PMC:7106203 JSONTXT

{"target":"https://pubannotation.org/docs/sourcedb/PMC/sourceid/7106203","sourcedb":"PMC","sourceid":"7106203","source_url":"https://www.ncbi.nlm.nih.gov/pmc/7106203","text":"Genetic diversity and evolution of SARS-CoV-2\n\nAbstract\nCOVID-19 is a viral respiratory illness caused by a new coronavirus called SARS-CoV-2. The World Health Organization declared the SARS-CoV-2 outbreak a global public health emergency. We performed genetic analyses of eighty-six complete or near-complete genomes of SARS-CoV-2 and revealed many mutations and deletions on coding and non-coding regions. These observations provided evidence of the genetic diversity and rapid evolution of this novel coronavirus.\n\nHighlights\n• A novel coronavirus is officially known as SARS-CoV-2.\n• Eighty-six complete or near-complete genomes of SARS-CoV-2 were analyzed.\n• Many mutations and deletions on coding and non-coding regions of SARS-CoV-2 were found.\n\n1 The study\nA new coronavirus SARS-CoV-2 is spreading cross the world (Phan, 2020). Since the virus emerged at the seafood wholesale market at the end of last year (Zhu et al., 2019), the number of infected cases has been rising dramatically (Velavan and Meyer, 2020). Human-to-human transmission of SARS-CoV-2 has been confirmed (Nishiura et al., 2020). The virus has been detected in bronchoalveolar-lavage (Zhu et al., 2019), sputum (Lin et al., 2020), saliva (K.K. To et al., 2020), throat (Bastola et al., 2020) and nasopharyngeal swabs (To et al., 2020).\nNucleotide substitution has been proposed to be one of the most important mechanisms of viral evolution in nature (Lauring and Andino, 2010). The rapid spread of SARS-CoV-2 raises intriguing questions such as whether its evolution is driven by mutations. To assess the genetic variation, eighty-six complete or near-complete genomes of SARS-CoV-2 were collected from GISAID [https://www.gisaid.org/]. These SARS-CoV-2 strains were detected in infected patients from China (50), USA (11), Australia (5), Japan (5), France (4), Singapore (3), England (2), Taiwan (2), South Korea (1), Belgium (1), Germany (1), and Vietnam (1). The pair-wise nucleotide sequence alignment was performed by ClustalX2 (Saitou and Nei, 1987), and the sequence of the strain China/WHU01/2020/EPI_ISL_406716 was used as a reference genome.\nLike other betacoronaviruses, the genome of SARS-CoV-2 has a long ORF1ab polyprotein at the 5′ end, followed by four major structural proteins, including the spike surface glycoprotein, small envelope protein, matrix protein, and nucleocapsid protein (Phan, 2020). Our genetic analysis discovered three deletions in the genomes of SARS-CoV-2 from Japan (Aichi), USA (Wisconsin), and Australia (Victoria) as shown in Fig. 1 . Two deletions (three nucleotides and twenty-four nucleotides) were in the ORF1ab polyprotein, and one deletion (ten nucleotides) was in the 3′ end of the genome.\nFig. 1 Genomic organization of SARS-CoV-2 and pairwise nucleotide sequence alignment showing deletions in the ORF1ab polyprotein and in the 3′ end of the genome.\nIt is interesting that our nucleotide sequence alignment also revealed ninety-three mutations over the entire genomes of SARS-CoV-2 (Table 1 ). Forty-two missense mutations were identified in all the major non-structural and structural proteins, except the envelope protein. Twenty-nine missense mutations were in the ORF1ab polyprotein, eight in the spike surface glycoprotein, one in the matrix protein, and four in the nucleocapsid protein. Of note, three mutations (D354, Y364, and F367) located in the spike surface glycoprotein receptor-binding domain. The spike surface glycoprotein plays an essential role in binding to receptors on the host cell and determines host tropism (Fung and Liu, 2019). It is also the major target of neutralizing antibodies (Yu et al., 2020). Mutations in the spike surface glycoprotein might induce its conformational changes, which probably led to the changing antigenicity. To date, a study on localization of amino acids involved in conformational changes of the SARS-CoV-2 spike surface glycoprotein structure is not available. The identification of these amino acids is of significance and should be investigated by further studies.\nTable 1 Mutations found in the entire genome of SARS-CoV-2 strains. The number in the parentheses indicated the location of amino acid in its protein.\nGenomic region No. nt mutations Missense mutation SARV-CoV-2 strain\n5′ UTR 8 N/A\nORF1ab polyprotein 48 29\nA (117) → T USA/CA3/2020/EPI_ISL_408008USA/CA4/2020/EPI_ISL_408009\nP (309) → S France/IDF0515/2020/EPI ISL_408430\nS (428) → N USA/CA1/2020/EPI_ISL_406034\nT (609) → I USA/CA5/2020/EPI_ISL_408010\nA (1176) → V Japan/TY-WK-012/2020/EPI_ISL_408665\nL (1599) → F Korea/KCDC03/2020/EPI_ISL_407193\nI (1607) → V USA/CA3/2020/EPI_ISL_408008USA/CA4/2020/EPI_ISL_408009\nM (2194) → T Shenzhen/SZTH-004/2020/EPI_ISL_406595\nL (2235) → I Wuhan/WH01/2019/EPI_ISL_406798\nI (2244) → T Wuhan/IPBCAMS-WH-03/2019/EPI_ISL_403930\nG (2251) → S Wuhan/WIV05/2019/EPI_ISL_402128\nA (2345) → V Shandong/IVDC-SD-001/2020/EPI_ISL_408482\nG (2534) → V Wuhan/IPBCAMS-WH-05/2020/EPI_ISL_403928\nD (2579) → A Wuhan/WIV07/2019/EPI_ISL_402130\nN (2708) → S Wuhan/IPBCAMS-WH-01/2019/EPI_ISL_402123\nF (2908) → I Wuhan/IPBCAMS-WH-01/2019/EPI_ISL_402123\nT (3058) → I France/IDF0515/2020/EPI_ISL_408430\nS (3099) → L Shenzhen/HKU-SZ-005/2020/EPI_ISL_405839\nL (3606) → F Yunnan/IVDC-YN-003/2020/EPI_ISL_408480Shandong/IVDC-SD-001/2020/EPI_ISL_408482Chongqing/IVDC-CQ-001/2020/EPI_ISL_408481Singapore/3/2020/EPI_ISL_407988France/IDF0515/2020/EPI_ISL_408430USA/AZ1/2020/EPI_ISL_406223\nE (3764) → D Japan/KY-V-029/2020/EPI_ISL_408669\nN (3833) → K Wuhan/WH01/2019/EPI_ISL_406798\nW (5308) → C Taiwan/2/2020/EPI_ISL_406031\nT (5579) → I USA/CA2/2020/EPI_ISL_406036\nI (6075) → T England/02/2020/EPI_ISL_407073England/01/2020/EPI_ISL_407071\nP (6083) → L Japan/AI/I-004/2020/EPI_ISL_407084\nF (6309) → Y Sichuan/IVDC-SC-001/2020/EPI_ISL_408484\nE (6565) → D Shenzhen/SZTH-004/2020/EPI_ISL_406595\nK (6958) → R Wuhan/WIV05/2019/EPI_ISL_402128\nD (7018) → N Wuhan/WIV02/2019/EPI_ISL_402127\nSpike polyprotein 14 8\nF (32) → I Wuhan/HBCDC-HB-01/2019/EPI_ISL_402132\nH (49) → Y Guangdong/20SF174/2020/EPI_ISL_406531Guangdong/20SF040/2020/EPI_ISL_403937Guangdong/20SF028/2020/EPI_ISL_403936\nS (247) → R Australia/VIC01/2020/EPI_ISL_406844\nN (354) → D Shenzhen/SZTH-004/2020/EPI_ISL_406595\nD (364) → Y Shenzhen/SZTH-004/2020/EPI_ISL_406595\nV (367) → F France/IDF0372/2020/EPI_ISL_406596France/IDF0373/2020/EPI_ISL_406597\nD (614) → G Germany/BavPat1/2020/EPI_ISL_406862\nP (1143) → L Australia/QLD02/2020/EPI_ISL_407896\nIntergenic region 5 N/A\nEnvelope protein 0 0\nMatrix protein 2 1\nD (209) → H Singapore/2/2020/EPI_ISL_407987\nIntergenic region 6 N/A\nNucleocapsid protein 7 4\nT (148) → I Shenzhen/SZTH-004/2020/EPI_ISL_406595\nS (194) → L Shenzhen/SZTH-003/2020/EPI_ISL_406594Foshan/20SF207/2020/EPI_ISL_406534USA/CA3/2020/EPI_ISL_408008USA/CA4/2020/EPI_ISL_408009\nS (202) → N Australia/QLD02/2020/EPI_ISL_407896\nP (344) → S Guangzhou/20SF206/2020/EPI_ISL_406533\n3′UTR 3 N/A\nComplete genome 93 42\n\nAcknowledgement\nWe acknowledge support from Division of Clinical Microbiology, 10.13039/100007408University of Pittsburgh Medical Center.\n\nDeclaration of Competing Interest\nThe author declares no competing financial interests.","divisions":[{"label":"article-title","span":{"begin":0,"end":45}},{"label":"abstract","span":{"begin":56,"end":516}},{"label":"p","span":{"begin":56,"end":516}},{"label":"abstract","span":{"begin":518,"end":754}},{"label":"title","span":{"begin":518,"end":528}},{"label":"p","span":{"begin":529,"end":754}},{"label":"label","span":{"begin":529,"end":530}},{"label":"p","span":{"begin":532,"end":586}},{"label":"label","span":{"begin":587,"end":588}},{"label":"p","span":{"begin":590,"end":663}},{"label":"label","span":{"begin":664,"end":665}},{"label":"p","span":{"begin":667,"end":754}},{"label":"body","span":{"begin":756,"end":6937}},{"label":"sec","span":{"begin":756,"end":6937}},{"label":"label","span":{"begin":756,"end":757}},{"label":"title","span":{"begin":759,"end":768}},{"label":"p","span":{"begin":769,"end":1317}},{"label":"p","span":{"begin":1318,"end":2133}},{"label":"p","span":{"begin":2134,"end":2882}},{"label":"figure","span":{"begin":2721,"end":2882}},{"label":"label","span":{"begin":2721,"end":2727}},{"label":"caption","span":{"begin":2728,"end":2882}},{"label":"p","span":{"begin":2728,"end":2882}},{"label":"p","span":{"begin":2883,"end":6937}},{"label":"table-wrap","span":{"begin":4058,"end":6937}},{"label":"label","span":{"begin":4058,"end":4065}},{"label":"caption","span":{"begin":4066,"end":4208}},{"label":"p","span":{"begin":4066,"end":4208}},{"label":"table","span":{"begin":4209,"end":6937}},{"label":"tr","span":{"begin":4209,"end":4276}},{"label":"th","span":{"begin":4209,"end":4223}},{"label":"th","span":{"begin":4224,"end":4240}},{"label":"th","span":{"begin":4241,"end":4258}},{"label":"th","span":{"begin":4259,"end":4276}},{"label":"tr","span":{"begin":4277,"end":4289}},{"label":"td","span":{"begin":4277,"end":4283}},{"label":"td","span":{"begin":4284,"end":4285}},{"label":"td","span":{"begin":4286,"end":4289}},{"label":"tr","span":{"begin":4290,"end":4314}},{"label":"td","span":{"begin":4290,"end":4308}},{"label":"td","span":{"begin":4309,"end":4311}},{"label":"td","span":{"begin":4312,"end":4314}},{"label":"tr","span":{"begin":4315,"end":4381}},{"label":"td","span":{"begin":4315,"end":4326}},{"label":"td","span":{"begin":4327,"end":4381}},{"label":"tr","span":{"begin":4382,"end":4428}},{"label":"td","span":{"begin":4382,"end":4393}},{"label":"td","span":{"begin":4394,"end":4428}},{"label":"tr","span":{"begin":4429,"end":4468}},{"label":"td","span":{"begin":4429,"end":4440}},{"label":"td","span":{"begin":4441,"end":4468}},{"label":"tr","span":{"begin":4469,"end":4508}},{"label":"td","span":{"begin":4469,"end":4480}},{"label":"td","span":{"begin":4481,"end":4508}},{"label":"tr","span":{"begin":4509,"end":4557}},{"label":"td","span":{"begin":4509,"end":4521}},{"label":"td","span":{"begin":4522,"end":4557}},{"label":"tr","span":{"begin":4558,"end":4603}},{"label":"td","span":{"begin":4558,"end":4570}},{"label":"td","span":{"begin":4571,"end":4603}},{"label":"tr","span":{"begin":4604,"end":4671}},{"label":"td","span":{"begin":4604,"end":4616}},{"label":"td","span":{"begin":4617,"end":4671}},{"label":"tr","span":{"begin":4672,"end":4722}},{"label":"td","span":{"begin":4672,"end":4684}},{"label":"td","span":{"begin":4685,"end":4722}},{"label":"tr","span":{"begin":4723,"end":4766}},{"label":"td","span":{"begin":4723,"end":4735}},{"label":"td","span":{"begin":4736,"end":4766}},{"label":"tr","span":{"begin":4767,"end":4819}},{"label":"td","span":{"begin":4767,"end":4779}},{"label":"td","span":{"begin":4780,"end":4819}},{"label":"tr","span":{"begin":4820,"end":4864}},{"label":"td","span":{"begin":4820,"end":4832}},{"label":"td","span":{"begin":4833,"end":4864}},{"label":"tr","span":{"begin":4865,"end":4918}},{"label":"td","span":{"begin":4865,"end":4877}},{"label":"td","span":{"begin":4878,"end":4918}},{"label":"tr","span":{"begin":4919,"end":4971}},{"label":"td","span":{"begin":4919,"end":4931}},{"label":"td","span":{"begin":4932,"end":4971}},{"label":"tr","span":{"begin":4972,"end":5016}},{"label":"td","span":{"begin":4972,"end":4984}},{"label":"td","span":{"begin":4985,"end":5016}},{"label":"tr","span":{"begin":5017,"end":5069}},{"label":"td","span":{"begin":5017,"end":5029}},{"label":"td","span":{"begin":5030,"end":5069}},{"label":"tr","span":{"begin":5070,"end":5122}},{"label":"td","span":{"begin":5070,"end":5082}},{"label":"td","span":{"begin":5083,"end":5122}},{"label":"tr","span":{"begin":5123,"end":5170}},{"label":"td","span":{"begin":5123,"end":5135}},{"label":"td","span":{"begin":5136,"end":5170}},{"label":"tr","span":{"begin":5171,"end":5223}},{"label":"td","span":{"begin":5171,"end":5183}},{"label":"td","span":{"begin":5184,"end":5223}},{"label":"tr","span":{"begin":5224,"end":5448}},{"label":"td","span":{"begin":5224,"end":5236}},{"label":"td","span":{"begin":5237,"end":5448}},{"label":"tr","span":{"begin":5449,"end":5496}},{"label":"td","span":{"begin":5449,"end":5461}},{"label":"td","span":{"begin":5462,"end":5496}},{"label":"tr","span":{"begin":5497,"end":5540}},{"label":"td","span":{"begin":5497,"end":5509}},{"label":"td","span":{"begin":5510,"end":5540}},{"label":"tr","span":{"begin":5541,"end":5582}},{"label":"td","span":{"begin":5541,"end":5553}},{"label":"td","span":{"begin":5554,"end":5582}},{"label":"tr","span":{"begin":5583,"end":5623}},{"label":"td","span":{"begin":5583,"end":5595}},{"label":"td","span":{"begin":5596,"end":5623}},{"label":"tr","span":{"begin":5624,"end":5697}},{"label":"td","span":{"begin":5624,"end":5636}},{"label":"td","span":{"begin":5637,"end":5697}},{"label":"tr","span":{"begin":5698,"end":5745}},{"label":"td","span":{"begin":5698,"end":5710}},{"label":"td","span":{"begin":5711,"end":5745}},{"label":"tr","span":{"begin":5746,"end":5798}},{"label":"td","span":{"begin":5746,"end":5758}},{"label":"td","span":{"begin":5759,"end":5798}},{"label":"tr","span":{"begin":5799,"end":5849}},{"label":"td","span":{"begin":5799,"end":5811}},{"label":"td","span":{"begin":5812,"end":5849}},{"label":"tr","span":{"begin":5850,"end":5894}},{"label":"td","span":{"begin":5850,"end":5862}},{"label":"td","span":{"begin":5863,"end":5894}},{"label":"tr","span":{"begin":5895,"end":5939}},{"label":"td","span":{"begin":5895,"end":5907}},{"label":"td","span":{"begin":5908,"end":5939}},{"label":"tr","span":{"begin":5940,"end":5962}},{"label":"td","span":{"begin":5940,"end":5957}},{"label":"td","span":{"begin":5958,"end":5960}},{"label":"td","span":{"begin":5961,"end":5962}},{"label":"tr","span":{"begin":5963,"end":6011}},{"label":"td","span":{"begin":5963,"end":5973}},{"label":"td","span":{"begin":5974,"end":6011}},{"label":"tr","span":{"begin":6012,"end":6134}},{"label":"td","span":{"begin":6012,"end":6022}},{"label":"td","span":{"begin":6023,"end":6134}},{"label":"tr","span":{"begin":6135,"end":6182}},{"label":"td","span":{"begin":6135,"end":6146}},{"label":"td","span":{"begin":6147,"end":6182}},{"label":"tr","span":{"begin":6183,"end":6232}},{"label":"td","span":{"begin":6183,"end":6194}},{"label":"td","span":{"begin":6195,"end":6232}},{"label":"tr","span":{"begin":6233,"end":6282}},{"label":"td","span":{"begin":6233,"end":6244}},{"label":"td","span":{"begin":6245,"end":6282}},{"label":"tr","span":{"begin":6283,"end":6363}},{"label":"td","span":{"begin":6283,"end":6294}},{"label":"td","span":{"begin":6295,"end":6363}},{"label":"tr","span":{"begin":6364,"end":6411}},{"label":"td","span":{"begin":6364,"end":6375}},{"label":"td","span":{"begin":6376,"end":6411}},{"label":"tr","span":{"begin":6412,"end":6460}},{"label":"td","span":{"begin":6412,"end":6424}},{"label":"td","span":{"begin":6425,"end":6460}},{"label":"tr","span":{"begin":6461,"end":6484}},{"label":"td","span":{"begin":6461,"end":6478}},{"label":"td","span":{"begin":6479,"end":6480}},{"label":"td","span":{"begin":6481,"end":6484}},{"label":"tr","span":{"begin":6485,"end":6505}},{"label":"td","span":{"begin":6485,"end":6501}},{"label":"td","span":{"begin":6502,"end":6503}},{"label":"td","span":{"begin":6504,"end":6505}},{"label":"tr","span":{"begin":6506,"end":6524}},{"label":"td","span":{"begin":6506,"end":6520}},{"label":"td","span":{"begin":6521,"end":6522}},{"label":"td","span":{"begin":6523,"end":6524}},{"label":"tr","span":{"begin":6525,"end":6568}},{"label":"td","span":{"begin":6525,"end":6536}},{"label":"td","span":{"begin":6537,"end":6568}},{"label":"tr","span":{"begin":6569,"end":6592}},{"label":"td","span":{"begin":6569,"end":6586}},{"label":"td","span":{"begin":6587,"end":6588}},{"label":"td","span":{"begin":6589,"end":6592}},{"label":"tr","span":{"begin":6593,"end":6617}},{"label":"td","span":{"begin":6593,"end":6613}},{"label":"td","span":{"begin":6614,"end":6615}},{"label":"td","span":{"begin":6616,"end":6617}},{"label":"tr","span":{"begin":6618,"end":6667}},{"label":"td","span":{"begin":6618,"end":6629}},{"label":"td","span":{"begin":6630,"end":6667}},{"label":"tr","span":{"begin":6668,"end":6805}},{"label":"td","span":{"begin":6668,"end":6679}},{"label":"td","span":{"begin":6680,"end":6805}},{"label":"tr","span":{"begin":6806,"end":6853}},{"label":"td","span":{"begin":6806,"end":6817}},{"label":"td","span":{"begin":6818,"end":6853}},{"label":"tr","span":{"begin":6854,"end":6903}},{"label":"td","span":{"begin":6854,"end":6865}},{"label":"td","span":{"begin":6866,"end":6903}},{"label":"tr","span":{"begin":6904,"end":6915}},{"label":"td","span":{"begin":6904,"end":6909}},{"label":"td","span":{"begin":6910,"end":6911}},{"label":"td","span":{"begin":6912,"end":6915}},{"label":"tr","span":{"begin":6916,"end":6937}},{"label":"td","span":{"begin":6916,"end":6931}},{"label":"td","span":{"begin":6932,"end":6934}},{"label":"td","span":{"begin":6935,"end":6937}},{"label":"back","span":{"begin":6939,"end":7165}},{"label":"ack","span":{"begin":6939,"end":7165}},{"label":"sec","span":{"begin":6939,"end":7076}},{"label":"title","span":{"begin":6939,"end":6954}},{"label":"p","span":{"begin":6955,"end":7076}},{"label":"sec","span":{"begin":7078,"end":7165}},{"label":"title","span":{"begin":7078,"end":7111}},{"label":"p","span":{"begin":7112,"end":7165}}],"tracks":[{"project":"LitCovid-PubTator","denotations":[{"id":"7","span":{"begin":108,"end":123},"obj":"Species"},{"id":"8","span":{"begin":131,"end":141},"obj":"Species"},{"id":"9","span":{"begin":186,"end":196},"obj":"Species"},{"id":"10","span":{"begin":321,"end":331},"obj":"Species"},{"id":"11","span":{"begin":498,"end":515},"obj":"Species"},{"id":"12","span":{"begin":56,"end":64},"obj":"Disease"},{"id":"13","span":{"begin":76,"end":95},"obj":"Disease"},{"id":"16","span":{"begin":534,"end":551},"obj":"Species"},{"id":"17","span":{"begin":575,"end":585},"obj":"Species"},{"id":"19","span":{"begin":638,"end":648},"obj":"Species"},{"id":"21","span":{"begin":732,"end":742},"obj":"Species"},{"id":"28","span":{"begin":771,"end":786},"obj":"Species"},{"id":"29","span":{"begin":787,"end":797},"obj":"Species"},{"id":"30","span":{"begin":1026,"end":1031},"obj":"Species"},{"id":"31","span":{"begin":1035,"end":1040},"obj":"Species"},{"id":"32","span":{"begin":1057,"end":1067},"obj":"Species"},{"id":"33","span":{"begin":955,"end":963},"obj":"Disease"},{"id":"39","span":{"begin":1480,"end":1490},"obj":"Species"},{"id":"40","span":{"begin":1654,"end":1664},"obj":"Species"},{"id":"41","span":{"begin":1725,"end":1735},"obj":"Species"},{"id":"42","span":{"begin":1770,"end":1778},"obj":"Species"},{"id":"43","span":{"begin":1761,"end":1769},"obj":"Disease"},{"id":"46","span":{"begin":2831,"end":2849},"obj":"Gene"},{"id":"47","span":{"begin":2752,"end":2762},"obj":"Species"},{"id":"53","span":{"begin":2200,"end":2218},"obj":"Gene"},{"id":"54","span":{"begin":2633,"end":2651},"obj":"Gene"},{"id":"55","span":{"begin":2145,"end":2162},"obj":"Species"},{"id":"56","span":{"begin":2178,"end":2188},"obj":"Species"},{"id":"57","span":{"begin":2465,"end":2475},"obj":"Species"},{"id":"60","span":{"begin":4264,"end":4269},"obj":"Species"},{"id":"61","span":{"begin":5980,"end":5991},"obj":"CellLine"},{"id":"63","span":{"begin":4106,"end":4116},"obj":"Species"},{"id":"67","span":{"begin":3201,"end":3219},"obj":"Gene"},{"id":"68","span":{"begin":3004,"end":3014},"obj":"Species"},{"id":"69","span":{"begin":3886,"end":3896},"obj":"Species"}],"attributes":[{"id":"A7","pred":"tao:has_database_id","subj":"7","obj":"Tax:2697049"},{"id":"A8","pred":"tao:has_database_id","subj":"8","obj":"Tax:2697049"},{"id":"A9","pred":"tao:has_database_id","subj":"9","obj":"Tax:2697049"},{"id":"A10","pred":"tao:has_database_id","subj":"10","obj":"Tax:2697049"},{"id":"A11","pred":"tao:has_database_id","subj":"11","obj":"Tax:2697049"},{"id":"A12","pred":"tao:has_database_id","subj":"12","obj":"MESH:C000657245"},{"id":"A13","pred":"tao:has_database_id","subj":"13","obj":"MESH:D012140"},{"id":"A16","pred":"tao:has_database_id","subj":"16","obj":"Tax:2697049"},{"id":"A17","pred":"tao:has_database_id","subj":"17","obj":"Tax:2697049"},{"id":"A19","pred":"tao:has_database_id","subj":"19","obj":"Tax:2697049"},{"id":"A21","pred":"tao:has_database_id","subj":"21","obj":"Tax:2697049"},{"id":"A28","pred":"tao:has_database_id","subj":"28","obj":"Tax:2697049"},{"id":"A29","pred":"tao:has_database_id","subj":"29","obj":"Tax:2697049"},{"id":"A30","pred":"tao:has_database_id","subj":"30","obj":"Tax:9606"},{"id":"A31","pred":"tao:has_database_id","subj":"31","obj":"Tax:9606"},{"id":"A32","pred":"tao:has_database_id","subj":"32","obj":"Tax:2697049"},{"id":"A33","pred":"tao:has_database_id","subj":"33","obj":"MESH:D007239"},{"id":"A39","pred":"tao:has_database_id","subj":"39","obj":"Tax:2697049"},{"id":"A40","pred":"tao:has_database_id","subj":"40","obj":"Tax:2697049"},{"id":"A41","pred":"tao:has_database_id","subj":"41","obj":"Tax:2697049"},{"id":"A42","pred":"tao:has_database_id","subj":"42","obj":"Tax:9606"},{"id":"A43","pred":"tao:has_database_id","subj":"43","obj":"MESH:D007239"},{"id":"A46","pred":"tao:has_database_id","subj":"46","obj":"Gene:43740578"},{"id":"A47","pred":"tao:has_database_id","subj":"47","obj":"Tax:2697049"},{"id":"A53","pred":"tao:has_database_id","subj":"53","obj":"Gene:43740578"},{"id":"A54","pred":"tao:has_database_id","subj":"54","obj":"Gene:43740578"},{"id":"A55","pred":"tao:has_database_id","subj":"55","obj":"Tax:694002"},{"id":"A56","pred":"tao:has_database_id","subj":"56","obj":"Tax:2697049"},{"id":"A57","pred":"tao:has_database_id","subj":"57","obj":"Tax:2697049"},{"id":"A60","pred":"tao:has_database_id","subj":"60","obj":"Tax:2697049"},{"id":"A61","pred":"tao:has_database_id","subj":"61","obj":"CVCL:3694"},{"id":"A63","pred":"tao:has_database_id","subj":"63","obj":"Tax:2697049"},{"id":"A67","pred":"tao:has_database_id","subj":"67","obj":"Gene:43740578"},{"id":"A68","pred":"tao:has_database_id","subj":"68","obj":"Tax:2697049"},{"id":"A69","pred":"tao:has_database_id","subj":"69","obj":"Tax:2697049"},{"subj":"7","pred":"source","obj":"LitCovid-PubTator"},{"subj":"8","pred":"source","obj":"LitCovid-PubTator"},{"subj":"9","pred":"source","obj":"LitCovid-PubTator"},{"subj":"10","pred":"source","obj":"LitCovid-PubTator"},{"subj":"11","pred":"source","obj":"LitCovid-PubTator"},{"subj":"12","pred":"source","obj":"LitCovid-PubTator"},{"subj":"13","pred":"source","obj":"LitCovid-PubTator"},{"subj":"16","pred":"source","obj":"LitCovid-PubTator"},{"subj":"17","pred":"source","obj":"LitCovid-PubTator"},{"subj":"19","pred":"source","obj":"LitCovid-PubTator"},{"subj":"21","pred":"source","obj":"LitCovid-PubTator"},{"subj":"28","pred":"source","obj":"LitCovid-PubTator"},{"subj":"29","pred":"source","obj":"LitCovid-PubTator"},{"subj":"30","pred":"source","obj":"LitCovid-PubTator"},{"subj":"31","pred":"source","obj":"LitCovid-PubTator"},{"subj":"32","pred":"source","obj":"LitCovid-PubTator"},{"subj":"33","pred":"source","obj":"LitCovid-PubTator"},{"subj":"39","pred":"source","obj":"LitCovid-PubTator"},{"subj":"40","pred":"source","obj":"LitCovid-PubTator"},{"subj":"41","pred":"source","obj":"LitCovid-PubTator"},{"subj":"42","pred":"source","obj":"LitCovid-PubTator"},{"subj":"43","pred":"source","obj":"LitCovid-PubTator"},{"subj":"46","pred":"source","obj":"LitCovid-PubTator"},{"subj":"47","pred":"source","obj":"LitCovid-PubTator"},{"subj":"53","pred":"source","obj":"LitCovid-PubTator"},{"subj":"54","pred":"source","obj":"LitCovid-PubTator"},{"subj":"55","pred":"source","obj":"LitCovid-PubTator"},{"subj":"56","pred":"source","obj":"LitCovid-PubTator"},{"subj":"57","pred":"source","obj":"LitCovid-PubTator"},{"subj":"60","pred":"source","obj":"LitCovid-PubTator"},{"subj":"61","pred":"source","obj":"LitCovid-PubTator"},{"subj":"63","pred":"source","obj":"LitCovid-PubTator"},{"subj":"67","pred":"source","obj":"LitCovid-PubTator"},{"subj":"68","pred":"source","obj":"LitCovid-PubTator"},{"subj":"69","pred":"source","obj":"LitCovid-PubTator"}],"namespaces":[{"prefix":"Tax","uri":"https://www.ncbi.nlm.nih.gov/taxonomy/"},{"prefix":"MESH","uri":"https://id.nlm.nih.gov/mesh/"},{"prefix":"Gene","uri":"https://www.ncbi.nlm.nih.gov/gene/"},{"prefix":"CVCL","uri":"https://web.expasy.org/cellosaurus/CVCL_"}]},{"project":"2_test","denotations":[{"id":"32092483-32007627-47573167","span":{"begin":834,"end":838},"obj":"32007627"},{"id":"32092483-3447015-47573168","span":{"begin":2032,"end":2036},"obj":"3447015"},{"id":"32092483-32007627-47573169","span":{"begin":2392,"end":2396},"obj":"32007627"},{"id":"32092483-31226023-47573170","span":{"begin":3581,"end":3585},"obj":"31226023"},{"id":"T4386","span":{"begin":834,"end":838},"obj":"32007627"},{"id":"T9126","span":{"begin":2032,"end":2036},"obj":"3447015"},{"id":"T34600","span":{"begin":2392,"end":2396},"obj":"32007627"},{"id":"T3166","span":{"begin":3581,"end":3585},"obj":"31226023"}],"attributes":[{"subj":"32092483-32007627-47573167","pred":"source","obj":"2_test"},{"subj":"32092483-3447015-47573168","pred":"source","obj":"2_test"},{"subj":"32092483-32007627-47573169","pred":"source","obj":"2_test"},{"subj":"32092483-31226023-47573170","pred":"source","obj":"2_test"},{"subj":"T4386","pred":"source","obj":"2_test"},{"subj":"T9126","pred":"source","obj":"2_test"},{"subj":"T34600","pred":"source","obj":"2_test"},{"subj":"T3166","pred":"source","obj":"2_test"}]},{"project":"LitCovid-PD-FMA-UBERON","denotations":[{"id":"T1","span":{"begin":310,"end":317},"obj":"Body_part"},{"id":"T2","span":{"begin":627,"end":634},"obj":"Body_part"},{"id":"T3","span":{"begin":1186,"end":1192},"obj":"Body_part"},{"id":"T4","span":{"begin":1213,"end":1219},"obj":"Body_part"},{"id":"T5","span":{"begin":1244,"end":1250},"obj":"Body_part"},{"id":"T6","span":{"begin":1318,"end":1328},"obj":"Body_part"},{"id":"T7","span":{"begin":1643,"end":1650},"obj":"Body_part"},{"id":"T8","span":{"begin":1958,"end":1968},"obj":"Body_part"},{"id":"T9","span":{"begin":2126,"end":2132},"obj":"Body_part"},{"id":"T10","span":{"begin":2168,"end":2174},"obj":"Body_part"},{"id":"T11","span":{"begin":2268,"end":2276},"obj":"Body_part"},{"id":"T12","span":{"begin":2306,"end":2318},"obj":"Body_part"},{"id":"T13","span":{"begin":2335,"end":2342},"obj":"Body_part"},{"id":"T14","span":{"begin":2351,"end":2358},"obj":"Body_part"},{"id":"T15","span":{"begin":2377,"end":2384},"obj":"Body_part"},{"id":"T16","span":{"begin":2454,"end":2461},"obj":"Body_part"},{"id":"T17","span":{"begin":2580,"end":2591},"obj":"Body_part"},{"id":"T18","span":{"begin":2608,"end":2619},"obj":"Body_part"},{"id":"T19","span":{"begin":2675,"end":2686},"obj":"Body_part"},{"id":"T20","span":{"begin":2713,"end":2719},"obj":"Body_part"},{"id":"T21","span":{"begin":2776,"end":2786},"obj":"Body_part"},{"id":"T22","span":{"begin":2875,"end":2881},"obj":"Body_part"},{"id":"T23","span":{"begin":2910,"end":2920},"obj":"Body_part"},{"id":"T24","span":{"begin":2993,"end":3000},"obj":"Body_part"},{"id":"T25","span":{"begin":3119,"end":3127},"obj":"Body_part"},{"id":"T26","span":{"begin":3149,"end":3156},"obj":"Body_part"},{"id":"T27","span":{"begin":3248,"end":3260},"obj":"Body_part"},{"id":"T28","span":{"begin":3280,"end":3287},"obj":"Body_part"},{"id":"T29","span":{"begin":3318,"end":3325},"obj":"Body_part"},{"id":"T30","span":{"begin":3404,"end":3416},"obj":"Body_part"},{"id":"T31","span":{"begin":3460,"end":3472},"obj":"Body_part"},{"id":"T32","span":{"begin":3533,"end":3537},"obj":"Body_part"},{"id":"T33","span":{"begin":3693,"end":3705},"obj":"Body_part"},{"id":"T34","span":{"begin":3832,"end":3843},"obj":"Body_part"},{"id":"T35","span":{"begin":3911,"end":3923},"obj":"Body_part"},{"id":"T36","span":{"begin":3980,"end":3991},"obj":"Body_part"},{"id":"T37","span":{"begin":4096,"end":4102},"obj":"Body_part"},{"id":"T38","span":{"begin":4182,"end":4192},"obj":"Body_part"},{"id":"T39","span":{"begin":4200,"end":4207},"obj":"Body_part"},{"id":"T40","span":{"begin":6494,"end":6501},"obj":"Body_part"},{"id":"T41","span":{"begin":6513,"end":6520},"obj":"Body_part"},{"id":"T42","span":{"begin":6606,"end":6613},"obj":"Body_part"},{"id":"T43","span":{"begin":6925,"end":6931},"obj":"Body_part"}],"attributes":[{"id":"A1","pred":"fma_id","subj":"T1","obj":"http://purl.org/sig/ont/fma/fma84116"},{"id":"A2","pred":"fma_id","subj":"T2","obj":"http://purl.org/sig/ont/fma/fma84116"},{"id":"A3","pred":"fma_id","subj":"T3","obj":"http://purl.org/sig/ont/fma/fma312401"},{"id":"A4","pred":"fma_id","subj":"T4","obj":"http://purl.org/sig/ont/fma/fma59862"},{"id":"A5","pred":"fma_id","subj":"T5","obj":"http://purl.org/sig/ont/fma/fma228738"},{"id":"A6","pred":"fma_id","subj":"T6","obj":"http://purl.org/sig/ont/fma/fma82740"},{"id":"A7","pred":"fma_id","subj":"T7","obj":"http://purl.org/sig/ont/fma/fma84116"},{"id":"A8","pred":"fma_id","subj":"T8","obj":"http://purl.org/sig/ont/fma/fma82740"},{"id":"A9","pred":"fma_id","subj":"T9","obj":"http://purl.org/sig/ont/fma/fma84116"},{"id":"A10","pred":"fma_id","subj":"T10","obj":"http://purl.org/sig/ont/fma/fma84116"},{"id":"A11","pred":"fma_id","subj":"T11","obj":"http://purl.org/sig/ont/fma/fma67257"},{"id":"A12","pred":"fma_id","subj":"T12","obj":"http://purl.org/sig/ont/fma/fma62925"},{"id":"A13","pred":"fma_id","subj":"T13","obj":"http://purl.org/sig/ont/fma/fma67257"},{"id":"A14","pred":"fma_id","subj":"T14","obj":"http://purl.org/sig/ont/fma/fma67257"},{"id":"A15","pred":"fma_id","subj":"T15","obj":"http://purl.org/sig/ont/fma/fma67257"},{"id":"A16","pred":"fma_id","subj":"T16","obj":"http://purl.org/sig/ont/fma/fma84116"},{"id":"A17","pred":"fma_id","subj":"T17","obj":"http://purl.org/sig/ont/fma/fma82740"},{"id":"A18","pred":"fma_id","subj":"T18","obj":"http://purl.org/sig/ont/fma/fma82740"},{"id":"A19","pred":"fma_id","subj":"T19","obj":"http://purl.org/sig/ont/fma/fma82740"},{"id":"A20","pred":"fma_id","subj":"T20","obj":"http://purl.org/sig/ont/fma/fma84116"},{"id":"A21","pred":"fma_id","subj":"T21","obj":"http://purl.org/sig/ont/fma/fma82740"},{"id":"A22","pred":"fma_id","subj":"T22","obj":"http://purl.org/sig/ont/fma/fma84116"},{"id":"A23","pred":"fma_id","subj":"T23","obj":"http://purl.org/sig/ont/fma/fma82740"},{"id":"A24","pred":"fma_id","subj":"T24","obj":"http://purl.org/sig/ont/fma/fma84116"},{"id":"A25","pred":"fma_id","subj":"T25","obj":"http://purl.org/sig/ont/fma/fma67257"},{"id":"A26","pred":"fma_id","subj":"T26","obj":"http://purl.org/sig/ont/fma/fma67257"},{"id":"A27","pred":"fma_id","subj":"T27","obj":"http://purl.org/sig/ont/fma/fma62925"},{"id":"A28","pred":"fma_id","subj":"T28","obj":"http://purl.org/sig/ont/fma/fma67257"},{"id":"A29","pred":"fma_id","subj":"T29","obj":"http://purl.org/sig/ont/fma/fma67257"},{"id":"A30","pred":"fma_id","subj":"T30","obj":"http://purl.org/sig/ont/fma/fma62925"},{"id":"A31","pred":"fma_id","subj":"T31","obj":"http://purl.org/sig/ont/fma/fma62925"},{"id":"A32","pred":"fma_id","subj":"T32","obj":"http://purl.org/sig/ont/fma/fma68646"},{"id":"A33","pred":"fma_id","subj":"T33","obj":"http://purl.org/sig/ont/fma/fma62925"},{"id":"A34","pred":"fma_id","subj":"T34","obj":"http://purl.org/sig/ont/fma/fma82739"},{"id":"A35","pred":"fma_id","subj":"T35","obj":"http://purl.org/sig/ont/fma/fma62925"},{"id":"A36","pred":"fma_id","subj":"T36","obj":"http://purl.org/sig/ont/fma/fma82739"},{"id":"A37","pred":"fma_id","subj":"T37","obj":"http://purl.org/sig/ont/fma/fma84116"},{"id":"A38","pred":"fma_id","subj":"T38","obj":"http://purl.org/sig/ont/fma/fma82739"},{"id":"A39","pred":"fma_id","subj":"T39","obj":"http://purl.org/sig/ont/fma/fma67257"},{"id":"A40","pred":"fma_id","subj":"T40","obj":"http://purl.org/sig/ont/fma/fma67257"},{"id":"A41","pred":"fma_id","subj":"T41","obj":"http://purl.org/sig/ont/fma/fma67257"},{"id":"A42","pred":"fma_id","subj":"T42","obj":"http://purl.org/sig/ont/fma/fma67257"},{"id":"A43","pred":"fma_id","subj":"T43","obj":"http://purl.org/sig/ont/fma/fma84116"},{"subj":"T1","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T2","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T3","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T4","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T5","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T6","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T7","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T8","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T9","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T10","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T11","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T12","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T13","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T14","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T15","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T16","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T17","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T18","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T19","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T20","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T21","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T22","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T23","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T24","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T25","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T26","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T27","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T28","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T29","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T30","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T31","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T32","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T33","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T34","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T35","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T36","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T37","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T38","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T39","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T40","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T41","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T42","pred":"source","obj":"LitCovid-PD-FMA-UBERON"},{"subj":"T43","pred":"source","obj":"LitCovid-PD-FMA-UBERON"}]},{"project":"LitCovid-PD-UBERON","denotations":[{"id":"T1","span":{"begin":1186,"end":1192},"obj":"Body_part"},{"id":"T2","span":{"begin":1213,"end":1219},"obj":"Body_part"},{"id":"T3","span":{"begin":1244,"end":1250},"obj":"Body_part"}],"attributes":[{"id":"A1","pred":"uberon_id","subj":"T1","obj":"http://purl.obolibrary.org/obo/UBERON_0007311"},{"id":"A2","pred":"uberon_id","subj":"T2","obj":"http://purl.obolibrary.org/obo/UBERON_0001836"},{"id":"A3","pred":"uberon_id","subj":"T3","obj":"http://purl.obolibrary.org/obo/UBERON_0000341"},{"subj":"T1","pred":"source","obj":"LitCovid-PD-UBERON"},{"subj":"T2","pred":"source","obj":"LitCovid-PD-UBERON"},{"subj":"T3","pred":"source","obj":"LitCovid-PD-UBERON"}]},{"project":"LitCovid_AGAC","denotations":[{"id":"p63588s9","span":{"begin":3723,"end":3745},"obj":"MPA"}],"attributes":[{"subj":"p63588s9","pred":"source","obj":"LitCovid_AGAC"}]},{"project":"LitCovid-PD-MONDO","denotations":[{"id":"T1","span":{"begin":35,"end":43},"obj":"Disease"},{"id":"T2","span":{"begin":56,"end":64},"obj":"Disease"},{"id":"T3","span":{"begin":131,"end":139},"obj":"Disease"},{"id":"T4","span":{"begin":186,"end":194},"obj":"Disease"},{"id":"T5","span":{"begin":321,"end":329},"obj":"Disease"},{"id":"T6","span":{"begin":575,"end":583},"obj":"Disease"},{"id":"T7","span":{"begin":638,"end":646},"obj":"Disease"},{"id":"T8","span":{"begin":732,"end":740},"obj":"Disease"},{"id":"T9","span":{"begin":787,"end":795},"obj":"Disease"},{"id":"T10","span":{"begin":1057,"end":1065},"obj":"Disease"},{"id":"T11","span":{"begin":1480,"end":1488},"obj":"Disease"},{"id":"T12","span":{"begin":1654,"end":1662},"obj":"Disease"},{"id":"T13","span":{"begin":1725,"end":1733},"obj":"Disease"},{"id":"T14","span":{"begin":2178,"end":2186},"obj":"Disease"},{"id":"T15","span":{"begin":2465,"end":2473},"obj":"Disease"},{"id":"T16","span":{"begin":2752,"end":2760},"obj":"Disease"},{"id":"T17","span":{"begin":3004,"end":3012},"obj":"Disease"},{"id":"T18","span":{"begin":3886,"end":3894},"obj":"Disease"},{"id":"T19","span":{"begin":4106,"end":4114},"obj":"Disease"}],"attributes":[{"id":"A1","pred":"mondo_id","subj":"T1","obj":"http://purl.obolibrary.org/obo/MONDO_0005091"},{"id":"A2","pred":"mondo_id","subj":"T2","obj":"http://purl.obolibrary.org/obo/MONDO_0100096"},{"id":"A3","pred":"mondo_id","subj":"T3","obj":"http://purl.obolibrary.org/obo/MONDO_0005091"},{"id":"A4","pred":"mondo_id","subj":"T4","obj":"http://purl.obolibrary.org/obo/MONDO_0005091"},{"id":"A5","pred":"mondo_id","subj":"T5","obj":"http://purl.obolibrary.org/obo/MONDO_0005091"},{"id":"A6","pred":"mondo_id","subj":"T6","obj":"http://purl.obolibrary.org/obo/MONDO_0005091"},{"id":"A7","pred":"mondo_id","subj":"T7","obj":"http://purl.obolibrary.org/obo/MONDO_0005091"},{"id":"A8","pred":"mondo_id","subj":"T8","obj":"http://purl.obolibrary.org/obo/MONDO_0005091"},{"id":"A9","pred":"mondo_id","subj":"T9","obj":"http://purl.obolibrary.org/obo/MONDO_0005091"},{"id":"A10","pred":"mondo_id","subj":"T10","obj":"http://purl.obolibrary.org/obo/MONDO_0005091"},{"id":"A11","pred":"mondo_id","subj":"T11","obj":"http://purl.obolibrary.org/obo/MONDO_0005091"},{"id":"A12","pred":"mondo_id","subj":"T12","obj":"http://purl.obolibrary.org/obo/MONDO_0005091"},{"id":"A13","pred":"mondo_id","subj":"T13","obj":"http://purl.obolibrary.org/obo/MONDO_0005091"},{"id":"A14","pred":"mondo_id","subj":"T14","obj":"http://purl.obolibrary.org/obo/MONDO_0005091"},{"id":"A15","pred":"mondo_id","subj":"T15","obj":"http://purl.obolibrary.org/obo/MONDO_0005091"},{"id":"A16","pred":"mondo_id","subj":"T16","obj":"http://purl.obolibrary.org/obo/MONDO_0005091"},{"id":"A17","pred":"mondo_id","subj":"T17","obj":"http://purl.obolibrary.org/obo/MONDO_0005091"},{"id":"A18","pred":"mondo_id","subj":"T18","obj":"http://purl.obolibrary.org/obo/MONDO_0005091"},{"id":"A19","pred":"mondo_id","subj":"T19","obj":"http://purl.obolibrary.org/obo/MONDO_0005091"},{"subj":"T1","pred":"source","obj":"LitCovid-PD-MONDO"},{"subj":"T2","pred":"source","obj":"LitCovid-PD-MONDO"},{"subj":"T3","pred":"source","obj":"LitCovid-PD-MONDO"},{"subj":"T4","pred":"source","obj":"LitCovid-PD-MONDO"},{"subj":"T5","pred":"source","obj":"LitCovid-PD-MONDO"},{"subj":"T6","pred":"source","obj":"LitCovid-PD-MONDO"},{"subj":"T7","pred":"source","obj":"LitCovid-PD-MONDO"},{"subj":"T8","pred":"source","obj":"LitCovid-PD-MONDO"},{"subj":"T9","pred":"source","obj":"LitCovid-PD-MONDO"},{"subj":"T10","pred":"source","obj":"LitCovid-PD-MONDO"},{"subj":"T11","pred":"source","obj":"LitCovid-PD-MONDO"},{"subj":"T12","pred":"source","obj":"LitCovid-PD-MONDO"},{"subj":"T13","pred":"source","obj":"LitCovid-PD-MONDO"},{"subj":"T14","pred":"source","obj":"LitCovid-PD-MONDO"},{"subj":"T15","pred":"source","obj":"LitCovid-PD-MONDO"},{"subj":"T16","pred":"source","obj":"LitCovid-PD-MONDO"},{"subj":"T17","pred":"source","obj":"LitCovid-PD-MONDO"},{"subj":"T18","pred":"source","obj":"LitCovid-PD-MONDO"},{"subj":"T19","pred":"source","obj":"LitCovid-PD-MONDO"}]},{"project":"LitCovid-PD-CLO","denotations":[{"id":"T1","span":{"begin":68,"end":69},"obj":"http://purl.obolibrary.org/obo/CLO_0001020"},{"id":"T2","span":{"begin":106,"end":107},"obj":"http://purl.obolibrary.org/obo/CLO_0001020"},{"id":"T3","span":{"begin":160,"end":172},"obj":"http://purl.obolibrary.org/obo/OBI_0000245"},{"id":"T4","span":{"begin":206,"end":207},"obj":"http://purl.obolibrary.org/obo/CLO_0001020"},{"id":"T5","span":{"begin":532,"end":533},"obj":"http://purl.obolibrary.org/obo/CLO_0001020"},{"id":"T6","span":{"begin":769,"end":770},"obj":"http://purl.obolibrary.org/obo/CLO_0001020"},{"id":"T7","span":{"begin":851,"end":856},"obj":"http://purl.obolibrary.org/obo/NCBITaxon_10239"},{"id":"T8","span":{"begin":970,"end":973},"obj":"http://purl.obolibrary.org/obo/CLO_0051582"},{"id":"T9","span":{"begin":1026,"end":1031},"obj":"http://purl.obolibrary.org/obo/NCBITaxon_9606"},{"id":"T10","span":{"begin":1035,"end":1040},"obj":"http://purl.obolibrary.org/obo/NCBITaxon_9606"},{"id":"T11","span":{"begin":1068,"end":1071},"obj":"http://purl.obolibrary.org/obo/CLO_0051582"},{"id":"T12","span":{"begin":1116,"end":1121},"obj":"http://purl.obolibrary.org/obo/NCBITaxon_10239"},{"id":"T13","span":{"begin":1122,"end":1125},"obj":"http://purl.obolibrary.org/obo/CLO_0051582"},{"id":"T14","span":{"begin":1342,"end":1345},"obj":"http://purl.obolibrary.org/obo/CLO_0051582"},{"id":"T15","span":{"begin":1801,"end":1803},"obj":"http://purl.obolibrary.org/obo/CLO_0053733"},{"id":"T16","span":{"begin":2114,"end":2115},"obj":"http://purl.obolibrary.org/obo/CLO_0001020"},{"id":"T17","span":{"begin":2189,"end":2192},"obj":"http://purl.obolibrary.org/obo/CLO_0051582"},{"id":"T18","span":{"begin":2193,"end":2194},"obj":"http://purl.obolibrary.org/obo/CLO_0001020"},{"id":"T19","span":{"begin":2671,"end":2674},"obj":"http://purl.obolibrary.org/obo/CLO_0050884"},{"id":"T20","span":{"begin":2736,"end":2748},"obj":"http://purl.obolibrary.org/obo/OBI_0000245"},{"id":"T21","span":{"begin":3533,"end":3537},"obj":"http://purl.obolibrary.org/obo/GO_0005623"},{"id":"T22","span":{"begin":3805,"end":3806},"obj":"http://purl.obolibrary.org/obo/CLO_0001020"},{"id":"T23","span":{"begin":4288,"end":4289},"obj":"http://purl.obolibrary.org/obo/CLO_0001020"},{"id":"T24","span":{"begin":4309,"end":4311},"obj":"http://purl.obolibrary.org/obo/CLO_0001382"},{"id":"T25","span":{"begin":4315,"end":4316},"obj":"http://purl.obolibrary.org/obo/CLO_0001020"},{"id":"T26","span":{"begin":4414,"end":4417},"obj":"http://purl.obolibrary.org/obo/CLO_0002941"},{"id":"T27","span":{"begin":4509,"end":4510},"obj":"http://purl.obolibrary.org/obo/CLO_0001020"},{"id":"T28","span":{"begin":4865,"end":4866},"obj":"http://purl.obolibrary.org/obo/CLO_0001020"},{"id":"T29","span":{"begin":4983,"end":4984},"obj":"http://purl.obolibrary.org/obo/CLO_0001020"},{"id":"T30","span":{"begin":5772,"end":5774},"obj":"http://purl.obolibrary.org/obo/CLO_0008954"},{"id":"T31","span":{"begin":5958,"end":5962},"obj":"http://purl.obolibrary.org/obo/CLO_0001079"},{"id":"T32","span":{"begin":5986,"end":5988},"obj":"http://purl.obolibrary.org/obo/CLO_0003622"},{"id":"T33","span":{"begin":6483,"end":6484},"obj":"http://purl.obolibrary.org/obo/CLO_0001020"},{"id":"T34","span":{"begin":6591,"end":6592},"obj":"http://purl.obolibrary.org/obo/CLO_0001020"},{"id":"T35","span":{"begin":6621,"end":6624},"obj":"http://purl.obolibrary.org/obo/CLO_0001079"},{"id":"T36","span":{"begin":6914,"end":6915},"obj":"http://purl.obolibrary.org/obo/CLO_0001020"}],"attributes":[{"subj":"T1","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T2","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T3","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T4","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T5","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T6","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T7","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T8","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T9","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T10","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T11","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T12","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T13","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T14","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T15","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T16","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T17","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T18","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T19","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T20","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T21","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T22","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T23","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T24","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T25","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T26","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T27","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T28","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T29","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T30","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T31","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T32","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T33","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T34","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T35","pred":"source","obj":"LitCovid-PD-CLO"},{"subj":"T36","pred":"source","obj":"LitCovid-PD-CLO"}]},{"project":"LitCovid-PD-CHEBI","denotations":[{"id":"T1","span":{"begin":1194,"end":1197},"obj":"Chemical"},{"id":"T2","span":{"begin":1318,"end":1328},"obj":"Chemical"},{"id":"T3","span":{"begin":1958,"end":1968},"obj":"Chemical"},{"id":"T4","span":{"begin":2268,"end":2276},"obj":"Chemical"},{"id":"T5","span":{"begin":2306,"end":2318},"obj":"Chemical"},{"id":"T6","span":{"begin":2335,"end":2342},"obj":"Chemical"},{"id":"T7","span":{"begin":2351,"end":2358},"obj":"Chemical"},{"id":"T8","span":{"begin":2377,"end":2384},"obj":"Chemical"},{"id":"T9","span":{"begin":2580,"end":2591},"obj":"Chemical"},{"id":"T10","span":{"begin":2608,"end":2619},"obj":"Chemical"},{"id":"T11","span":{"begin":2675,"end":2686},"obj":"Chemical"},{"id":"T12","span":{"begin":2776,"end":2786},"obj":"Chemical"},{"id":"T13","span":{"begin":2910,"end":2920},"obj":"Chemical"},{"id":"T14","span":{"begin":3119,"end":3127},"obj":"Chemical"},{"id":"T15","span":{"begin":3149,"end":3156},"obj":"Chemical"},{"id":"T16","span":{"begin":3248,"end":3260},"obj":"Chemical"},{"id":"T17","span":{"begin":3280,"end":3287},"obj":"Chemical"},{"id":"T18","span":{"begin":3318,"end":3325},"obj":"Chemical"},{"id":"T19","span":{"begin":3404,"end":3416},"obj":"Chemical"},{"id":"T20","span":{"begin":3460,"end":3472},"obj":"Chemical"},{"id":"T21","span":{"begin":3693,"end":3705},"obj":"Chemical"},{"id":"T22","span":{"begin":3832,"end":3843},"obj":"Chemical"},{"id":"T23","span":{"begin":3832,"end":3837},"obj":"Chemical"},{"id":"T24","span":{"begin":3838,"end":3843},"obj":"Chemical"},{"id":"T25","span":{"begin":3911,"end":3923},"obj":"Chemical"},{"id":"T26","span":{"begin":3980,"end":3991},"obj":"Chemical"},{"id":"T27","span":{"begin":3980,"end":3985},"obj":"Chemical"},{"id":"T28","span":{"begin":3986,"end":3991},"obj":"Chemical"},{"id":"T29","span":{"begin":4182,"end":4192},"obj":"Chemical"},{"id":"T30","span":{"begin":4182,"end":4187},"obj":"Chemical"},{"id":"T31","span":{"begin":4188,"end":4192},"obj":"Chemical"},{"id":"T32","span":{"begin":4200,"end":4207},"obj":"Chemical"},{"id":"T33","span":{"begin":4528,"end":4530},"obj":"Chemical"},{"id":"T34","span":{"begin":4531,"end":4533},"obj":"Chemical"},{"id":"T35","span":{"begin":4892,"end":4894},"obj":"Chemical"},{"id":"T36","span":{"begin":5249,"end":5251},"obj":"Chemical"},{"id":"T37","span":{"begin":5289,"end":5291},"obj":"Chemical"},{"id":"T38","span":{"begin":5468,"end":5470},"obj":"Chemical"},{"id":"T39","span":{"begin":5717,"end":5719},"obj":"Chemical"},{"id":"T42","span":{"begin":6494,"end":6501},"obj":"Chemical"},{"id":"T43","span":{"begin":6513,"end":6520},"obj":"Chemical"},{"id":"T44","span":{"begin":6606,"end":6613},"obj":"Chemical"}],"attributes":[{"id":"A1","pred":"chebi_id","subj":"T1","obj":"http://purl.obolibrary.org/obo/CHEBI_32386"},{"id":"A2","pred":"chebi_id","subj":"T2","obj":"http://purl.obolibrary.org/obo/CHEBI_36976"},{"id":"A3","pred":"chebi_id","subj":"T3","obj":"http://purl.obolibrary.org/obo/CHEBI_36976"},{"id":"A4","pred":"chebi_id","subj":"T4","obj":"http://purl.obolibrary.org/obo/CHEBI_36080"},{"id":"A5","pred":"chebi_id","subj":"T5","obj":"http://purl.obolibrary.org/obo/CHEBI_17089"},{"id":"A6","pred":"chebi_id","subj":"T6","obj":"http://purl.obolibrary.org/obo/CHEBI_36080"},{"id":"A7","pred":"chebi_id","subj":"T7","obj":"http://purl.obolibrary.org/obo/CHEBI_36080"},{"id":"A8","pred":"chebi_id","subj":"T8","obj":"http://purl.obolibrary.org/obo/CHEBI_36080"},{"id":"A9","pred":"chebi_id","subj":"T9","obj":"http://purl.obolibrary.org/obo/CHEBI_36976"},{"id":"A10","pred":"chebi_id","subj":"T10","obj":"http://purl.obolibrary.org/obo/CHEBI_36976"},{"id":"A11","pred":"chebi_id","subj":"T11","obj":"http://purl.obolibrary.org/obo/CHEBI_36976"},{"id":"A12","pred":"chebi_id","subj":"T12","obj":"http://purl.obolibrary.org/obo/CHEBI_36976"},{"id":"A13","pred":"chebi_id","subj":"T13","obj":"http://purl.obolibrary.org/obo/CHEBI_36976"},{"id":"A14","pred":"chebi_id","subj":"T14","obj":"http://purl.obolibrary.org/obo/CHEBI_36080"},{"id":"A15","pred":"chebi_id","subj":"T15","obj":"http://purl.obolibrary.org/obo/CHEBI_36080"},{"id":"A16","pred":"chebi_id","subj":"T16","obj":"http://purl.obolibrary.org/obo/CHEBI_17089"},{"id":"A17","pred":"chebi_id","subj":"T17","obj":"http://purl.obolibrary.org/obo/CHEBI_36080"},{"id":"A18","pred":"chebi_id","subj":"T18","obj":"http://purl.obolibrary.org/obo/CHEBI_36080"},{"id":"A19","pred":"chebi_id","subj":"T19","obj":"http://purl.obolibrary.org/obo/CHEBI_17089"},{"id":"A20","pred":"chebi_id","subj":"T20","obj":"http://purl.obolibrary.org/obo/CHEBI_17089"},{"id":"A21","pred":"chebi_id","subj":"T21","obj":"http://purl.obolibrary.org/obo/CHEBI_17089"},{"id":"A22","pred":"chebi_id","subj":"T22","obj":"http://purl.obolibrary.org/obo/CHEBI_33709"},{"id":"A23","pred":"chebi_id","subj":"T23","obj":"http://purl.obolibrary.org/obo/CHEBI_46882"},{"id":"A24","pred":"chebi_id","subj":"T24","obj":"http://purl.obolibrary.org/obo/CHEBI_37527"},{"id":"A25","pred":"chebi_id","subj":"T25","obj":"http://purl.obolibrary.org/obo/CHEBI_17089"},{"id":"A26","pred":"chebi_id","subj":"T26","obj":"http://purl.obolibrary.org/obo/CHEBI_33709"},{"id":"A27","pred":"chebi_id","subj":"T27","obj":"http://purl.obolibrary.org/obo/CHEBI_46882"},{"id":"A28","pred":"chebi_id","subj":"T28","obj":"http://purl.obolibrary.org/obo/CHEBI_37527"},{"id":"A29","pred":"chebi_id","subj":"T29","obj":"http://purl.obolibrary.org/obo/CHEBI_33709"},{"id":"A30","pred":"chebi_id","subj":"T30","obj":"http://purl.obolibrary.org/obo/CHEBI_46882"},{"id":"A31","pred":"chebi_id","subj":"T31","obj":"http://purl.obolibrary.org/obo/CHEBI_37527"},{"id":"A32","pred":"chebi_id","subj":"T32","obj":"http://purl.obolibrary.org/obo/CHEBI_36080"},{"id":"A33","pred":"chebi_id","subj":"T33","obj":"http://purl.obolibrary.org/obo/CHEBI_73667"},{"id":"A34","pred":"chebi_id","subj":"T34","obj":"http://purl.obolibrary.org/obo/CHEBI_74872"},{"id":"A35","pred":"chebi_id","subj":"T35","obj":"http://purl.obolibrary.org/obo/CHEBI_74807"},{"id":"A36","pred":"chebi_id","subj":"T36","obj":"http://purl.obolibrary.org/obo/CHEBI_141452"},{"id":"A37","pred":"chebi_id","subj":"T37","obj":"http://purl.obolibrary.org/obo/CHEBI_74807"},{"id":"A38","pred":"chebi_id","subj":"T38","obj":"http://purl.obolibrary.org/obo/CHEBI_73608"},{"id":"A39","pred":"chebi_id","subj":"T39","obj":"http://purl.obolibrary.org/obo/CHEBI_73805"},{"id":"A40","pred":"chebi_id","subj":"T39","obj":"http://purl.obolibrary.org/obo/CHEBI_74007"},{"id":"A41","pred":"chebi_id","subj":"T39","obj":"http://purl.obolibrary.org/obo/CHEBI_74088"},{"id":"A42","pred":"chebi_id","subj":"T42","obj":"http://purl.obolibrary.org/obo/CHEBI_36080"},{"id":"A43","pred":"chebi_id","subj":"T43","obj":"http://purl.obolibrary.org/obo/CHEBI_36080"},{"id":"A44","pred":"chebi_id","subj":"T44","obj":"http://purl.obolibrary.org/obo/CHEBI_36080"},{"subj":"T1","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T2","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T3","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T4","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T5","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T6","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T7","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T8","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T9","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T10","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T11","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T12","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T13","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T14","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T15","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T16","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T17","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T18","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T19","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T20","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T21","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T22","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T23","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T24","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T25","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T26","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T27","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T28","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T29","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T30","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T31","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T32","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T33","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T34","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T35","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T36","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T37","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T38","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T39","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T42","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T43","pred":"source","obj":"LitCovid-PD-CHEBI"},{"subj":"T44","pred":"source","obj":"LitCovid-PD-CHEBI"}]},{"project":"LitCovid-PD-GO-BP","denotations":[{"id":"T1","span":{"begin":3558,"end":3565},"obj":"http://purl.obolibrary.org/obo/GO_0009606"},{"id":"T2","span":{"begin":3816,"end":3828},"obj":"http://purl.obolibrary.org/obo/GO_0051179"}],"attributes":[{"subj":"T1","pred":"source","obj":"LitCovid-PD-GO-BP"},{"subj":"T2","pred":"source","obj":"LitCovid-PD-GO-BP"}]},{"project":"LitCovid-sentences","denotations":[{"id":"T1","span":{"begin":0,"end":45},"obj":"Sentence"},{"id":"T2","span":{"begin":47,"end":55},"obj":"Sentence"},{"id":"T3","span":{"begin":56,"end":142},"obj":"Sentence"},{"id":"T4","span":{"begin":143,"end":239},"obj":"Sentence"},{"id":"T5","span":{"begin":240,"end":407},"obj":"Sentence"},{"id":"T6","span":{"begin":408,"end":516},"obj":"Sentence"},{"id":"T7","span":{"begin":518,"end":528},"obj":"Sentence"},{"id":"T8","span":{"begin":529,"end":586},"obj":"Sentence"},{"id":"T9","span":{"begin":587,"end":663},"obj":"Sentence"},{"id":"T10","span":{"begin":664,"end":754},"obj":"Sentence"},{"id":"T11","span":{"begin":756,"end":768},"obj":"Sentence"},{"id":"T12","span":{"begin":769,"end":840},"obj":"Sentence"},{"id":"T13","span":{"begin":841,"end":1025},"obj":"Sentence"},{"id":"T14","span":{"begin":1026,"end":1111},"obj":"Sentence"},{"id":"T15","span":{"begin":1112,"end":1225},"obj":"Sentence"},{"id":"T16","span":{"begin":1226,"end":1317},"obj":"Sentence"},{"id":"T17","span":{"begin":1318,"end":1459},"obj":"Sentence"},{"id":"T18","span":{"begin":1460,"end":1572},"obj":"Sentence"},{"id":"T19","span":{"begin":1573,"end":1718},"obj":"Sentence"},{"id":"T20","span":{"begin":1719,"end":1943},"obj":"Sentence"},{"id":"T21","span":{"begin":1944,"end":2133},"obj":"Sentence"},{"id":"T22","span":{"begin":2134,"end":2398},"obj":"Sentence"},{"id":"T23","span":{"begin":2399,"end":2558},"obj":"Sentence"},{"id":"T24","span":{"begin":2559,"end":2720},"obj":"Sentence"},{"id":"T25","span":{"begin":2721,"end":2882},"obj":"Sentence"},{"id":"T26","span":{"begin":2883,"end":3026},"obj":"Sentence"},{"id":"T27","span":{"begin":3027,"end":3157},"obj":"Sentence"},{"id":"T28","span":{"begin":3158,"end":3326},"obj":"Sentence"},{"id":"T29","span":{"begin":3327,"end":3441},"obj":"Sentence"},{"id":"T30","span":{"begin":3442,"end":3587},"obj":"Sentence"},{"id":"T31","span":{"begin":3588,"end":3661},"obj":"Sentence"},{"id":"T32","span":{"begin":3662,"end":3795},"obj":"Sentence"},{"id":"T33","span":{"begin":3796,"end":3951},"obj":"Sentence"},{"id":"T34","span":{"begin":3952,"end":4057},"obj":"Sentence"},{"id":"T35","span":{"begin":4058,"end":4125},"obj":"Sentence"},{"id":"T36","span":{"begin":4126,"end":4208},"obj":"Sentence"},{"id":"T37","span":{"begin":4209,"end":4276},"obj":"Sentence"},{"id":"T38","span":{"begin":4277,"end":4289},"obj":"Sentence"},{"id":"T39","span":{"begin":4290,"end":4314},"obj":"Sentence"},{"id":"T40","span":{"begin":4315,"end":4381},"obj":"Sentence"},{"id":"T41","span":{"begin":4382,"end":4428},"obj":"Sentence"},{"id":"T42","span":{"begin":4429,"end":4468},"obj":"Sentence"},{"id":"T43","span":{"begin":4469,"end":4508},"obj":"Sentence"},{"id":"T44","span":{"begin":4509,"end":4557},"obj":"Sentence"},{"id":"T45","span":{"begin":4558,"end":4603},"obj":"Sentence"},{"id":"T46","span":{"begin":4604,"end":4671},"obj":"Sentence"},{"id":"T47","span":{"begin":4672,"end":4722},"obj":"Sentence"},{"id":"T48","span":{"begin":4723,"end":4766},"obj":"Sentence"},{"id":"T49","span":{"begin":4767,"end":4819},"obj":"Sentence"},{"id":"T50","span":{"begin":4820,"end":4864},"obj":"Sentence"},{"id":"T51","span":{"begin":4865,"end":4918},"obj":"Sentence"},{"id":"T52","span":{"begin":4919,"end":4971},"obj":"Sentence"},{"id":"T53","span":{"begin":4972,"end":5016},"obj":"Sentence"},{"id":"T54","span":{"begin":5017,"end":5069},"obj":"Sentence"},{"id":"T55","span":{"begin":5070,"end":5122},"obj":"Sentence"},{"id":"T56","span":{"begin":5123,"end":5170},"obj":"Sentence"},{"id":"T57","span":{"begin":5171,"end":5223},"obj":"Sentence"},{"id":"T58","span":{"begin":5224,"end":5448},"obj":"Sentence"},{"id":"T59","span":{"begin":5449,"end":5496},"obj":"Sentence"},{"id":"T60","span":{"begin":5497,"end":5540},"obj":"Sentence"},{"id":"T61","span":{"begin":5541,"end":5582},"obj":"Sentence"},{"id":"T62","span":{"begin":5583,"end":5623},"obj":"Sentence"},{"id":"T63","span":{"begin":5624,"end":5697},"obj":"Sentence"},{"id":"T64","span":{"begin":5698,"end":5745},"obj":"Sentence"},{"id":"T65","span":{"begin":5746,"end":5798},"obj":"Sentence"},{"id":"T66","span":{"begin":5799,"end":5849},"obj":"Sentence"},{"id":"T67","span":{"begin":5850,"end":5894},"obj":"Sentence"},{"id":"T68","span":{"begin":5895,"end":5939},"obj":"Sentence"},{"id":"T69","span":{"begin":5940,"end":5962},"obj":"Sentence"},{"id":"T70","span":{"begin":5963,"end":6011},"obj":"Sentence"},{"id":"T71","span":{"begin":6012,"end":6134},"obj":"Sentence"},{"id":"T72","span":{"begin":6135,"end":6182},"obj":"Sentence"},{"id":"T73","span":{"begin":6183,"end":6232},"obj":"Sentence"},{"id":"T74","span":{"begin":6233,"end":6282},"obj":"Sentence"},{"id":"T75","span":{"begin":6283,"end":6363},"obj":"Sentence"},{"id":"T76","span":{"begin":6364,"end":6411},"obj":"Sentence"},{"id":"T77","span":{"begin":6412,"end":6460},"obj":"Sentence"},{"id":"T78","span":{"begin":6461,"end":6484},"obj":"Sentence"},{"id":"T79","span":{"begin":6485,"end":6505},"obj":"Sentence"},{"id":"T80","span":{"begin":6506,"end":6524},"obj":"Sentence"},{"id":"T81","span":{"begin":6525,"end":6568},"obj":"Sentence"},{"id":"T82","span":{"begin":6569,"end":6592},"obj":"Sentence"},{"id":"T83","span":{"begin":6593,"end":6617},"obj":"Sentence"},{"id":"T84","span":{"begin":6618,"end":6667},"obj":"Sentence"},{"id":"T85","span":{"begin":6668,"end":6805},"obj":"Sentence"},{"id":"T86","span":{"begin":6806,"end":6853},"obj":"Sentence"},{"id":"T87","span":{"begin":6854,"end":6903},"obj":"Sentence"},{"id":"T88","span":{"begin":6904,"end":6915},"obj":"Sentence"},{"id":"T89","span":{"begin":6916,"end":6937},"obj":"Sentence"},{"id":"T90","span":{"begin":6939,"end":6954},"obj":"Sentence"},{"id":"T91","span":{"begin":6955,"end":7076},"obj":"Sentence"},{"id":"T92","span":{"begin":7078,"end":7111},"obj":"Sentence"},{"id":"T93","span":{"begin":7112,"end":7165},"obj":"Sentence"}],"namespaces":[{"prefix":"_base","uri":"http://pubannotation.org/ontology/tao.owl#"}],"attributes":[{"subj":"T1","pred":"source","obj":"LitCovid-sentences"},{"subj":"T2","pred":"source","obj":"LitCovid-sentences"},{"subj":"T3","pred":"source","obj":"LitCovid-sentences"},{"subj":"T4","pred":"source","obj":"LitCovid-sentences"},{"subj":"T5","pred":"source","obj":"LitCovid-sentences"},{"subj":"T6","pred":"source","obj":"LitCovid-sentences"},{"subj":"T7","pred":"source","obj":"LitCovid-sentences"},{"subj":"T8","pred":"source","obj":"LitCovid-sentences"},{"subj":"T9","pred":"source","obj":"LitCovid-sentences"},{"subj":"T10","pred":"source","obj":"LitCovid-sentences"},{"subj":"T11","pred":"source","obj":"LitCovid-sentences"},{"subj":"T12","pred":"source","obj":"LitCovid-sentences"},{"subj":"T13","pred":"source","obj":"LitCovid-sentences"},{"subj":"T14","pred":"source","obj":"LitCovid-sentences"},{"subj":"T15","pred":"source","obj":"LitCovid-sentences"},{"subj":"T16","pred":"source","obj":"LitCovid-sentences"},{"subj":"T17","pred":"source","obj":"LitCovid-sentences"},{"subj":"T18","pred":"source","obj":"LitCovid-sentences"},{"subj":"T19","pred":"source","obj":"LitCovid-sentences"},{"subj":"T20","pred":"source","obj":"LitCovid-sentences"},{"subj":"T21","pred":"source","obj":"LitCovid-sentences"},{"subj":"T22","pred":"source","obj":"LitCovid-sentences"},{"subj":"T23","pred":"source","obj":"LitCovid-sentences"},{"subj":"T24","pred":"source","obj":"LitCovid-sentences"},{"subj":"T25","pred":"source","obj":"LitCovid-sentences"},{"subj":"T26","pred":"source","obj":"LitCovid-sentences"},{"subj":"T27","pred":"source","obj":"LitCovid-sentences"},{"subj":"T28","pred":"source","obj":"LitCovid-sentences"},{"subj":"T29","pred":"source","obj":"LitCovid-sentences"},{"subj":"T30","pred":"source","obj":"LitCovid-sentences"},{"subj":"T31","pred":"source","obj":"LitCovid-sentences"},{"subj":"T32","pred":"source","obj":"LitCovid-sentences"},{"subj":"T33","pred":"source","obj":"LitCovid-sentences"},{"subj":"T34","pred":"source","obj":"LitCovid-sentences"},{"subj":"T35","pred":"source","obj":"LitCovid-sentences"},{"subj":"T36","pred":"source","obj":"LitCovid-sentences"},{"subj":"T37","pred":"source","obj":"LitCovid-sentences"},{"subj":"T38","pred":"source","obj":"LitCovid-sentences"},{"subj":"T39","pred":"source","obj":"LitCovid-sentences"},{"subj":"T40","pred":"source","obj":"LitCovid-sentences"},{"subj":"T41","pred":"source","obj":"LitCovid-sentences"},{"subj":"T42","pred":"source","obj":"LitCovid-sentences"},{"subj":"T43","pred":"source","obj":"LitCovid-sentences"},{"subj":"T44","pred":"source","obj":"LitCovid-sentences"},{"subj":"T45","pred":"source","obj":"LitCovid-sentences"},{"subj":"T46","pred":"source","obj":"LitCovid-sentences"},{"subj":"T47","pred":"source","obj":"LitCovid-sentences"},{"subj":"T48","pred":"source","obj":"LitCovid-sentences"},{"subj":"T49","pred":"source","obj":"LitCovid-sentences"},{"subj":"T50","pred":"source","obj":"LitCovid-sentences"},{"subj":"T51","pred":"source","obj":"LitCovid-sentences"},{"subj":"T52","pred":"source","obj":"LitCovid-sentences"},{"subj":"T53","pred":"source","obj":"LitCovid-sentences"},{"subj":"T54","pred":"source","obj":"LitCovid-sentences"},{"subj":"T55","pred":"source","obj":"LitCovid-sentences"},{"subj":"T56","pred":"source","obj":"LitCovid-sentences"},{"subj":"T57","pred":"source","obj":"LitCovid-sentences"},{"subj":"T58","pred":"source","obj":"LitCovid-sentences"},{"subj":"T59","pred":"source","obj":"LitCovid-sentences"},{"subj":"T60","pred":"source","obj":"LitCovid-sentences"},{"subj":"T61","pred":"source","obj":"LitCovid-sentences"},{"subj":"T62","pred":"source","obj":"LitCovid-sentences"},{"subj":"T63","pred":"source","obj":"LitCovid-sentences"},{"subj":"T64","pred":"source","obj":"LitCovid-sentences"},{"subj":"T65","pred":"source","obj":"LitCovid-sentences"},{"subj":"T66","pred":"source","obj":"LitCovid-sentences"},{"subj":"T67","pred":"source","obj":"LitCovid-sentences"},{"subj":"T68","pred":"source","obj":"LitCovid-sentences"},{"subj":"T69","pred":"source","obj":"LitCovid-sentences"},{"subj":"T70","pred":"source","obj":"LitCovid-sentences"},{"subj":"T71","pred":"source","obj":"LitCovid-sentences"},{"subj":"T72","pred":"source","obj":"LitCovid-sentences"},{"subj":"T73","pred":"source","obj":"LitCovid-sentences"},{"subj":"T74","pred":"source","obj":"LitCovid-sentences"},{"subj":"T75","pred":"source","obj":"LitCovid-sentences"},{"subj":"T76","pred":"source","obj":"LitCovid-sentences"},{"subj":"T77","pred":"source","obj":"LitCovid-sentences"},{"subj":"T78","pred":"source","obj":"LitCovid-sentences"},{"subj":"T79","pred":"source","obj":"LitCovid-sentences"},{"subj":"T80","pred":"source","obj":"LitCovid-sentences"},{"subj":"T81","pred":"source","obj":"LitCovid-sentences"},{"subj":"T82","pred":"source","obj":"LitCovid-sentences"},{"subj":"T83","pred":"source","obj":"LitCovid-sentences"},{"subj":"T84","pred":"source","obj":"LitCovid-sentences"},{"subj":"T85","pred":"source","obj":"LitCovid-sentences"},{"subj":"T86","pred":"source","obj":"LitCovid-sentences"},{"subj":"T87","pred":"source","obj":"LitCovid-sentences"},{"subj":"T88","pred":"source","obj":"LitCovid-sentences"},{"subj":"T89","pred":"source","obj":"LitCovid-sentences"},{"subj":"T90","pred":"source","obj":"LitCovid-sentences"},{"subj":"T91","pred":"source","obj":"LitCovid-sentences"},{"subj":"T92","pred":"source","obj":"LitCovid-sentences"},{"subj":"T93","pred":"source","obj":"LitCovid-sentences"}]},{"project":"LitCovid-PD-HP","denotations":[{"id":"T1","span":{"begin":76,"end":95},"obj":"Phenotype"}],"attributes":[{"id":"A1","pred":"hp_id","subj":"T1","obj":"http://purl.obolibrary.org/obo/HP_0002086"},{"subj":"T1","pred":"source","obj":"LitCovid-PD-HP"}]}],"config":{"attribute types":[{"pred":"source","value type":"selection","values":[{"id":"LitCovid-PubTator","color":"#93b0ec","default":true},{"id":"2_test","color":"#caec93"},{"id":"LitCovid-PD-FMA-UBERON","color":"#ec93e4"},{"id":"LitCovid-PD-UBERON","color":"#93ecda"},{"id":"LitCovid_AGAC","color":"#ecc093"},{"id":"LitCovid-PD-MONDO","color":"#a593ec"},{"id":"LitCovid-PD-CLO","color":"#9aec93"},{"id":"LitCovid-PD-CHEBI","color":"#ec93b4"},{"id":"LitCovid-PD-GO-BP","color":"#93ceec"},{"id":"LitCovid-sentences","color":"#e8ec93"},{"id":"LitCovid-PD-HP","color":"#d693ec"}]}]}}