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PMC:1860061 / 1282-2738 JSONTXT 5 Projects

Annnotations TAB TSV DIC JSON TextAE-old TextAE

Id Subject Object Predicate Lexical cue
T2422 0-14 NN denotes Susceptibility
T2424 15-17 IN denotes to
T2425 18-22 RBS denotes most
T2427 23-30 JJ denotes complex
T2426 31-39 NNS denotes diseases
T2428 40-42 VBZ denotes is
T2423 43-53 VBN denotes controlled
T2429 54-56 IN denotes by
T2430 57-61 JJ denotes many
T2431 62-67 NNS denotes genes
T2432 67-69 , denotes ,
T2433 69-73 RB denotes each
T2434 74-80 VBG denotes having
T2435 81-82 DT denotes a
T2437 83-88 JJ denotes small
T2436 89-95 NN denotes effect
T2438 96-98 IN denotes on
T2439 99-102 DT denotes the
T2440 103-110 NN denotes disease
T2441 110-111 . denotes .
T2442 111-205 sentence denotes One example is rheumatoid arthritis (RA), a common complex multifactorial autoimmune disease.
T2443 112-115 CD denotes One
T2444 116-123 NN denotes example
T2445 124-126 VBZ denotes is
T2446 127-137 JJ denotes rheumatoid
T2447 138-147 NN denotes arthritis
T2448 148-149 -LRB- denotes (
T2449 149-151 NN denotes RA
T2450 151-152 -RRB- denotes )
T2451 152-154 , denotes ,
T2452 154-155 DT denotes a
T2454 156-162 JJ denotes common
T2455 163-170 NN denotes complex
T2456 171-185 JJ denotes multifactorial
T2457 186-196 JJ denotes autoimmune
T2453 197-204 NN denotes disease
T2458 204-205 . denotes .
T2459 205-358 sentence denotes Several studies have been carried out to detect the genetic basis of RA, and more than 30 genomic regions have shown evidence of linkage to the disease.
T2460 206-213 JJ denotes Several
T2461 214-221 NNS denotes studies
T2463 222-226 VBP denotes have
T2464 227-231 VBN denotes been
T2462 232-239 VBN denotes carried
T2465 240-243 RP denotes out
T2466 244-246 TO denotes to
T2467 247-253 VB denotes detect
T2468 254-257 DT denotes the
T2470 258-265 JJ denotes genetic
T2469 266-271 NN denotes basis
T2471 272-274 IN denotes of
T2472 275-277 NN denotes RA
T2473 277-279 , denotes ,
T2474 279-282 CC denotes and
T2475 283-287 JJR denotes more
T2477 288-292 IN denotes than
T2476 293-295 CD denotes 30
T2479 296-303 JJ denotes genomic
T2478 304-311 NNS denotes regions
T2481 312-316 VBP denotes have
T2480 317-322 VBN denotes shown
T2482 323-331 NN denotes evidence
T2483 332-334 IN denotes of
T2484 335-342 NN denotes linkage
T2485 343-345 IN denotes to
T2486 346-349 DT denotes the
T2487 350-357 NN denotes disease
T2488 357-358 . denotes .
T2489 358-500 sentence denotes Most of these genomic regions did not reach a genome-wide significant threshold value of linkage, with P values between 0.05 and 0.001 [1-5].
T2490 359-363 JJS denotes Most
T2492 364-366 IN denotes of
T2493 367-372 DT denotes these
T2495 373-380 JJ denotes genomic
T2494 381-388 NNS denotes regions
T2496 389-392 VBD denotes did
T2497 393-396 RB denotes not
T2491 397-402 VB denotes reach
T2498 403-404 DT denotes a
T2500 405-411 NN denotes genome
T2502 411-412 HYPH denotes -
T2501 412-416 JJ denotes wide
T2503 417-428 JJ denotes significant
T2504 429-438 NN denotes threshold
T2499 439-444 NN denotes value
T2505 445-447 IN denotes of
T2506 448-455 NN denotes linkage
T2507 455-457 , denotes ,
T2508 457-461 IN denotes with
T2509 462-463 NN denotes P
T2510 464-470 NNS denotes values
T2511 471-478 IN denotes between
T2512 479-483 CD denotes 0.05
T2513 484-487 CC denotes and
T2514 488-493 CD denotes 0.001
T2515 494-495 -LRB- denotes [
T2516 495-496 CD denotes 1
T2517 496-497 SYM denotes -
T2518 497-498 CD denotes 5
T2519 498-499 -RRB- denotes ]
T2520 499-500 . denotes .
T2521 500-549 sentence denotes Thus, these loci only have a small effect on RA.
T2522 501-505 RB denotes Thus
T2524 505-507 , denotes ,
T2525 507-512 DT denotes these
T2526 513-517 NNS denotes loci
T2527 518-522 RB denotes only
T2523 523-527 VBP denotes have
T2528 528-529 DT denotes a
T2530 530-535 JJ denotes small
T2529 536-542 NN denotes effect
T2531 543-545 IN denotes on
T2532 546-548 NN denotes RA
T2533 548-549 . denotes .
T2534 549-701 sentence denotes Small genetic contributions could also be seen from the susceptibility genes of RA identified so far, including HLA-DR4, PADI4, PTPN22 and FCRL3 [6-9].
T2535 550-555 JJ denotes Small
T2537 556-563 JJ denotes genetic
T2536 564-577 NNS denotes contributions
T2539 578-583 MD denotes could
T2540 584-588 RB denotes also
T2541 589-591 VB denotes be
T2538 592-596 VBN denotes seen
T2542 597-601 IN denotes from
T2543 602-605 DT denotes the
T2545 606-620 NN denotes susceptibility
T2544 621-626 NNS denotes genes
T2546 627-629 IN denotes of
T2547 630-632 NN denotes RA
T2548 633-643 VBN denotes identified
T2549 644-646 RB denotes so
T2550 647-650 RB denotes far
T2551 650-652 , denotes ,
T2552 652-661 VBG denotes including
T2553 662-665 NN denotes HLA
T2555 665-666 HYPH denotes -
T2554 666-669 NN denotes DR4
T2556 669-671 , denotes ,
T2557 671-676 NN denotes PADI4
T2558 676-678 , denotes ,
T2559 678-684 NN denotes PTPN22
T2560 685-688 CC denotes and
T2561 689-694 NN denotes FCRL3
T2562 695-696 -LRB- denotes [
T2563 696-697 CD denotes 6
T2564 697-698 SYM denotes -
T2565 698-699 CD denotes 9
T2566 699-700 -RRB- denotes ]
T2567 700-701 . denotes .
T2568 701-835 sentence denotes Except for HLA-DR4, which is strongly associated with RA, all the other susceptibility genes have only a small effect on the disease.
T2569 702-708 IN denotes Except
T2571 709-712 IN denotes for
T2572 713-716 NN denotes HLA
T2574 716-717 HYPH denotes -
T2573 717-720 NN denotes DR4
T2575 720-722 , denotes ,
T2576 722-727 WDT denotes which
T2578 728-730 VBZ denotes is
T2579 731-739 RB denotes strongly
T2577 740-750 VBN denotes associated
T2580 751-755 IN denotes with
T2581 756-758 NN denotes RA
T2582 758-760 , denotes ,
T2583 760-763 PDT denotes all
T2585 764-767 DT denotes the
T2586 768-773 JJ denotes other
T2587 774-788 NN denotes susceptibility
T2584 789-794 NNS denotes genes
T2570 795-799 VBP denotes have
T2588 800-804 RB denotes only
T2590 805-806 DT denotes a
T2591 807-812 JJ denotes small
T2589 813-819 NN denotes effect
T2592 820-822 IN denotes on
T2593 823-826 DT denotes the
T2594 827-834 NN denotes disease
T2595 834-835 . denotes .
T2596 835-914 sentence denotes In the mouse model of RA, small genetic contributions are also often observed.
T2597 836-838 IN denotes In
T2599 839-842 DT denotes the
T2601 843-848 NN denotes mouse
T2600 849-854 NN denotes model
T2602 855-857 IN denotes of
T2603 858-860 NN denotes RA
T2604 860-862 , denotes ,
T2605 862-867 JJ denotes small
T2607 868-875 JJ denotes genetic
T2606 876-889 NNS denotes contributions
T2608 890-893 VBP denotes are
T2609 894-898 RB denotes also
T2610 899-904 RB denotes often
T2598 905-913 VBN denotes observed
T2611 913-914 . denotes .
T2612 914-1105 sentence denotes For example, in a previous study, we carried out a genome screen to identify the quantitative trait loci (QTL) in collagen-induced arthritis (CIA), which is a widely used animal model of RA.
T2613 915-918 IN denotes For
T2615 919-926 NN denotes example
T2616 926-928 , denotes ,
T2617 928-930 IN denotes in
T2618 931-932 DT denotes a
T2620 933-941 JJ denotes previous
T2619 942-947 NN denotes study
T2621 947-949 , denotes ,
T2622 949-951 PRP denotes we
T2614 952-959 VBD denotes carried
T2623 960-963 RP denotes out
T2624 964-965 DT denotes a
T2626 966-972 NN denotes genome
T2625 973-979 NN denotes screen
T2627 980-982 TO denotes to
T2628 983-991 VB denotes identify
T2629 992-995 DT denotes the
T2631 996-1008 JJ denotes quantitative
T2632 1009-1014 NN denotes trait
T2630 1015-1019 NNS denotes loci
T2633 1020-1021 -LRB- denotes (
T2634 1021-1024 NN denotes QTL
T2635 1024-1025 -RRB- denotes )
T2636 1026-1028 IN denotes in
T2637 1029-1037 NN denotes collagen
T2639 1037-1038 HYPH denotes -
T2638 1038-1045 VBN denotes induced
T2640 1046-1055 NN denotes arthritis
T2641 1056-1057 -LRB- denotes (
T2642 1057-1060 NN denotes CIA
T2643 1060-1061 -RRB- denotes )
T2644 1061-1063 , denotes ,
T2645 1063-1068 WDT denotes which
T2646 1069-1071 VBZ denotes is
T2647 1072-1073 DT denotes a
T2649 1074-1080 RB denotes widely
T2650 1081-1085 VBN denotes used
T2651 1086-1092 NN denotes animal
T2648 1093-1098 NN denotes model
T2652 1099-1101 IN denotes of
T2653 1102-1104 NN denotes RA
T2654 1104-1105 . denotes .
T2655 1105-1283 sentence denotes Only one QTL, Cia2, was identified for the phenotype of CIA severity, but this QTL contributes to only 16% of the phenotype variations for CIA susceptibility in F2 progeny [10].
T2656 1106-1110 RB denotes Only
T2658 1111-1114 CD denotes one
T2657 1115-1118 NN denotes QTL
T2660 1118-1120 , denotes ,
T2661 1120-1124 NN denotes Cia2
T2662 1124-1126 , denotes ,
T2663 1126-1129 VBD denotes was
T2659 1130-1140 VBN denotes identified
T2664 1141-1144 IN denotes for
T2665 1145-1148 DT denotes the
T2666 1149-1158 NN denotes phenotype
T2667 1159-1161 IN denotes of
T2668 1162-1165 NN denotes CIA
T2669 1166-1174 NN denotes severity
T2670 1174-1176 , denotes ,
T2671 1176-1179 CC denotes but
T2672 1180-1184 DT denotes this
T2673 1185-1188 NN denotes QTL
T2674 1189-1200 VBZ denotes contributes
T2675 1201-1203 IN denotes to
T2676 1204-1208 RB denotes only
T2677 1209-1211 CD denotes 16
T2678 1211-1212 NN denotes %
T2679 1213-1215 IN denotes of
T2680 1216-1219 DT denotes the
T2682 1220-1229 NN denotes phenotype
T2681 1230-1240 NNS denotes variations
T2683 1241-1244 IN denotes for
T2684 1245-1248 NN denotes CIA
T2685 1249-1263 NN denotes susceptibility
T2686 1264-1266 IN denotes in
T2687 1267-1269 NN denotes F2
T2688 1270-1277 NN denotes progeny
T2689 1278-1279 -LRB- denotes [
T2690 1279-1281 CD denotes 10
T2691 1281-1282 -RRB- denotes ]
T2692 1282-1283 . denotes .
T2693 1283-1456 sentence denotes This suggests that there must be other susceptibility genes whose contributions were not big enough to reach the stringent significance threshold value of linkage analysis.
T2694 1284-1288 DT denotes This
T2695 1289-1297 VBZ denotes suggests
T2696 1298-1302 IN denotes that
T2698 1303-1308 EX denotes there
T2699 1309-1313 MD denotes must
T2697 1314-1316 VB denotes be
T2700 1317-1322 JJ denotes other
T2702 1323-1337 NN denotes susceptibility
T2701 1338-1343 NNS denotes genes
T2703 1344-1349 WP$ denotes whose
T2704 1350-1363 NNS denotes contributions
T2705 1364-1368 VBD denotes were
T2706 1369-1372 RB denotes not
T2707 1373-1376 JJ denotes big
T2708 1377-1383 RB denotes enough
T2709 1384-1386 TO denotes to
T2710 1387-1392 VB denotes reach
T2711 1393-1396 DT denotes the
T2713 1397-1406 JJ denotes stringent
T2714 1407-1419 NN denotes significance
T2715 1420-1429 NN denotes threshold
T2712 1430-1435 NN denotes value
T2716 1436-1438 IN denotes of
T2717 1439-1446 NN denotes linkage
T2718 1447-1455 NN denotes analysis
T2719 1455-1456 . denotes .
R210 T2422 T2423 nsubjpass Susceptibility,controlled
R211 T2424 T2422 prep to,Susceptibility
R212 T2425 T2426 advmod most,diseases
R213 T2426 T2424 pobj diseases,to
R214 T2427 T2426 amod complex,diseases
R215 T2428 T2423 auxpass is,controlled
R216 T2429 T2423 agent by,controlled
R217 T2430 T2431 amod many,genes
R218 T2431 T2429 pobj genes,by
R219 T2432 T2431 punct ", ",genes
R220 T2433 T2434 advmod each,having
R221 T2434 T2431 acl having,genes
R222 T2435 T2436 det a,effect
R223 T2436 T2434 dobj effect,having
R224 T2437 T2436 amod small,effect
R225 T2438 T2436 prep on,effect
R226 T2439 T2440 det the,disease
R227 T2440 T2438 pobj disease,on
R228 T2441 T2423 punct .,controlled
R229 T2443 T2444 nummod One,example
R230 T2444 T2445 nsubj example,is
R231 T2446 T2447 amod rheumatoid,arthritis
R232 T2447 T2445 attr arthritis,is
R233 T2448 T2447 punct (,arthritis
R234 T2449 T2447 appos RA,arthritis
R235 T2450 T2447 punct ),arthritis
R236 T2451 T2447 punct ", ",arthritis
R237 T2452 T2453 det a,disease
R238 T2453 T2447 appos disease,arthritis
R239 T2454 T2453 amod common,disease
R240 T2455 T2453 nmod complex,disease
R241 T2456 T2453 amod multifactorial,disease
R242 T2457 T2453 amod autoimmune,disease
R243 T2458 T2445 punct .,is
R244 T2460 T2461 amod Several,studies
R245 T2461 T2462 nsubjpass studies,carried
R246 T2463 T2462 aux have,carried
R247 T2464 T2462 auxpass been,carried
R248 T2465 T2462 prt out,carried
R249 T2466 T2467 aux to,detect
R250 T2467 T2462 advcl detect,carried
R251 T2468 T2469 det the,basis
R252 T2469 T2467 dobj basis,detect
R253 T2470 T2469 amod genetic,basis
R254 T2471 T2469 prep of,basis
R255 T2472 T2471 pobj RA,of
R256 T2473 T2462 punct ", ",carried
R257 T2474 T2462 cc and,carried
R258 T2475 T2476 amod more,30
R259 T2476 T2478 nummod 30,regions
R260 T2477 T2476 quantmod than,30
R261 T2478 T2480 nsubj regions,shown
R262 T2479 T2478 amod genomic,regions
R263 T2480 T2462 conj shown,carried
R264 T2481 T2480 aux have,shown
R265 T2482 T2480 dobj evidence,shown
R266 T2483 T2482 prep of,evidence
R267 T2484 T2483 pobj linkage,of
R268 T2485 T2484 prep to,linkage
R269 T2486 T2487 det the,disease
R270 T2487 T2485 pobj disease,to
R271 T2488 T2480 punct .,shown
R272 T2490 T2491 nsubj Most,reach
R273 T2492 T2490 prep of,Most
R274 T2493 T2494 det these,regions
R275 T2494 T2492 pobj regions,of
R276 T2495 T2494 amod genomic,regions
R277 T2496 T2491 aux did,reach
R278 T2497 T2491 neg not,reach
R279 T2498 T2499 det a,value
R280 T2499 T2491 dobj value,reach
R281 T2500 T2501 npadvmod genome,wide
R282 T2501 T2499 amod wide,value
R283 T2502 T2501 punct -,wide
R284 T2503 T2499 amod significant,value
R285 T2504 T2499 compound threshold,value
R286 T2505 T2499 prep of,value
R287 T2506 T2505 pobj linkage,of
R288 T2507 T2499 punct ", ",value
R289 T2508 T2499 prep with,value
R290 T2509 T2510 compound P,values
R291 T2510 T2508 pobj values,with
R292 T2511 T2510 prep between,values
R293 T2512 T2511 pobj 0.05,between
R294 T2513 T2512 cc and,0.05
R295 T2514 T2512 conj 0.001,0.05
R296 T2515 T2516 punct [,1
R297 T2516 T2491 parataxis 1,reach
R298 T2517 T2518 punct -,5
R299 T2518 T2516 prep 5,1
R300 T2519 T2516 punct ],1
R301 T2520 T2491 punct .,reach
R302 T2522 T2523 advmod Thus,have
R303 T2524 T2523 punct ", ",have
R304 T2525 T2526 det these,loci
R305 T2526 T2523 nsubj loci,have
R306 T2527 T2523 advmod only,have
R307 T2528 T2529 det a,effect
R308 T2529 T2523 dobj effect,have
R309 T2530 T2529 amod small,effect
R310 T2531 T2529 prep on,effect
R311 T2532 T2531 pobj RA,on
R312 T2533 T2523 punct .,have
R313 T2535 T2536 amod Small,contributions
R314 T2536 T2538 nsubjpass contributions,seen
R315 T2537 T2536 amod genetic,contributions
R316 T2539 T2538 aux could,seen
R317 T2540 T2538 advmod also,seen
R318 T2541 T2538 auxpass be,seen
R319 T2542 T2538 prep from,seen
R320 T2543 T2544 det the,genes
R321 T2544 T2542 pobj genes,from
R322 T2545 T2544 compound susceptibility,genes
R323 T2546 T2544 prep of,genes
R324 T2547 T2546 pobj RA,of
R325 T2548 T2544 acl identified,genes
R326 T2549 T2550 advmod so,far
R327 T2550 T2548 advmod far,identified
R328 T2551 T2544 punct ", ",genes
R329 T2552 T2544 prep including,genes
R330 T2553 T2554 compound HLA,DR4
R331 T2554 T2552 pobj DR4,including
R332 T2555 T2554 punct -,DR4
R333 T2556 T2554 punct ", ",DR4
R334 T2557 T2554 conj PADI4,DR4
R335 T2558 T2557 punct ", ",PADI4
R336 T2559 T2557 conj PTPN22,PADI4
R337 T2560 T2559 cc and,PTPN22
R338 T2561 T2559 conj FCRL3,PTPN22
R339 T2562 T2563 punct [,6
R340 T2563 T2538 parataxis 6,seen
R341 T2564 T2565 punct -,9
R342 T2565 T2563 prep 9,6
R343 T2566 T2563 punct ],6
R344 T2567 T2538 punct .,seen
R345 T2569 T2570 prep Except,have
R346 T2571 T2569 prep for,Except
R347 T2572 T2573 compound HLA,DR4
R348 T2573 T2571 pobj DR4,for
R349 T2574 T2573 punct -,DR4
R350 T2575 T2573 punct ", ",DR4
R351 T2576 T2577 dep which,associated
R352 T2577 T2573 relcl associated,DR4
R353 T2578 T2577 auxpass is,associated
R354 T2579 T2577 advmod strongly,associated
R355 T2580 T2577 prep with,associated
R356 T2581 T2580 pobj RA,with
R357 T2582 T2570 punct ", ",have
R358 T2583 T2584 predet all,genes
R359 T2584 T2570 nsubj genes,have
R360 T2585 T2584 det the,genes
R361 T2586 T2584 amod other,genes
R362 T2587 T2584 compound susceptibility,genes
R363 T2588 T2589 advmod only,effect
R364 T2589 T2570 dobj effect,have
R365 T2590 T2589 det a,effect
R366 T2591 T2589 amod small,effect
R367 T2592 T2589 prep on,effect
R368 T2593 T2594 det the,disease
R369 T2660 T2657 punct ", ",QTL
R370 T2594 T2592 pobj disease,on
R371 T2661 T2657 appos Cia2,QTL
R372 T2595 T2570 punct .,have
R373 T2662 T2659 punct ", ",identified
R374 T2663 T2659 auxpass was,identified
R375 T2597 T2598 prep In,observed
R376 T2664 T2659 prep for,identified
R377 T2665 T2666 det the,phenotype
R378 T2666 T2664 pobj phenotype,for
R379 T2599 T2600 det the,model
R380 T2667 T2666 prep of,phenotype
R381 T2668 T2669 compound CIA,severity
R382 T2669 T2667 pobj severity,of
R383 T2670 T2659 punct ", ",identified
R384 T2600 T2597 pobj model,In
R385 T2671 T2659 cc but,identified
R386 T2672 T2673 det this,QTL
R387 T2673 T2674 nsubj QTL,contributes
R388 T2674 T2659 conj contributes,identified
R389 T2601 T2600 compound mouse,model
R390 T2675 T2674 prep to,contributes
R391 T2676 T2677 advmod only,16
R392 T2602 T2600 prep of,model
R393 T2677 T2678 nummod 16,%
R394 T2678 T2675 pobj %,to
R395 T2679 T2678 prep of,%
R396 T2603 T2602 pobj RA,of
R397 T2680 T2681 det the,variations
R398 T2681 T2679 pobj variations,of
R399 T2682 T2681 compound phenotype,variations
R400 T2604 T2598 punct ", ",observed
R401 T2683 T2681 prep for,variations
R402 T2684 T2685 compound CIA,susceptibility
R403 T2685 T2683 pobj susceptibility,for
R404 T2605 T2606 amod small,contributions
R405 T2686 T2674 prep in,contributes
R406 T2687 T2688 compound F2,progeny
R407 T2606 T2598 nsubjpass contributions,observed
R408 T2688 T2686 pobj progeny,in
R409 T2607 T2606 amod genetic,contributions
R410 T2608 T2598 auxpass are,observed
R411 T2689 T2690 punct [,10
R412 T2609 T2610 advmod also,often
R413 T2690 T2674 parataxis 10,contributes
R414 T2691 T2690 punct ],10
R415 T2692 T2674 punct .,contributes
R416 T2610 T2598 advmod often,observed
R417 T2694 T2695 nsubj This,suggests
R418 T2696 T2697 mark that,be
R419 T2697 T2695 ccomp be,suggests
R420 T2698 T2697 expl there,be
R421 T2611 T2598 punct .,observed
R422 T2699 T2697 aux must,be
R423 T2700 T2701 amod other,genes
R424 T2613 T2614 prep For,carried
R425 T2701 T2697 attr genes,be
R426 T2702 T2701 compound susceptibility,genes
R427 T2703 T2704 poss whose,contributions
R428 T2704 T2705 dep contributions,were
R429 T2705 T2701 relcl were,genes
R430 T2615 T2613 pobj example,For
R431 T2706 T2705 neg not,were
R432 T2707 T2705 acomp big,were
R433 T2708 T2707 advmod enough,big
R434 T2616 T2614 punct ", ",carried
R435 T2709 T2710 aux to,reach
R436 T2710 T2707 xcomp reach,big
R437 T2711 T2712 det the,value
R438 T2617 T2614 prep in,carried
R439 T2712 T2710 dobj value,reach
R440 T2713 T2712 amod stringent,value
R441 T2714 T2712 compound significance,value
R442 T2715 T2712 compound threshold,value
R443 T2618 T2619 det a,study
R444 T2716 T2712 prep of,value
R445 T2717 T2718 compound linkage,analysis
R446 T2619 T2617 pobj study,in
R447 T2718 T2716 pobj analysis,of
R448 T2719 T2695 punct .,suggests
R449 T2620 T2619 amod previous,study
R452 T2621 T2614 punct ", ",carried
R455 T2622 T2614 nsubj we,carried
R459 T2623 T2614 prt out,carried
R462 T2624 T2625 det a,screen
R465 T2625 T2614 dobj screen,carried
R473 T2626 T2625 compound genome,screen
R475 T2627 T2628 aux to,identify
R478 T2628 T2614 advcl identify,carried
R482 T2629 T2630 det the,loci
R486 T2630 T2628 dobj loci,identify
R490 T2631 T2632 amod quantitative,trait
R494 T2632 T2630 compound trait,loci
R495 T2633 T2630 punct (,loci
R497 T2634 T2630 appos QTL,loci
R500 T2635 T2628 punct ),identify
R504 T2636 T2628 prep in,identify
R508 T2637 T2638 npadvmod collagen,induced
R509 T2638 T2640 amod induced,arthritis
R510 T2639 T2638 punct -,induced
R511 T2640 T2636 pobj arthritis,in
R515 T2641 T2640 punct (,arthritis
R518 T2642 T2640 appos CIA,arthritis
R524 T2643 T2640 punct ),arthritis
R525 T2644 T2640 punct ", ",arthritis
R529 T2645 T2646 dep which,is
R531 T2646 T2640 relcl is,arthritis
R535 T2647 T2648 det a,model
R538 T2648 T2646 attr model,is
R542 T2649 T2650 advmod widely,used
R546 T2650 T2648 amod used,model
R549 T2651 T2648 compound animal,model
R554 T2652 T2648 prep of,model
R557 T2653 T2652 pobj RA,of
R561 T2654 T2614 punct .,carried
R563 T2656 T2657 advmod Only,QTL
R568 T2657 T2659 nsubjpass QTL,identified
R572 T2658 T2657 nummod one,QTL