PubMed:32198201 JSONTXT

{"target":"https://pubannotation.org/docs/sourcedb/PubMed/sourceid/32198201","sourcedb":"PubMed","sourceid":"32198201","source_url":"https://www.ncbi.nlm.nih.gov/pubmed/32198201","text":"Coronavirus endoribonuclease targets viral polyuridine sequences to evade activating host sensors.\nCoronaviruses (CoVs) are positive-sense RNA viruses that can emerge from endemic reservoirs and infect zoonotically, causing significant morbidity and mortality. CoVs encode an endoribonuclease designated EndoU that facilitates evasion of host pattern recognition receptor MDA5, but the target of EndoU activity was not known. Here, we report that EndoU cleaves the 5'-polyuridines from negative-sense viral RNA, termed PUN RNA, which is the product of polyA-templated RNA synthesis. Using a virus containing an EndoU catalytic-inactive mutation, we detected a higher abundance of PUN RNA in the cytoplasm compared to wild-type-infected cells. Furthermore, we found that transfecting PUN RNA into cells stimulates a robust, MDA5-dependent interferon response, and that removal of the polyuridine extension on the RNA dampens the response. Overall, the results of this study reveal the PUN RNA to be a CoV MDA5-dependent pathogen-associated molecular pattern (PAMP). We also establish a mechanism for EndoU activity to cleave and limit the accumulation of this PAMP. Since EndoU activity is highly conserved in all CoVs, inhibiting this activity may serve as an approach for therapeutic interventions against existing and emerging CoV infections.","tracks":[{"project":"LitCovid-OGER","denotations":[{"id":"T1","span":{"begin":568,"end":581},"obj":"GO:0032774"},{"id":"T1","span":{"begin":37,"end":42},"obj":"GO:0019012"},{"id":"T2","span":{"begin":85,"end":89},"obj":"GO:0018995"},{"id":"T3","span":{"begin":338,"end":342},"obj":"GO:0018995"},{"id":"T4","span":{"begin":501,"end":506},"obj":"GO:0019012"},{"id":"T5","span":{"begin":591,"end":596},"obj":"GO:0019012"},{"id":"T6","span":{"begin":695,"end":704},"obj":"GO:0005737"},{"id":"T1","span":{"begin":304,"end":309},"obj":"PR:000030363"},{"id":"T2","span":{"begin":372,"end":376},"obj":"PR:000008910"},{"id":"T3","span":{"begin":396,"end":401},"obj":"PR:000030363"},{"id":"T4","span":{"begin":447,"end":452},"obj":"PR:000030363"},{"id":"T5","span":{"begin":611,"end":616},"obj":"PR:000030363"},{"id":"T6","span":{"begin":823,"end":827},"obj":"PR:000008910"},{"id":"T7","span":{"begin":1004,"end":1008},"obj":"PR:000008910"},{"id":"T8","span":{"begin":1058,"end":1062},"obj":"PR:000017540"},{"id":"T9","span":{"begin":1099,"end":1104},"obj":"PR:000030363"},{"id":"T10","span":{"begin":1159,"end":1163},"obj":"PR:000017540"},{"id":"T11","span":{"begin":1171,"end":1176},"obj":"PR:000030363"},{"id":"T1","span":{"begin":120,"end":123},"obj":"SO:0001853"},{"id":"T2","span":{"begin":995,"end":997},"obj":"SO:0000999"},{"id":"T1","span":{"begin":895,"end":904},"obj":"UBERON:2000106"}],"namespaces":[{"prefix":"PR","uri":"http://purl.obolibrary.org/obo/PR_"},{"prefix":"UBERON","uri":"http://purl.obolibrary.org/obo/UBERON_"}],"attributes":[{"subj":"T1","pred":"source","obj":"LitCovid-OGER"},{"subj":"T1","pred":"source","obj":"LitCovid-OGER"},{"subj":"T2","pred":"source","obj":"LitCovid-OGER"},{"subj":"T3","pred":"source","obj":"LitCovid-OGER"},{"subj":"T4","pred":"source","obj":"LitCovid-OGER"},{"subj":"T5","pred":"source","obj":"LitCovid-OGER"},{"subj":"T6","pred":"source","obj":"LitCovid-OGER"},{"subj":"T1","pred":"source","obj":"LitCovid-OGER"},{"subj":"T2","pred":"source","obj":"LitCovid-OGER"},{"subj":"T3","pred":"source","obj":"LitCovid-OGER"},{"subj":"T4","pred":"source","obj":"LitCovid-OGER"},{"subj":"T5","pred":"source","obj":"LitCovid-OGER"},{"subj":"T6","pred":"source","obj":"LitCovid-OGER"},{"subj":"T7","pred":"source","obj":"LitCovid-OGER"},{"subj":"T8","pred":"source","obj":"LitCovid-OGER"},{"subj":"T9","pred":"source","obj":"LitCovid-OGER"},{"subj":"T10","pred":"source","obj":"LitCovid-OGER"},{"subj":"T11","pred":"source","obj":"LitCovid-OGER"},{"subj":"T1","pred":"source","obj":"LitCovid-OGER"},{"subj":"T2","pred":"source","obj":"LitCovid-OGER"},{"subj":"T1","pred":"source","obj":"LitCovid-OGER"}]},{"project":"LitCovid-OGER-BB","denotations":[{"id":"T1","span":{"begin":0,"end":11},"obj":"NCBITaxon:11118"},{"id":"T2","span":{"begin":37,"end":42},"obj":"NCBITaxon:10239"},{"id":"T3","span":{"begin":99,"end":112},"obj":"NCBITaxon:11118"},{"id":"T4","span":{"begin":114,"end":118},"obj":"NCBITaxon:11118"},{"id":"T5","span":{"begin":143,"end":150},"obj":"NCBITaxon:10239"},{"id":"T6","span":{"begin":304,"end":309},"obj":"PR:000030363"},{"id":"T7","span":{"begin":372,"end":376},"obj":"PR:000008910"},{"id":"T8","span":{"begin":396,"end":401},"obj":"PR:000030363"},{"id":"T9","span":{"begin":453,"end":460},"obj":"MOP:0000780"},{"id":"T10","span":{"begin":501,"end":506},"obj":"NCBITaxon:10239"},{"id":"T11","span":{"begin":568,"end":581},"obj":"GO:0032774"},{"id":"T12","span":{"begin":591,"end":596},"obj":"NCBITaxon:10239"},{"id":"T13","span":{"begin":695,"end":704},"obj":"GO:0005737"},{"id":"T14","span":{"begin":770,"end":782},"obj":"GO:0009294"},{"id":"T15","span":{"begin":823,"end":827},"obj":"PR:000008910"},{"id":"T16","span":{"begin":1004,"end":1008},"obj":"PR:000008910"},{"id":"T17","span":{"begin":1058,"end":1062},"obj":"PR:000017540"},{"id":"T18","span":{"begin":1117,"end":1123},"obj":"MOP:0000780"},{"id":"T19","span":{"begin":1159,"end":1163},"obj":"PR:000017540"}],"namespaces":[{"prefix":"NCBITaxon","uri":"https://www.ncbi.nlm.nih.gov/taxonomy/"}],"attributes":[{"subj":"T1","pred":"source","obj":"LitCovid-OGER-BB"},{"subj":"T2","pred":"source","obj":"LitCovid-OGER-BB"},{"subj":"T3","pred":"source","obj":"LitCovid-OGER-BB"},{"subj":"T4","pred":"source","obj":"LitCovid-OGER-BB"},{"subj":"T5","pred":"source","obj":"LitCovid-OGER-BB"},{"subj":"T6","pred":"source","obj":"LitCovid-OGER-BB"},{"subj":"T7","pred":"source","obj":"LitCovid-OGER-BB"},{"subj":"T8","pred":"source","obj":"LitCovid-OGER-BB"},{"subj":"T9","pred":"source","obj":"LitCovid-OGER-BB"},{"subj":"T10","pred":"source","obj":"LitCovid-OGER-BB"},{"subj":"T11","pred":"source","obj":"LitCovid-OGER-BB"},{"subj":"T12","pred":"source","obj":"LitCovid-OGER-BB"},{"subj":"T13","pred":"source","obj":"LitCovid-OGER-BB"},{"subj":"T14","pred":"source","obj":"LitCovid-OGER-BB"},{"subj":"T15","pred":"source","obj":"LitCovid-OGER-BB"},{"subj":"T16","pred":"source","obj":"LitCovid-OGER-BB"},{"subj":"T17","pred":"source","obj":"LitCovid-OGER-BB"},{"subj":"T18","pred":"source","obj":"LitCovid-OGER-BB"},{"subj":"T19","pred":"source","obj":"LitCovid-OGER-BB"}]},{"project":"LitCovid-sentences-v1","denotations":[{"id":"TextSentencer_T1","span":{"begin":0,"end":98},"obj":"Sentence"},{"id":"TextSentencer_T2","span":{"begin":99,"end":260},"obj":"Sentence"},{"id":"TextSentencer_T3","span":{"begin":261,"end":425},"obj":"Sentence"},{"id":"TextSentencer_T4","span":{"begin":426,"end":582},"obj":"Sentence"},{"id":"TextSentencer_T5","span":{"begin":583,"end":742},"obj":"Sentence"},{"id":"TextSentencer_T6","span":{"begin":743,"end":937},"obj":"Sentence"},{"id":"TextSentencer_T7","span":{"begin":938,"end":1064},"obj":"Sentence"},{"id":"TextSentencer_T8","span":{"begin":1065,"end":1164},"obj":"Sentence"},{"id":"TextSentencer_T9","span":{"begin":1165,"end":1344},"obj":"Sentence"}],"attributes":[{"subj":"TextSentencer_T1","pred":"source","obj":"LitCovid-sentences-v1"},{"subj":"TextSentencer_T2","pred":"source","obj":"LitCovid-sentences-v1"},{"subj":"TextSentencer_T3","pred":"source","obj":"LitCovid-sentences-v1"},{"subj":"TextSentencer_T4","pred":"source","obj":"LitCovid-sentences-v1"},{"subj":"TextSentencer_T5","pred":"source","obj":"LitCovid-sentences-v1"},{"subj":"TextSentencer_T6","pred":"source","obj":"LitCovid-sentences-v1"},{"subj":"TextSentencer_T7","pred":"source","obj":"LitCovid-sentences-v1"},{"subj":"TextSentencer_T8","pred":"source","obj":"LitCovid-sentences-v1"},{"subj":"TextSentencer_T9","pred":"source","obj":"LitCovid-sentences-v1"}]},{"project":"LitCovid-TimeML","denotations":[{"id":"tok1","span":{"begin":0,"end":11},"obj":"NNP"},{"id":"tok2","span":{"begin":12,"end":28},"obj":"NN"},{"id":"tok3","span":{"begin":29,"end":36},"obj":"NNS"},{"id":"tok4","span":{"begin":37,"end":42},"obj":"JJ"},{"id":"tok5","span":{"begin":43,"end":54},"obj":"NN"},{"id":"tok6","span":{"begin":55,"end":64},"obj":"NNS"},{"id":"tok7","span":{"begin":65,"end":67},"obj":"TO"},{"id":"tok8","span":{"begin":68,"end":73},"obj":"VB"},{"id":"tok9","span":{"begin":74,"end":84},"obj":"VBG"},{"id":"tok10","span":{"begin":85,"end":89},"obj":"NN"},{"id":"tok11","span":{"begin":90,"end":97},"obj":"NNS"},{"id":"tok12","span":{"begin":97,"end":98},"obj":"."},{"id":"tok13","span":{"begin":99,"end":112},"obj":"NNP"},{"id":"tok14","span":{"begin":113,"end":114},"obj":"("},{"id":"tok15","span":{"begin":114,"end":118},"obj":"NNP"},{"id":"tok16","span":{"begin":118,"end":119},"obj":")"},{"id":"tok17","span":{"begin":120,"end":123},"obj":"VBP"},{"id":"tok18","span":{"begin":124,"end":138},"obj":"JJ"},{"id":"tok19","span":{"begin":139,"end":142},"obj":"NNP"},{"id":"tok20","span":{"begin":143,"end":150},"obj":"NNS"},{"id":"tok21","span":{"begin":151,"end":155},"obj":"WDT"},{"id":"tok22","span":{"begin":156,"end":159},"obj":"MD"},{"id":"tok23","span":{"begin":160,"end":166},"obj":"VB"},{"id":"tok24","span":{"begin":167,"end":171},"obj":"IN"},{"id":"tok25","span":{"begin":172,"end":179},"obj":"JJ"},{"id":"tok26","span":{"begin":180,"end":190},"obj":"NNS"},{"id":"tok27","span":{"begin":191,"end":194},"obj":"CC"},{"id":"tok28","span":{"begin":195,"end":201},"obj":"NN"},{"id":"tok29","span":{"begin":202,"end":214},"obj":"RB"},{"id":"tok30","span":{"begin":214,"end":215},"obj":","},{"id":"tok31","span":{"begin":216,"end":223},"obj":"VBG"},{"id":"tok32","span":{"begin":224,"end":235},"obj":"JJ"},{"id":"tok33","span":{"begin":236,"end":245},"obj":"NN"},{"id":"tok34","span":{"begin":246,"end":249},"obj":"CC"},{"id":"tok35","span":{"begin":250,"end":259},"obj":"NN"},{"id":"tok36","span":{"begin":259,"end":260},"obj":"."},{"id":"tok37","span":{"begin":261,"end":265},"obj":"NNP"},{"id":"tok38","span":{"begin":266,"end":272},"obj":"NN"},{"id":"tok39","span":{"begin":273,"end":275},"obj":"DT"},{"id":"tok40","span":{"begin":276,"end":292},"obj":"NN"},{"id":"tok41","span":{"begin":293,"end":303},"obj":"VBN"},{"id":"tok42","span":{"begin":304,"end":309},"obj":"NNP"},{"id":"tok43","span":{"begin":310,"end":314},"obj":"IN"},{"id":"tok44","span":{"begin":315,"end":326},"obj":"NNS"},{"id":"tok45","span":{"begin":327,"end":334},"obj":"NN"},{"id":"tok46","span":{"begin":335,"end":337},"obj":"IN"},{"id":"tok47","span":{"begin":338,"end":342},"obj":"NN"},{"id":"tok48","span":{"begin":343,"end":350},"obj":"NN"},{"id":"tok49","span":{"begin":351,"end":362},"obj":"NN"},{"id":"tok50","span":{"begin":363,"end":371},"obj":"NN"},{"id":"tok51","span":{"begin":372,"end":375},"obj":"NNP"},{"id":"tok52","span":{"begin":375,"end":376},"obj":"CD"},{"id":"tok53","span":{"begin":376,"end":377},"obj":","},{"id":"tok54","span":{"begin":378,"end":381},"obj":"CC"},{"id":"tok55","span":{"begin":382,"end":385},"obj":"DT"},{"id":"tok56","span":{"begin":386,"end":392},"obj":"NN"},{"id":"tok57","span":{"begin":393,"end":395},"obj":"IN"},{"id":"tok58","span":{"begin":396,"end":401},"obj":"NNP"},{"id":"tok59","span":{"begin":402,"end":410},"obj":"NN"},{"id":"tok60","span":{"begin":411,"end":414},"obj":"VBD"},{"id":"tok61","span":{"begin":415,"end":418},"obj":"RB"},{"id":"tok62","span":{"begin":419,"end":424},"obj":"VBN"},{"id":"tok63","span":{"begin":424,"end":425},"obj":"."},{"id":"tok64","span":{"begin":426,"end":430},"obj":"RB"},{"id":"tok65","span":{"begin":430,"end":431},"obj":","},{"id":"tok66","span":{"begin":432,"end":434},"obj":"PRP"},{"id":"tok67","span":{"begin":435,"end":441},"obj":"VBP"},{"id":"tok68","span":{"begin":442,"end":446},"obj":"IN"},{"id":"tok69","span":{"begin":447,"end":452},"obj":"NNP"},{"id":"tok70","span":{"begin":453,"end":460},"obj":"NNS"},{"id":"tok71","span":{"begin":461,"end":464},"obj":"DT"},{"id":"tok72","span":{"begin":465,"end":466},"obj":"CD"},{"id":"tok73","span":{"begin":466,"end":467},"obj":"POS"},{"id":"tok74","span":{"begin":467,"end":468},"obj":":"},{"id":"tok75","span":{"begin":468,"end":480},"obj":"NNS"},{"id":"tok76","span":{"begin":481,"end":485},"obj":"IN"},{"id":"tok77","span":{"begin":486,"end":500},"obj":"JJ"},{"id":"tok78","span":{"begin":501,"end":506},"obj":"JJ"},{"id":"tok79","span":{"begin":507,"end":510},"obj":"NNP"},{"id":"tok80","span":{"begin":510,"end":511},"obj":","},{"id":"tok81","span":{"begin":512,"end":518},"obj":"VBD"},{"id":"tok82","span":{"begin":519,"end":522},"obj":"NNP"},{"id":"tok83","span":{"begin":523,"end":526},"obj":"NNP"},{"id":"tok84","span":{"begin":526,"end":527},"obj":","},{"id":"tok85","span":{"begin":528,"end":533},"obj":"WDT"},{"id":"tok86","span":{"begin":534,"end":536},"obj":"VBZ"},{"id":"tok87","span":{"begin":537,"end":540},"obj":"DT"},{"id":"tok88","span":{"begin":541,"end":548},"obj":"NN"},{"id":"tok89","span":{"begin":549,"end":551},"obj":"IN"},{"id":"tok90","span":{"begin":552,"end":567},"obj":"JJ"},{"id":"tok91","span":{"begin":568,"end":571},"obj":"NNP"},{"id":"tok92","span":{"begin":572,"end":581},"obj":"NN"},{"id":"tok93","span":{"begin":581,"end":582},"obj":"."},{"id":"tok94","span":{"begin":583,"end":588},"obj":"VBG"},{"id":"tok95","span":{"begin":589,"end":590},"obj":"DT"},{"id":"tok96","span":{"begin":591,"end":596},"obj":"NN"},{"id":"tok97","span":{"begin":597,"end":607},"obj":"VBG"},{"id":"tok98","span":{"begin":608,"end":610},"obj":"DT"},{"id":"tok99","span":{"begin":611,"end":616},"obj":"NNP"},{"id":"tok100","span":{"begin":617,"end":635},"obj":"JJ"},{"id":"tok101","span":{"begin":636,"end":644},"obj":"NN"},{"id":"tok102","span":{"begin":644,"end":645},"obj":","},{"id":"tok103","span":{"begin":646,"end":648},"obj":"PRP"},{"id":"tok104","span":{"begin":649,"end":657},"obj":"VBD"},{"id":"tok105","span":{"begin":658,"end":659},"obj":"DT"},{"id":"tok106","span":{"begin":660,"end":666},"obj":"JJR"},{"id":"tok107","span":{"begin":667,"end":676},"obj":"NN"},{"id":"tok108","span":{"begin":677,"end":679},"obj":"IN"},{"id":"tok109","span":{"begin":680,"end":683},"obj":"NNP"},{"id":"tok110","span":{"begin":684,"end":687},"obj":"NNP"},{"id":"tok111","span":{"begin":688,"end":690},"obj":"IN"},{"id":"tok112","span":{"begin":691,"end":694},"obj":"DT"},{"id":"tok113","span":{"begin":695,"end":704},"obj":"NN"},{"id":"tok114","span":{"begin":705,"end":713},"obj":"VBN"},{"id":"tok115","span":{"begin":714,"end":716},"obj":"TO"},{"id":"tok116","span":{"begin":717,"end":735},"obj":"JJ"},{"id":"tok117","span":{"begin":736,"end":741},"obj":"NNS"},{"id":"tok118","span":{"begin":741,"end":742},"obj":"."},{"id":"tok119","span":{"begin":743,"end":754},"obj":"RB"},{"id":"tok120","span":{"begin":754,"end":755},"obj":","},{"id":"tok121","span":{"begin":756,"end":758},"obj":"PRP"},{"id":"tok122","span":{"begin":759,"end":764},"obj":"VBD"},{"id":"tok123","span":{"begin":765,"end":769},"obj":"IN"},{"id":"tok124","span":{"begin":770,"end":782},"obj":"VBG"},{"id":"tok125","span":{"begin":783,"end":786},"obj":"NNP"},{"id":"tok126","span":{"begin":787,"end":790},"obj":"NNP"},{"id":"tok127","span":{"begin":791,"end":795},"obj":"IN"},{"id":"tok128","span":{"begin":796,"end":801},"obj":"NNS"},{"id":"tok129","span":{"begin":802,"end":812},"obj":"NNS"},{"id":"tok130","span":{"begin":813,"end":814},"obj":"DT"},{"id":"tok131","span":{"begin":815,"end":821},"obj":"JJ"},{"id":"tok132","span":{"begin":821,"end":822},"obj":","},{"id":"tok133","span":{"begin":823,"end":826},"obj":"NNP"},{"id":"tok134","span":{"begin":826,"end":827},"obj":"CD"},{"id":"tok135","span":{"begin":827,"end":828},"obj":":"},{"id":"tok136","span":{"begin":828,"end":837},"obj":"JJ"},{"id":"tok137","span":{"begin":838,"end":848},"obj":"NN"},{"id":"tok138","span":{"begin":849,"end":857},"obj":"NN"},{"id":"tok139","span":{"begin":857,"end":858},"obj":","},{"id":"tok140","span":{"begin":859,"end":862},"obj":"CC"},{"id":"tok141","span":{"begin":863,"end":867},"obj":"DT"},{"id":"tok142","span":{"begin":868,"end":875},"obj":"NN"},{"id":"tok143","span":{"begin":876,"end":878},"obj":"IN"},{"id":"tok144","span":{"begin":879,"end":882},"obj":"DT"},{"id":"tok145","span":{"begin":883,"end":894},"obj":"NN"},{"id":"tok146","span":{"begin":895,"end":904},"obj":"NN"},{"id":"tok147","span":{"begin":905,"end":907},"obj":"IN"},{"id":"tok148","span":{"begin":908,"end":911},"obj":"DT"},{"id":"tok149","span":{"begin":912,"end":915},"obj":"NNP"},{"id":"tok150","span":{"begin":916,"end":923},"obj":"NNS"},{"id":"tok151","span":{"begin":924,"end":927},"obj":"DT"},{"id":"tok152","span":{"begin":928,"end":936},"obj":"NN"},{"id":"tok153","span":{"begin":936,"end":937},"obj":"."},{"id":"tok154","span":{"begin":938,"end":945},"obj":"RB"},{"id":"tok155","span":{"begin":945,"end":946},"obj":","},{"id":"tok156","span":{"begin":947,"end":950},"obj":"DT"},{"id":"tok157","span":{"begin":951,"end":958},"obj":"NNS"},{"id":"tok158","span":{"begin":959,"end":961},"obj":"IN"},{"id":"tok159","span":{"begin":962,"end":966},"obj":"DT"},{"id":"tok160","span":{"begin":967,"end":972},"obj":"NN"},{"id":"tok161","span":{"begin":973,"end":979},"obj":"VBP"},{"id":"tok162","span":{"begin":980,"end":983},"obj":"DT"},{"id":"tok163","span":{"begin":984,"end":987},"obj":"NNP"},{"id":"tok164","span":{"begin":988,"end":991},"obj":"NNP"},{"id":"tok165","span":{"begin":992,"end":994},"obj":"TO"},{"id":"tok166","span":{"begin":995,"end":997},"obj":"VB"},{"id":"tok167","span":{"begin":998,"end":999},"obj":"DT"},{"id":"tok168","span":{"begin":1000,"end":1003},"obj":"NNP"},{"id":"tok169","span":{"begin":1004,"end":1007},"obj":"NNP"},{"id":"tok170","span":{"begin":1007,"end":1008},"obj":"CD"},{"id":"tok171","span":{"begin":1008,"end":1009},"obj":":"},{"id":"tok172","span":{"begin":1009,"end":1018},"obj":"JJ"},{"id":"tok173","span":{"begin":1019,"end":1038},"obj":"JJ"},{"id":"tok174","span":{"begin":1039,"end":1048},"obj":"JJ"},{"id":"tok175","span":{"begin":1049,"end":1056},"obj":"NN"},{"id":"tok176","span":{"begin":1057,"end":1058},"obj":"("},{"id":"tok177","span":{"begin":1058,"end":1062},"obj":"NNP"},{"id":"tok178","span":{"begin":1062,"end":1063},"obj":")"},{"id":"tok179","span":{"begin":1063,"end":1064},"obj":"."},{"id":"tok180","span":{"begin":1065,"end":1067},"obj":"PRP"},{"id":"tok181","span":{"begin":1068,"end":1072},"obj":"RB"},{"id":"tok182","span":{"begin":1073,"end":1082},"obj":"VB"},{"id":"tok183","span":{"begin":1083,"end":1084},"obj":"DT"},{"id":"tok184","span":{"begin":1085,"end":1094},"obj":"NN"},{"id":"tok185","span":{"begin":1095,"end":1098},"obj":"IN"},{"id":"tok186","span":{"begin":1099,"end":1104},"obj":"NNP"},{"id":"tok187","span":{"begin":1105,"end":1113},"obj":"NN"},{"id":"tok188","span":{"begin":1114,"end":1116},"obj":"TO"},{"id":"tok189","span":{"begin":1117,"end":1123},"obj":"NN"},{"id":"tok190","span":{"begin":1124,"end":1127},"obj":"CC"},{"id":"tok191","span":{"begin":1128,"end":1133},"obj":"VB"},{"id":"tok192","span":{"begin":1134,"end":1137},"obj":"DT"},{"id":"tok193","span":{"begin":1138,"end":1150},"obj":"NN"},{"id":"tok194","span":{"begin":1151,"end":1153},"obj":"IN"},{"id":"tok195","span":{"begin":1154,"end":1158},"obj":"DT"},{"id":"tok196","span":{"begin":1159,"end":1163},"obj":"NNP"},{"id":"tok197","span":{"begin":1163,"end":1164},"obj":"."},{"id":"tok198","span":{"begin":1165,"end":1170},"obj":"IN"},{"id":"tok199","span":{"begin":1171,"end":1176},"obj":"NNP"},{"id":"tok200","span":{"begin":1177,"end":1185},"obj":"NN"},{"id":"tok201","span":{"begin":1186,"end":1188},"obj":"VBZ"},{"id":"tok202","span":{"begin":1189,"end":1195},"obj":"RB"},{"id":"tok203","span":{"begin":1196,"end":1205},"obj":"JJ"},{"id":"tok204","span":{"begin":1206,"end":1208},"obj":"IN"},{"id":"tok205","span":{"begin":1209,"end":1212},"obj":"DT"},{"id":"tok206","span":{"begin":1213,"end":1217},"obj":"NNP"},{"id":"tok207","span":{"begin":1217,"end":1218},"obj":","},{"id":"tok208","span":{"begin":1219,"end":1229},"obj":"VBG"},{"id":"tok209","span":{"begin":1230,"end":1234},"obj":"DT"},{"id":"tok210","span":{"begin":1235,"end":1243},"obj":"NN"},{"id":"tok211","span":{"begin":1244,"end":1247},"obj":"MD"},{"id":"tok212","span":{"begin":1248,"end":1253},"obj":"VB"},{"id":"tok213","span":{"begin":1254,"end":1256},"obj":"IN"},{"id":"tok214","span":{"begin":1257,"end":1259},"obj":"DT"},{"id":"tok215","span":{"begin":1260,"end":1268},"obj":"NN"},{"id":"tok216","span":{"begin":1269,"end":1272},"obj":"IN"},{"id":"tok217","span":{"begin":1273,"end":1284},"obj":"JJ"},{"id":"tok218","span":{"begin":1285,"end":1298},"obj":"NNS"},{"id":"tok219","span":{"begin":1299,"end":1306},"obj":"IN"},{"id":"tok220","span":{"begin":1307,"end":1315},"obj":"VBG"},{"id":"tok221","span":{"begin":1316,"end":1319},"obj":"CC"},{"id":"tok222","span":{"begin":1320,"end":1328},"obj":"VBG"},{"id":"tok223","span":{"begin":1329,"end":1332},"obj":"NNP"},{"id":"tok224","span":{"begin":1333,"end":1343},"obj":"NNS"},{"id":"tok225","span":{"begin":1343,"end":1344},"obj":"."},{"id":"lookup1","span":{"begin":65,"end":67},"obj":"country_code"},{"id":"lookup2","span":{"begin":273,"end":275},"obj":"country_code"},{"id":"lookup3","span":{"begin":426,"end":430},"obj":"stop"},{"id":"lookup4","span":{"begin":534,"end":536},"obj":"country_code"},{"id":"lookup5","span":{"begin":608,"end":610},"obj":"country_code"},{"id":"lookup6","span":{"begin":688,"end":690},"obj":"country_code"},{"id":"lookup7","span":{"begin":714,"end":716},"obj":"country_code"},{"id":"lookup8","span":{"begin":962,"end":966},"obj":"time_modifier"},{"id":"lookup9","span":{"begin":992,"end":994},"obj":"country_code"},{"id":"lookup10","span":{"begin":995,"end":997},"obj":"country_code"},{"id":"lookup11","span":{"begin":1065,"end":1067},"obj":"stop"},{"id":"lookup12","span":{"begin":1114,"end":1116},"obj":"country_code"},{"id":"lookup13","span":{"begin":1154,"end":1158},"obj":"time_modifier"},{"id":"lookup14","span":{"begin":1186,"end":1188},"obj":"country_code"},{"id":"lookup15","span":{"begin":1206,"end":1208},"obj":"country_code"},{"id":"lookup16","span":{"begin":1230,"end":1234},"obj":"time_modifier"},{"id":"lookup17","span":{"begin":1254,"end":1256},"obj":"country_code"},{"id":"lookup18","span":{"begin":1257,"end":1259},"obj":"country_code"},{"id":"event1","span":{"begin":649,"end":657},"obj":"OCCURRENCE"},{"id":"event2","span":{"begin":1073,"end":1082},"obj":"OCCURRENCE"},{"id":"event3","span":{"begin":293,"end":303},"obj":"OCCURRENCE"},{"id":"event4","span":{"begin":216,"end":223},"obj":"OCCURRENCE"},{"id":"event5","span":{"begin":68,"end":73},"obj":"OCCURRENCE"},{"id":"event6","span":{"begin":1248,"end":1253},"obj":"OCCURRENCE"},{"id":"event7","span":{"begin":1128,"end":1133},"obj":"OCCURRENCE"},{"id":"event8","span":{"begin":1320,"end":1328},"obj":"OCCURRENCE"},{"id":"event9","span":{"begin":419,"end":424},"obj":"OCCURRENCE"},{"id":"event10","span":{"begin":597,"end":607},"obj":"OCCURRENCE"},{"id":"event11","span":{"begin":1219,"end":1229},"obj":"OCCURRENCE"},{"id":"event12","span":{"begin":435,"end":441},"obj":"REPORTING"},{"id":"event13","span":{"begin":160,"end":166},"obj":"OCCURRENCE"},{"id":"event14","span":{"begin":759,"end":764},"obj":"OCCURRENCE"}],"attributes":[{"subj":"tok1","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok2","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok3","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok4","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok5","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok6","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok7","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok8","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok9","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok10","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok11","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok12","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok13","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok14","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok15","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok16","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok17","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok18","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok19","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok20","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok21","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok22","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok23","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok24","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok25","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok26","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok27","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok28","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok29","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok30","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok31","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok32","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok33","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok34","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok35","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok36","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok37","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok38","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok39","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok40","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok41","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok42","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok43","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok44","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok45","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok46","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok47","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok48","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok49","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok50","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok51","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok52","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok53","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok54","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok55","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok56","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok57","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok58","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok59","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok60","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok61","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok62","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok63","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok64","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok65","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok66","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok67","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok68","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok69","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok70","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok71","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok72","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok73","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok74","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok75","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok76","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok77","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok78","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok79","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok80","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok81","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok82","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok83","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok84","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok85","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok86","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok87","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok88","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok89","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok90","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok91","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok92","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok93","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok94","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok95","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok96","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok97","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok98","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok99","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok100","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok101","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok102","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok103","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok104","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok105","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok106","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok107","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok108","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok109","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok110","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok111","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok112","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok113","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok114","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok115","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok116","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok117","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok118","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok119","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok120","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok121","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok122","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok123","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok124","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok125","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok126","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok127","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok128","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok129","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok130","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok131","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok132","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok133","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok134","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok135","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok136","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok137","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok138","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok139","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok140","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok141","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok142","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok143","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok144","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok145","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok146","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok147","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok148","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok149","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok150","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok151","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok152","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok153","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok154","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok155","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok156","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok157","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok158","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok159","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok160","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok161","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok162","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok163","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok164","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok165","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok166","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok167","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok168","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok169","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok170","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok171","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok172","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok173","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok174","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok175","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok176","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok177","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok178","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok179","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok180","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok181","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok182","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok183","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok184","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok185","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok186","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok187","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok188","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok189","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok190","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok191","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok192","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok193","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok194","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok195","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok196","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok197","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok198","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok199","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok200","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok201","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok202","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok203","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok204","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok205","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok206","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok207","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok208","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok209","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok210","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok211","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok212","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok213","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok214","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok215","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok216","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok217","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok218","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok219","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok220","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok221","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok222","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok223","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok224","pred":"source","obj":"LitCovid-TimeML"},{"subj":"tok225","pred":"source","obj":"LitCovid-TimeML"},{"subj":"lookup1","pred":"source","obj":"LitCovid-TimeML"},{"subj":"lookup2","pred":"source","obj":"LitCovid-TimeML"},{"subj":"lookup3","pred":"source","obj":"LitCovid-TimeML"},{"subj":"lookup4","pred":"source","obj":"LitCovid-TimeML"},{"subj":"lookup5","pred":"source","obj":"LitCovid-TimeML"},{"subj":"lookup6","pred":"source","obj":"LitCovid-TimeML"},{"subj":"lookup7","pred":"source","obj":"LitCovid-TimeML"},{"subj":"lookup8","pred":"source","obj":"LitCovid-TimeML"},{"subj":"lookup9","pred":"source","obj":"LitCovid-TimeML"},{"subj":"lookup10","pred":"source","obj":"LitCovid-TimeML"},{"subj":"lookup11","pred":"source","obj":"LitCovid-TimeML"},{"subj":"lookup12","pred":"source","obj":"LitCovid-TimeML"},{"subj":"lookup13","pred":"source","obj":"LitCovid-TimeML"},{"subj":"lookup14","pred":"source","obj":"LitCovid-TimeML"},{"subj":"lookup15","pred":"source","obj":"LitCovid-TimeML"},{"subj":"lookup16","pred":"source","obj":"LitCovid-TimeML"},{"subj":"lookup17","pred":"source","obj":"LitCovid-TimeML"},{"subj":"lookup18","pred":"source","obj":"LitCovid-TimeML"},{"subj":"event1","pred":"source","obj":"LitCovid-TimeML"},{"subj":"event2","pred":"source","obj":"LitCovid-TimeML"},{"subj":"event3","pred":"source","obj":"LitCovid-TimeML"},{"subj":"event4","pred":"source","obj":"LitCovid-TimeML"},{"subj":"event5","pred":"source","obj":"LitCovid-TimeML"},{"subj":"event6","pred":"source","obj":"LitCovid-TimeML"},{"subj":"event7","pred":"source","obj":"LitCovid-TimeML"},{"subj":"event8","pred":"source","obj":"LitCovid-TimeML"},{"subj":"event9","pred":"source","obj":"LitCovid-TimeML"},{"subj":"event10","pred":"source","obj":"LitCovid-TimeML"},{"subj":"event11","pred":"source","obj":"LitCovid-TimeML"},{"subj":"event12","pred":"source","obj":"LitCovid-TimeML"},{"subj":"event13","pred":"source","obj":"LitCovid-TimeML"},{"subj":"event14","pred":"source","obj":"LitCovid-TimeML"}]},{"project":"LitCovid-PD-MONDO-v1","denotations":[{"id":"T1","span":{"begin":1333,"end":1343},"obj":"Disease"}],"attributes":[{"id":"A1","pred":"mondo_id","subj":"T1","obj":"http://purl.obolibrary.org/obo/MONDO_0005550"},{"subj":"T1","pred":"source","obj":"LitCovid-PD-MONDO-v1"}]},{"project":"LitCovid-PD-FMA-UBERON-v1","denotations":[{"id":"T34653","span":{"begin":895,"end":904},"obj":"Body_part"},{"id":"T1","span":{"begin":139,"end":142},"obj":"Body_part"},{"id":"T2","span":{"begin":507,"end":510},"obj":"Body_part"},{"id":"T3","span":{"begin":523,"end":526},"obj":"Body_part"},{"id":"T4","span":{"begin":568,"end":571},"obj":"Body_part"},{"id":"T5","span":{"begin":684,"end":687},"obj":"Body_part"},{"id":"T6","span":{"begin":695,"end":704},"obj":"Body_part"},{"id":"T7","span":{"begin":736,"end":741},"obj":"Body_part"},{"id":"T8","span":{"begin":787,"end":790},"obj":"Body_part"},{"id":"T9","span":{"begin":796,"end":801},"obj":"Body_part"},{"id":"T10","span":{"begin":912,"end":915},"obj":"Body_part"},{"id":"T11","span":{"begin":988,"end":991},"obj":"Body_part"}],"attributes":[{"id":"A3","pred":"fma_id","subj":"T3","obj":"http://purl.org/sig/ont/fma/fma67095"},{"id":"A9","pred":"fma_id","subj":"T9","obj":"http://purl.org/sig/ont/fma/fma68646"},{"id":"A11","pred":"fma_id","subj":"T11","obj":"http://purl.org/sig/ont/fma/fma67095"},{"id":"A1","pred":"fma_id","subj":"T1","obj":"http://purl.org/sig/ont/fma/fma67095"},{"id":"A8","pred":"fma_id","subj":"T8","obj":"http://purl.org/sig/ont/fma/fma67095"},{"id":"A5","pred":"fma_id","subj":"T5","obj":"http://purl.org/sig/ont/fma/fma67095"},{"id":"A6","pred":"fma_id","subj":"T6","obj":"http://purl.org/sig/ont/fma/fma66835"},{"id":"A2","pred":"fma_id","subj":"T2","obj":"http://purl.org/sig/ont/fma/fma67095"},{"id":"A10","pred":"fma_id","subj":"T10","obj":"http://purl.org/sig/ont/fma/fma67095"},{"id":"A63207","pred":"uberon_id","subj":"T34653","obj":"http://purl.obolibrary.org/obo/UBERON_2000106"},{"id":"A4","pred":"fma_id","subj":"T4","obj":"http://purl.org/sig/ont/fma/fma67095"},{"id":"A7","pred":"fma_id","subj":"T7","obj":"http://purl.org/sig/ont/fma/fma68646"},{"subj":"T34653","pred":"source","obj":"LitCovid-PD-FMA-UBERON-v1"},{"subj":"T1","pred":"source","obj":"LitCovid-PD-FMA-UBERON-v1"},{"subj":"T2","pred":"source","obj":"LitCovid-PD-FMA-UBERON-v1"},{"subj":"T3","pred":"source","obj":"LitCovid-PD-FMA-UBERON-v1"},{"subj":"T4","pred":"source","obj":"LitCovid-PD-FMA-UBERON-v1"},{"subj":"T5","pred":"source","obj":"LitCovid-PD-FMA-UBERON-v1"},{"subj":"T6","pred":"source","obj":"LitCovid-PD-FMA-UBERON-v1"},{"subj":"T7","pred":"source","obj":"LitCovid-PD-FMA-UBERON-v1"},{"subj":"T8","pred":"source","obj":"LitCovid-PD-FMA-UBERON-v1"},{"subj":"T9","pred":"source","obj":"LitCovid-PD-FMA-UBERON-v1"},{"subj":"T10","pred":"source","obj":"LitCovid-PD-FMA-UBERON-v1"},{"subj":"T11","pred":"source","obj":"LitCovid-PD-FMA-UBERON-v1"}]}],"config":{"attribute types":[{"pred":"source","value type":"selection","values":[{"id":"LitCovid-OGER","color":"#eccb93","default":true},{"id":"LitCovid-OGER-BB","color":"#b193ec"},{"id":"LitCovid-sentences-v1","color":"#93ec97"},{"id":"LitCovid-TimeML","color":"#ec93a8"},{"id":"LitCovid-PD-MONDO-v1","color":"#93c2ec"},{"id":"LitCovid-PD-FMA-UBERON-v1","color":"#dcec93"}]}]}}