	were,normalized
R8143	T12695	T12689	prep	for,normalized
R8144	T12696	T12697	det	each,mouse
R8145	T12697	T12695	pobj	mouse,for
R8146	T12698	T12699	mark	so,were
R8147	T12699	T12689	advcl	were,normalized
R8148	T12700	T12699	mark	that,were
R8149	T12701	T12702	compound	S16,levels
R815	T1721	T1722	punct	[,19
R8150	T12702	T12699	nsubj	levels,were
R8151	T12703	T12699	acomp	equal,were
R8152	T12704	T12703	prep	to,equal
R8153	T12705	T12704	pobj	1,to
R8154	T12706	T12707	punct	(,units
R8155	T12707	T12705	parataxis	units,1
R8156	T12708	T12707	amod	arbitrary,units
R8157	T12709	T12707	punct	),units
R8158	T12710	T12689	punct	.,normalized
R8159	T12805	T12806	advmod	In,situ
R816	T1722	T1714	parataxis	19,proposed
R8160	T12806	T12807	amod	situ,hybridization
R8161	T12809	T12810	amod	Coronal,sections
R8162	T12810	T12811	nsubjpass	sections,cut
R8163	T12812	T12811	auxpass	were,cut
R8164	T12813	T12811	prep	from,cut
R8165	T12814	T12815	det	the,epithelia
R8166	T12815	T12813	pobj	epithelia,from
R8167	T12816	T12815	amod	olfactory,epithelia
R8168	T12817	T12815	prep	of,epithelia
R8169	T12818	T12819	det	an,mouse
R817	T1723	T1722	nummod	14,19
R8170	T12819	T12817	pobj	mouse,of
R8171	T12820	T12819	amod	adult,mouse
R8172	T12821	T12822	punct	(,Figure
R8173	T12822	T12819	parataxis	Figure,mouse
R8174	T12823	T12822	nummod	4,Figure
R8175	T12824	T12822	punct	),Figure
R8176	T12825	T12819	cc	and,mouse
R8177	T12826	T12827	det	a,mouse
R8178	T12827	T12819	conj	mouse,mouse
R8179	T12828	T12827	amod	young,mouse
R818	T1724	T1722	punct	",",19
R8180	T12829	T12830	punct	(,P6
R8181	T12830	T12828	parataxis	P6,young
R8182	T12831	T12830	punct	),P6
R8183	T12832	T12827	nmod	C57BL,mouse
R8184	T12833	T12832	punct	/,C57BL
R8185	T12834	T12832	nummod	6,C57BL
R8186	T12835	T12811	punct	.,cut
R8187	T12837	T12838	nmod	RNA,hybridization
R8188	T12838	T12841	nsubjpass	hybridization,carried
R8189	T12839	T12840	advmod	in,situ
R819	T1725	T1722	nummod	16,19
R8190	T12840	T12838	amod	situ,hybridization
R8191	T12842	T12841	auxpass	was,carried
R8192	T12843	T12841	prt	out,carried
R8193	T12844	T12845	mark	as,described
R8194	T12845	T12841	advcl	described,carried
R8195	T12846	T12845	advmod	previously,described
R8196	T12847	T12848	punct	[,53
R8197	T12848	T12845	parataxis	53,described
R8198	T12849	T12848	nummod	15,53
R8199	T12850	T12848	punct	",",53
R82	T386	T371	nsubj	we,find
R820	T1726	T1722	punct	",",19
R8200	T12851	T12848	punct	],53
R8201	T12852	T12841	prep	with,carried
R8202	T12853	T12854	npadvmod	digoxigenin,labeled
R8203	T12854	T12856	amod	labeled,riboprobes
R8204	T12855	T12854	punct	-,labeled
R8205	T12856	T12852	pobj	riboprobes,with
R8206	T12857	T12856	amod	antisense,riboprobes
R8207	T12858	T12856	amod	specific,riboprobes
R8208	T12859	T12858	prep	for,specific
R8209	T12860	T12861	det	the,UTRs
R821	T1727	T1722	punct	],19
R8210	T12861	T12859	pobj	UTRs,for
R8211	T12862	T12861	nummod	3,UTRs
R8212	T12863	T12862	punct	',3
R8213	T12864	T12861	prep	of,UTRs
R8214	T12865	T12866	compound	genes,AY318555
R8215	T12866	T12864	pobj	AY318555,of
R8216	T12867	T12868	punct	(,kb
R8217	T12868	T12866	parataxis	kb,AY318555
R8218	T12869	T12868	nummod	0.5,kb
R8219	T12870	T12868	punct	),kb
R822	T1728	T1714	punct	.,proposed
R8220	T12871	T12866	cc	and,AY318555
R8221	T12872	T12866	conj	AY317365,AY318555
R8222	T12873	T12874	punct	(,kb
R8223	T12874	T12872	parataxis	kb,AY317365
R8224	T12875	T12874	nummod	0.5,kb
R8225	T12876	T12874	punct	),kb
R8226	T12877	T12841	punct	.,carried
R8227	T12879	T12880	compound	Riboprobe,sequences
R8228	T12880	T12881	nsubjpass	sequences,generated
R8229	T12882	T12881	auxpass	were,generated
R823	T1730	T1731	prep	In,bring
R8230	T12883	T12881	prep	by,generated
R8231	T12884	T12883	pobj	PCR,by
R8232	T12885	T12881	advcl	using,generated
R8233	T12886	T12887	compound	primer,pairs
R8234	T12887	T12885	dobj	pairs,using
R8235	T12888	T12889	nummod	5,TCTTCCAAACCTGGACCCCCC
R8236	T12889	T12887	appos	TCTTCCAAACCTGGACCCCCC,pairs
R8237	T12890	T12888	punct	',5
R8238	T12891	T12889	punct	-,TCTTCCAAACCTGGACCCCCC
R8239	T12892	T12889	punct	-,TCTTCCAAACCTGGACCCCCC
R824	T1732	T1733	nummod	one,model
R8240	T12893	T12889	nummod	3,TCTTCCAAACCTGGACCCCCC
R8241	T12894	T12889	punct	',TCTTCCAAACCTGGACCCCCC
R8242	T12895	T12889	cc	and,TCTTCCAAACCTGGACCCCCC
R8243	T12896	T12897	nummod	5,ATCTCTCCAGCACCTTACTTG
R8244	T12897	T12889	conj	ATCTCTCCAGCACCTTACTTG,TCTTCCAAACCTGGACCCCCC
R8245	T12898	T12896	punct	',5
R8246	T12899	T12897	punct	-,ATCTCTCCAGCACCTTACTTG
R8247	T12900	T12897	punct	-,ATCTCTCCAGCACCTTACTTG
R8248	T12901	T12897	nummod	3,ATCTCTCCAGCACCTTACTTG
R8249	T12902	T12897	punct	',ATCTCTCCAGCACCTTACTTG
R825	T1733	T1730	pobj	model,In
R8250	T12903	T12885	prep	for,using
R8251	T12904	T12903	pobj	AY318555,for
R8252	T12905	T12885	cc	and,using
R8253	T12906	T12907	compound	primer,pairs
R8254	T12907	T12885	conj	pairs,using
R8255	T12908	T12909	nummod	5,TAAGATGTAAGTGATAATTTAGATTACAGG
R8256	T12909	T12907	appos	TAAGATGTAAGTGATAATTTAGATTACAGG,pairs
R8257	T12910	T12908	punct	',5
R8258	T12911	T12909	punct	-,TAAGATGTAAGTGATAATTTAGATTACAGG
R8259	T12912	T12909	punct	-,TAAGATGTAAGTGATAATTTAGATTACAGG
R826	T1734	T1731	punct	", ",bring
R8260	T12913	T12909	nummod	3,TAAGATGTAAGTGATAATTTAGATTACAGG
R8261	T12914	T12909	punct	',TAAGATGTAAGTGATAATTTAGATTACAGG
R8262	T12915	T12909	cc	and,TAAGATGTAAGTGATAATTTAGATTACAGG
R8263	T12916	T12917	nummod	5,TTTCTGCCTCAGCTATGACAG
R8264	T12917	T12909	conj	TTTCTGCCTCAGCTATGACAG,TAAGATGTAAGTGATAATTTAGATTACAGG
R8265	T12918	T12916	punct	',5
R8266	T12919	T12917	punct	-,TTTCTGCCTCAGCTATGACAG
R8267	T12920	T12917	punct	-,TTTCTGCCTCAGCTATGACAG
R8268	T12921	T12917	nummod	3,TTTCTGCCTCAGCTATGACAG
R8269	T12922	T12917	punct	',TTTCTGCCTCAGCTATGACAG
R827	T1735	T1736	amod	somatic,recombination
R8270	T12923	T12907	prep	for,pairs
R8271	T12924	T12923	pobj	AY317365,for
R8272	T12925	T12881	punct	.,generated
R8273	T12927	T12928	nsubjpass	Hybridization,carried
R8274	T12929	T12928	auxpass	was,carried
R8275	T12930	T12928	prt	out,carried
R8276	T12931	T12928	prep	in,carried
R8277	T12932	T12933	nummod	50,%
R8278	T12933	T12934	compound	%,formamide
R8279	T12934	T12931	pobj	formamide,in
R828	T1736	T1731	nsubj	recombination,bring
R8280	T12935	T12928	prep	at,carried
R8281	T12936	T12937	nummod	65,°C
R8282	T12937	T12935	pobj	°C,at
R8283	T12938	T12928	punct	", ",carried
R8284	T12939	T12928	cc	and,carried
R8285	T12940	T12941	nsubjpass	slides,washed
R8286	T12941	T12928	conj	washed,carried
R8287	T12942	T12941	auxpass	were,washed
R8288	T12943	T12941	prep	at,washed
R8289	T12944	T12945	amod	high,stringency
R829	T1737	T1736	compound	DNA,recombination
R8290	T12945	T12943	pobj	stringency,at
R8291	T12946	T12947	punct	(,°C
R8292	T12947	T12945	parataxis	°C,stringency
R8293	T12948	T12947	nummod	65,°C
R8294	T12949	T12947	punct	", ",°C
R8295	T12950	T12951	nsubj	0.2,SSC
R8296	T12951	T12947	ccomp	SSC,°C
R8297	T12952	T12951	punct	×,SSC
R8298	T12953	T12947	punct	),°C
R8299	T12954	T12941	punct	.,washed
R83	T387	T388	mark	that,differ
R830	T1738	T1731	aux	would,bring
R8300	T12956	T12957	det	The,probes
R8301	T12957	T12958	nsubj	probes,hybridize
R8302	T12959	T12957	appos	each,probes
R8303	T12960	T12958	prep	to,hybridize
R8304	T12961	T12962	advmod	only,band
R8305	T12962	T12960	pobj	band,to
R8306	T12963	T12962	nummod	one,band
R8307	T12964	T12962	prep	on,band
R8308	T12965	T12966	det	a,blot
R8309	T12966	T12964	pobj	blot,on
R831	T1739	T1740	nummod	one,gene
R8310	T12967	T12966	compound	Southern,blot
R8311	T12968	T12958	punct	", ",hybridize
R8312	T12969	T12958	advcl	indicating,hybridize
R8313	T12970	T12971	mark	that,detects
R8314	T12971	T12969	ccomp	detects,indicating
R8315	T12972	T12973	det	each,probe
R8316	T12973	T12971	nsubj	probe,detects
R8317	T12974	T12971	advmod	only,detects
R8318	T12975	T12976	nummod	one,gene
R8319	T12976	T12971	dobj	gene,detects
R832	T1740	T1731	dobj	gene,bring
R8320	T12977	T12978	amod	olfactory,receptor
R8321	T12978	T12976	compound	receptor,gene
R8322	T12979	T12958	punct	.,hybridize
R8323	T12981	T12982	compound	BLAST,analyses
R8324	T12982	T12983	nsubj	analyses,show
R8325	T12984	T12985	mark	that,is
R8326	T12985	T12983	advcl	is,show
R8327	T12986	T12987	det	the,probe
R8328	T12987	T12985	nsubj	probe,is
R8329	T12988	T12987	compound	AY318555,probe
R833	T1741	T1742	amod	olfactory,receptor
R8330	T12989	T12985	acomp	unique,is
R8331	T12990	T12985	prep	in,is
R8332	T12991	T12992	poss	Celera,assembly
R8333	T12992	T12990	pobj	assembly,in
R8334	T12993	T12991	case	's,Celera
R8335	T12994	T12995	compound	mouse,genome
R8336	T12995	T12992	compound	genome,assembly
R8337	T12996	T12997	punct	(,Release
R8338	T12997	T12985	parataxis	Release,is
R8339	T12998	T12997	nummod	13,Release
R834	T1742	T1740	compound	receptor,gene
R8340	T12999	T12997	punct	),Release
R8341	T13000	T12985	punct	", ",is
R8342	T13001	T12985	cc	and,is
R8343	T13002	T13003	mark	that,is
R8344	T13003	T12985	conj	is,is
R8345	T13004	T13005	det	the,probe
R8346	T13005	T13003	nsubj	probe,is
R8347	T13006	T13005	compound	AY317365,probe
R8348	T13007	T13003	acomp	similar,is
R8349	T13008	T13007	prep	to,similar
R835	T1743	T1731	prep	into,bring
R8350	T13009	T13010	advmod	only,region
R8351	T13010	T13008	pobj	region,to
R8352	T13011	T13010	nummod	one,region
R8353	T13012	T13010	amod	other,region
R8354	T13013	T13010	amod	genomic,region
R8355	T13014	T12983	punct	.,show
R8356	T13016	T13017	det	This,region
R8357	T13017	T13020	nsubj	region,is
R8358	T13018	T13017	amod	potential,region
R8359	T13019	T13017	amod	cross-hybridizing,region
R836	T1744	T1745	det	a,configuration
R8360	T13021	T13022	quantmod	over,10
R8361	T13022	T13023	nummod	10,Mb
R8362	T13023	T13024	npadvmod	Mb,from
R8363	T13024	T13020	prep	from,is
R8364	T13025	T13026	det	the,region
R8365	T13026	T13024	pobj	region,from
R8366	T13027	T13026	amod	nearest,region
R8367	T13028	T13029	amod	olfactory,receptor
R8368	T13029	T13026	compound	receptor,region
R8369	T13030	T13026	compound	coding,region
R837	T1745	T1743	pobj	configuration,into
R8370	T13031	T13020	cc	and,is
R8371	T13032	T13020	conj	is,is
R8372	T13033	T13032	advmod	thus,is
R8373	T13034	T13035	advmod	highly,unlikely
R8374	T13035	T13032	acomp	unlikely,is
R8375	T13036	T13037	aux	to,included
R8376	T13037	T13035	xcomp	included,unlikely
R8377	T13038	T13037	auxpass	be,included
R8378	T13039	T13037	prep	in,included
R8379	T13040	T13041	det	any,transcript
R838	T1797	T1798	mark	as,expressed
R8380	T13041	T13039	pobj	transcript,in
R8381	T13042	T13043	amod	olfactory,receptor
R8382	T13043	T13041	compound	receptor,transcript
R8383	T13044	T13020	punct	.,is
R8384	T13046	T13047	amod	Low,power
R8385	T13047	T13049	compound	power,images
R8386	T13048	T13047	punct	-,power
R8387	T13049	T13050	nsubjpass	images,composed
R8388	T13051	T13050	auxpass	are,composed
R8389	T13052	T13050	prep	of,composed
R839	T1746	T1747	advmod	transcriptionally,active
R8390	T13053	T13054	nummod	three,micrographs
R8391	T13054	T13052	pobj	micrographs,of
R8392	T13055	T13054	amod	overlapping,micrographs
R8393	T13056	T13057	punct	(,40
R8394	T13057	T13054	parataxis	40,micrographs
R8395	T13058	T13057	punct	×,40
R8396	T13059	T13057	punct	),40
R8397	T13060	T13054	acl	assembled,micrographs
R8398	T13061	T13060	prep	in,assembled
R8399	T13062	T13063	compound	Adobe,Photoshop
R84	T388	T371	ccomp	differ,find
R840	T1798	T1792	advcl	expressed,is
R8400	T13063	T13061	pobj	Photoshop,in
R8401	T13064	T13063	nummod	7.0,Photoshop
R8402	T13065	T13050	punct	.,composed
R8403	T13084	T13085	det	A,list
R8404	T13085	T13086	nsubjpass	list,found
R8405	T13087	T13085	prep	of,list
R8406	T13088	T13089	det	the,primers
R8407	T13089	T13087	pobj	primers,of
R8408	T13090	T13089	acl	used,primers
R8409	T13091	T13092	aux	to,confirm
R841	T1747	T1745	amod	active,configuration
R8410	T13092	T13090	advcl	confirm,used
R8411	T13093	T13094	det	the,expression
R8412	T13094	T13092	dobj	expression,confirm
R8413	T13095	T13094	prep	of,expression
R8414	T13096	T13097	amod	olfactory,receptor
R8415	T13097	T13098	compound	receptor,genes
R8416	T13098	T13095	pobj	genes,of
R8417	T13099	T13092	prep	by,confirm
R8418	T13100	T13101	compound	RT,PCR
R8419	T13101	T13099	pobj	PCR,by
R842	T1748	T1745	amod	genomic,configuration
R8420	T13102	T13101	punct	-,PCR
R8421	T13103	T13101	cc	and,PCR
R8422	T13104	T13101	conj	PCR,PCR
R8423	T13105	T13101	prep	from,PCR
R8424	T13106	T13107	compound	cDNA,templates
R8425	T13107	T13105	pobj	templates,from
R8426	T13108	T13107	compound	library,templates
R8427	T13109	T13086	aux	can,found
R8428	T13110	T13086	auxpass	be,found
R8429	T13111	T13086	prep	in,found
R843	T1799	T1800	advmod	even,transgenes
R8430	T13112	T13113	amod	Additional,file
R8431	T13113	T13111	pobj	file,in
R8432	T13114	T13113	compound	data,file
R8433	T13115	T13113	nummod	1,file
R8434	T13116	T13086	punct	.,found
R8435	T13118	T13119	det	The,conditions
R8436	T13119	T13121	nsubjpass	conditions,found
R8437	T13120	T13119	amod	experimental,conditions
R8438	T13122	T13119	acl	used,conditions
R8439	T13123	T13122	prep	for,used
R844	T1800	T1798	nsubjpass	transgenes,expressed
R8440	T13124	T13125	amod	real,time
R8441	T13125	T13127	compound	time,PCR
R8442	T13126	T13125	punct	-,time
R8443	T13127	T13123	pobj	PCR,for
R8444	T13128	T13121	aux	can,found
R8445	T13129	T13121	auxpass	be,found
R8446	T13130	T13121	prep	in,found
R8447	T13131	T13132	amod	Additional,file
R8448	T13132	T13130	pobj	file,in
R8449	T13133	T13132	compound	data,file
R845	T1801	T1802	amod	olfactory,receptor
R8450	T13134	T13132	nummod	2,file
R8451	T13135	T13121	punct	.,found
R846	T1749	T1731	punct	", ",bring
R847	T1802	T1800	compound	receptor,transgenes
R848	T1803	T1800	prep	with,transgenes
R849	T1804	T1805	amod	identical,regions
R85	T389	T390	compound	transcript,levels
R850	T1750	T1751	mark	as,observed
R851	T1805	T1803	pobj	regions,with
R852	T1806	T1805	amod	upstream,regions
R853	T1807	T1798	auxpass	are,expressed
R854	T1751	T1731	advcl	observed,bring
R855	T1808	T1798	prep	in,expressed
R856	T1809	T1810	amod	different,neurons
R857	T1810	T1808	pobj	neurons,in
R858	T1752	T1751	prep	for,observed
R859	T1811	T1812	punct	[,18
R86	T390	T388	nsubj	levels,differ
R860	T1812	T1798	parataxis	18,expressed
R861	T1813	T1812	punct	],18
R862	T1753	T1754	det	the,locus
R863	T1814	T1792	punct	.,is
R864	T1754	T1752	pobj	locus,for
R865	T1816	T1817	prep	In,contain
R866	T1818	T1819	det	a,model
R867	T1755	T1756	compound	yeast,mating
R868	T1819	T1816	pobj	model,In
R869	T1820	T1819	amod	third,model
R87	T391	T390	prep	in,levels
R870	T1756	T1757	compound	mating,type
R871	T1821	T1817	punct	", ",contain
R872	T1822	T1823	expl	there,be
R873	T1823	T1817	ccomp	be,contain
R874	T1757	T1754	compound	type,locus
R875	T1824	T1823	aux	would,be
R876	T1825	T1826	det	a,quantity
R877	T1826	T1823	attr	quantity,be
R878	T1758	T1759	punct	[,20
R879	T1827	T1826	amod	limiting,quantity
R88	T392	T393	det	the,epithelium
R880	T1828	T1826	prep	of,quantity
R881	T1759	T1754	parataxis	20,locus
R882	T1829	T1830	compound	transcription,factors
R883	T1830	T1828	pobj	factors,of
R884	T1831	T1817	punct	-,contain
R885	T1760	T1759	punct	],20
R886	T1832	T1833	det	the,cell
R887	T1833	T1817	nsubj	cell,contain
R888	T1834	T1817	aux	might,contain
R889	T1761	T1754	cc	and,locus
R89	T393	T391	pobj	epithelium,in
R890	T1835	T1836	det	a,machine
R891	T1836	T1817	dobj	machine,contain
R892	T1837	T1836	amod	single,machine
R893	T1762	T1763	det	the,genes
R894	T1838	T1836	amod	transcriptional,machine
R895	T1839	T1836	punct	',machine
R896	T1840	T1836	punct	',machine
R897	T1841	T1842	dep	that,is
R898	T1842	T1836	relcl	is,machine
R899	T1843	T1842	acomp	capable,is
R9	T305	T301	acomp	available,is
R90	T394	T393	amod	olfactory,epithelium
R900	T1763	T1754	conj	genes,locus
R901	T1764	T1763	amod	mammalian,genes
R902	T1844	T1843	prep	of,capable
R903	T1765	T1763	compound	immunoglobulin,genes
R904	T1845	T1844	pcomp	accommodating,of
R905	T1846	T1847	det	the,promoter
R906	T1847	T1845	dobj	promoter,accommodating
R907	T1766	T1767	punct	[,21
R908	T1848	T1847	prep	of,promoter
R909	T1849	T1850	advmod	only,gene
R91	T395	T388	aux	can,differ
R910	T1767	T1763	parataxis	21,genes
R911	T1850	T1848	pobj	gene,of
R912	T1851	T1850	nummod	one,gene
R913	T1852	T1853	amod	olfactory,receptor
R914	T1768	T1767	punct	],21
R915	T1853	T1850	compound	receptor,gene
R916	T1854	T1836	punct	", ",machine
R917	T1769	T1731	punct	.,bring
R918	T1855	T1836	amod	similar,machine
R919	T1856	T1855	prep	to,similar
R92	T396	T388	prep	between,differ
R920	T1857	T1858	det	the,body
R921	T1771	T1772	advmod	Alternatively,invokes
R922	T1858	T1856	pobj	body,to
R923	T1859	T1860	compound	expression,site
R924	T1860	T1858	compound	site,body
R925	T1861	T1858	acl	used,body
R926	T1773	T1772	punct	", ",invokes
R927	T1862	T1861	agent	by,used
R928	T1863	T1864	amod	African,trypanosomes
R929	T1864	T1862	pobj	trypanosomes,by
R93	T397	T398	amod	olfactory,receptors
R930	T1865	T1866	aux	to,ensure
R931	T1774	T1775	det	a,model
R932	T1866	T1861	advcl	ensure,used
R933	T1867	T1868	amod	singular,expression
R934	T1775	T1772	nsubj	model,invokes
R935	T1868	T1866	dobj	expression,ensure
R936	T1869	T1868	prep	of,expression
R937	T1776	T1775	amod	second,model
R938	T1870	T1871	advmod	only,set
R939	T1871	T1869	pobj	set,of
R94	T398	T396	pobj	receptors,between
R940	T1872	T1871	nummod	one,set
R941	T1777	T1778	det	a,code
R942	T1873	T1871	prep	of,set
R943	T1874	T1875	amod	variant,glycoprotein
R944	T1875	T1877	compound	glycoprotein,genes
R945	T1778	T1772	dobj	code,invokes
R946	T1876	T1875	compound	surface,glycoprotein
R947	T1877	T1873	pobj	genes,of
R948	T1878	T1879	punct	[,22
R949	T1879	T1817	parataxis	22,contain
R95	T399	T388	prep	by,differ
R950	T1880	T1879	punct	],22
R951	T1779	T1778	amod	combinatorial,code
R952	T1881	T1817	punct	.,contain
R953	T1883	T1884	advmod	Finally,send
R954	T1780	T1778	prep	of,code
R955	T1885	T1884	punct	", ",send
R956	T1886	T1884	prep	in,send
R957	T1781	T1782	compound	transcription,factor
R958	T1887	T1888	det	a,model
R959	T1888	T1886	pobj	model,in
R96	T400	T401	advmod	up,300-fold
R960	T1782	T1783	compound	factor,sites
R961	T1889	T1888	amod	fourth,model
R962	T1890	T1884	punct	", ",send
R963	T1891	T1892	amod	transcriptional,activity
R964	T1783	T1780	pobj	sites,of
R965	T1892	T1884	nsubj	activity,send
R966	T1893	T1892	prep	at,activity
R967	T1894	T1895	nummod	one,allele
R968	T1784	T1783	compound	binding,sites
R969	T1895	T1893	pobj	allele,at
R97	T401	T399	pcomp	300-fold,by
R970	T1896	T1897	advmod	stochastically,chosen
R971	T1785	T1778	amod	unique,code
R972	T1897	T1895	amod	chosen,allele
R973	T1898	T1899	amod	olfactory,receptor
R974	T1899	T1895	compound	receptor,allele
R975	T1786	T1785	prep	to,unique
R976	T1900	T1884	aux	might,send
R977	T1901	T1902	amod	negative,feedback
R978	T1902	T1884	dobj	feedback,send
R979	T1787	T1788	det	each,gene
R98	T402	T401	quantmod	to,300-fold
R980	T1788	T1786	pobj	gene,to
R981	T1789	T1772	punct	.,invokes
R982	T1791	T1792	nsubj	This,is
R983	T1903	T1904	aux	to,repress
R984	T1904	T1884	advcl	repress,send
R985	T1793	T1792	acomp	unlikely,is
R986	T1905	T1904	dobj	activity,repress
R987	T1906	T1905	prep	of,activity
R988	T1907	T1908	det	all,receptors
R989	T1794	T1792	punct	", ",is
R99	T403	T371	punct	.,find
R990	T1908	T1906	pobj	receptors,of
R991	T1909	T1908	amod	other,receptors
R992	T1795	T1792	advmod	however,is
R993	T1910	T1908	amod	olfactory,receptors
R994	T1911	T1884	cc	and,send
R995	T1912	T1911	punct	/,and
R996	T1796	T1792	punct	", ",is
R997	T1913	T1911	cc	or,and
R998	T1914	T1915	amod	positive,feedback
R999	T2009	T2010	amod	many,genes

PMC:329117 JSONTXT 8 Projects

Annnotations TAB TSV DIC JSON TextAE Lectin_function IAV-Glycan

PMC:329117 JSON TXT 8 Projects