Id |
Subject |
Object |
Predicate |
Lexical cue |
T359 |
3-5 |
IN |
denotes |
of |
T360 |
6-15 |
JJ |
denotes |
different |
T361 |
16-26 |
NNS |
denotes |
morphogens |
T362 |
27-30 |
CC |
denotes |
and |
T363 |
31-41 |
JJ |
denotes |
downstream |
T364 |
58-64 |
NNS |
denotes |
events |
T365 |
42-57 |
JJ |
denotes |
transcriptional |
T366 |
65-69 |
WDT |
denotes |
that |
T367 |
70-75 |
VBP |
denotes |
guide |
T368 |
76-80 |
NN |
denotes |
hair |
T369 |
81-84 |
NN |
denotes |
bud |
T370 |
85-94 |
NN |
denotes |
formation |
T371 |
95-101 |
IN |
denotes |
within |
T372 |
102-105 |
DT |
denotes |
the |
T373 |
117-121 |
NN |
denotes |
skin |
T374 |
106-116 |
VBG |
denotes |
developing |
T375 |
121-122 |
. |
denotes |
. |
T1019 |
318-327 |
JJ |
denotes |
Mammalian |
T1020 |
328-339 |
NN |
denotes |
development |
T1021 |
340-348 |
VBZ |
denotes |
involves |
T1022 |
349-352 |
DT |
denotes |
the |
T1023 |
353-366 |
NN |
denotes |
morphogenesis |
T1024 |
367-369 |
IN |
denotes |
of |
T1025 |
370-377 |
JJ |
denotes |
complex |
T1026 |
396-406 |
NNS |
denotes |
structures |
T1027 |
378-383 |
CD |
denotes |
three |
T1028 |
384-395 |
JJ |
denotes |
dimensional |
T1029 |
383-384 |
HYPH |
denotes |
- |
T1030 |
407-411 |
IN |
denotes |
from |
T1031 |
412-421 |
RB |
denotes |
seemingly |
T1032 |
422-429 |
JJ |
denotes |
uniform |
T1033 |
430-436 |
NNS |
denotes |
sheets |
T1034 |
437-439 |
CC |
denotes |
or |
T1035 |
440-446 |
NNS |
denotes |
masses |
T1036 |
447-449 |
IN |
denotes |
of |
T1037 |
450-455 |
NNS |
denotes |
cells |
T1038 |
455-456 |
. |
denotes |
. |
T1039 |
456-594 |
sentence |
denotes |
A simple bud-like structure initiates the formation of many organs, including lungs, spinal cord, mammary glands, and hair follicles [1]. |
T1040 |
457-458 |
DT |
denotes |
A |
T1041 |
475-484 |
NN |
denotes |
structure |
T1042 |
459-465 |
JJ |
denotes |
simple |
T1043 |
466-469 |
NN |
denotes |
bud |
T1044 |
470-474 |
JJ |
denotes |
like |
T1045 |
469-470 |
HYPH |
denotes |
- |
T1046 |
485-494 |
VBZ |
denotes |
initiates |
T1047 |
495-498 |
DT |
denotes |
the |
T1048 |
499-508 |
NN |
denotes |
formation |
T1049 |
509-511 |
IN |
denotes |
of |
T1050 |
512-516 |
JJ |
denotes |
many |
T1051 |
517-523 |
NNS |
denotes |
organs |
T1052 |
523-525 |
, |
denotes |
, |
T1053 |
525-534 |
VBG |
denotes |
including |
T1054 |
535-540 |
NNS |
denotes |
lungs |
T1055 |
540-542 |
, |
denotes |
, |
T1056 |
542-548 |
JJ |
denotes |
spinal |
T1057 |
549-553 |
NN |
denotes |
cord |
T1058 |
553-555 |
, |
denotes |
, |
T1059 |
555-562 |
JJ |
denotes |
mammary |
T1060 |
563-569 |
NNS |
denotes |
glands |
T1061 |
569-571 |
, |
denotes |
, |
T1062 |
571-574 |
CC |
denotes |
and |
T1063 |
575-579 |
NN |
denotes |
hair |
T1064 |
580-589 |
NNS |
denotes |
follicles |
T1065 |
590-591 |
-LRB- |
denotes |
[ |
T1066 |
591-592 |
CD |
denotes |
1 |
T1067 |
592-593 |
-RRB- |
denotes |
] |
T1068 |
593-594 |
. |
denotes |
. |
T1069 |
594-772 |
sentence |
denotes |
The multipotent, adhering epithelial cells are typically attached to an underlying basal lamina that polarizes the epithelial sheet and separates it from surrounding mesenchyme. |
T1070 |
595-598 |
DT |
denotes |
The |
T1071 |
632-637 |
NNS |
denotes |
cells |
T1072 |
599-610 |
JJ |
denotes |
multipotent |
T1073 |
610-612 |
, |
denotes |
, |
T1074 |
612-620 |
VBG |
denotes |
adhering |
T1075 |
621-631 |
JJ |
denotes |
epithelial |
T1076 |
652-660 |
VBN |
denotes |
attached |
T1077 |
638-641 |
VBP |
denotes |
are |
T1078 |
642-651 |
RB |
denotes |
typically |
T1079 |
661-663 |
IN |
denotes |
to |
T1080 |
664-666 |
DT |
denotes |
an |
T1081 |
684-690 |
NN |
denotes |
lamina |
T1082 |
667-677 |
VBG |
denotes |
underlying |
T1083 |
678-683 |
JJ |
denotes |
basal |
T1084 |
691-695 |
WDT |
denotes |
that |
T1085 |
696-705 |
VBZ |
denotes |
polarizes |
T1086 |
706-709 |
DT |
denotes |
the |
T1087 |
721-726 |
NN |
denotes |
sheet |
T1088 |
710-720 |
JJ |
denotes |
epithelial |
T1089 |
727-730 |
CC |
denotes |
and |
T1090 |
731-740 |
VBZ |
denotes |
separates |
T1091 |
741-743 |
PRP |
denotes |
it |
T1092 |
744-748 |
IN |
denotes |
from |
T1093 |
749-760 |
VBG |
denotes |
surrounding |
T1094 |
761-771 |
NN |
denotes |
mesenchyme |
T1095 |
771-772 |
. |
denotes |
. |
T1096 |
772-952 |
sentence |
denotes |
Budding morphogenesis is guided by a reciprocal exchange of signals between epithelium and mesenchyme to specify the identity of the organ that will form and to govern its growth. |
T1097 |
773-780 |
VBG |
denotes |
Budding |
T1098 |
781-794 |
NN |
denotes |
morphogenesis |
T1099 |
798-804 |
VBN |
denotes |
guided |
T1100 |
795-797 |
VBZ |
denotes |
is |
T1101 |
805-807 |
IN |
denotes |
by |
T1102 |
808-809 |
DT |
denotes |
a |
T1103 |
821-829 |
NN |
denotes |
exchange |
T1104 |
810-820 |
JJ |
denotes |
reciprocal |
T1105 |
830-832 |
IN |
denotes |
of |
T1106 |
833-840 |
NNS |
denotes |
signals |
T1107 |
841-848 |
IN |
denotes |
between |
T1108 |
849-859 |
NN |
denotes |
epithelium |
T1109 |
860-863 |
CC |
denotes |
and |
T1110 |
864-874 |
NN |
denotes |
mesenchyme |
T1111 |
875-877 |
TO |
denotes |
to |
T1112 |
878-885 |
VB |
denotes |
specify |
T1113 |
886-889 |
DT |
denotes |
the |
T1114 |
890-898 |
NN |
denotes |
identity |
T1115 |
899-901 |
IN |
denotes |
of |
T1116 |
902-905 |
DT |
denotes |
the |
T1117 |
906-911 |
NN |
denotes |
organ |
T1118 |
912-916 |
WDT |
denotes |
that |
T1119 |
922-926 |
VB |
denotes |
form |
T1120 |
917-921 |
MD |
denotes |
will |
T1121 |
927-930 |
CC |
denotes |
and |
T1122 |
931-933 |
TO |
denotes |
to |
T1123 |
934-940 |
VB |
denotes |
govern |
T1124 |
941-944 |
PRP$ |
denotes |
its |
T1125 |
945-951 |
NN |
denotes |
growth |
T1126 |
951-952 |
. |
denotes |
. |
T1127 |
952-1128 |
sentence |
denotes |
At the helm of these molecular communication pathways are Wnts, bone morphogenic proteins (BMPs), transforming growth factor βs (TGF-βs), and fibroblast growth factors (FGFs). |
T1128 |
953-955 |
IN |
denotes |
At |
T1129 |
1007-1010 |
VBP |
denotes |
are |
T1130 |
956-959 |
DT |
denotes |
the |
T1131 |
960-964 |
NN |
denotes |
helm |
T1132 |
965-967 |
IN |
denotes |
of |
T1133 |
968-973 |
DT |
denotes |
these |
T1134 |
998-1006 |
NNS |
denotes |
pathways |
T1135 |
974-983 |
JJ |
denotes |
molecular |
T1136 |
984-997 |
NN |
denotes |
communication |
T1137 |
1011-1015 |
NNS |
denotes |
Wnts |
T1138 |
1015-1017 |
, |
denotes |
, |
T1139 |
1017-1021 |
NN |
denotes |
bone |
T1140 |
1034-1042 |
NN |
denotes |
proteins |
T1141 |
1022-1033 |
JJ |
denotes |
morphogenic |
T1142 |
1043-1044 |
-LRB- |
denotes |
( |
T1143 |
1044-1048 |
NNS |
denotes |
BMPs |
T1144 |
1048-1049 |
-RRB- |
denotes |
) |
T1145 |
1049-1051 |
, |
denotes |
, |
T1146 |
1051-1063 |
VBG |
denotes |
transforming |
T1147 |
1078-1080 |
NNS |
denotes |
βs |
T1148 |
1064-1070 |
NN |
denotes |
growth |
T1149 |
1071-1077 |
NN |
denotes |
factor |
T1150 |
1081-1082 |
-LRB- |
denotes |
( |
T1151 |
1082-1085 |
NN |
denotes |
TGF |
T1152 |
1086-1088 |
NNS |
denotes |
βs |
T1153 |
1085-1086 |
HYPH |
denotes |
- |
T1154 |
1088-1089 |
-RRB- |
denotes |
) |
T1155 |
1089-1091 |
, |
denotes |
, |
T1156 |
1091-1094 |
CC |
denotes |
and |
T1157 |
1095-1105 |
NN |
denotes |
fibroblast |
T1158 |
1113-1120 |
NNS |
denotes |
factors |
T1159 |
1106-1112 |
NN |
denotes |
growth |
T1160 |
1121-1122 |
-LRB- |
denotes |
( |
T1161 |
1122-1126 |
NNS |
denotes |
FGFs |
T1162 |
1126-1127 |
-RRB- |
denotes |
) |
T1163 |
1127-1128 |
. |
denotes |
. |
T1164 |
1128-1352 |
sentence |
denotes |
Through activation of cell surface transmembrane receptors, these external signaling molecules trigger distinct cascades of intracellular events that culminate in changes in gene expression, growth, and differentiation [2]. |
T1165 |
1129-1136 |
IN |
denotes |
Through |
T1166 |
1224-1231 |
VBP |
denotes |
trigger |
T1167 |
1137-1147 |
NN |
denotes |
activation |
T1168 |
1148-1150 |
IN |
denotes |
of |
T1169 |
1151-1155 |
NN |
denotes |
cell |
T1170 |
1156-1163 |
NN |
denotes |
surface |
T1171 |
1178-1187 |
NNS |
denotes |
receptors |
T1172 |
1164-1177 |
NN |
denotes |
transmembrane |
T1173 |
1187-1189 |
, |
denotes |
, |
T1174 |
1189-1194 |
DT |
denotes |
these |
T1175 |
1214-1223 |
NNS |
denotes |
molecules |
T1176 |
1195-1203 |
JJ |
denotes |
external |
T1177 |
1204-1213 |
NN |
denotes |
signaling |
T1178 |
1232-1240 |
JJ |
denotes |
distinct |
T1179 |
1241-1249 |
NNS |
denotes |
cascades |
T1180 |
1250-1252 |
IN |
denotes |
of |
T1181 |
1253-1266 |
JJ |
denotes |
intracellular |
T1182 |
1267-1273 |
NNS |
denotes |
events |
T1183 |
1274-1278 |
WDT |
denotes |
that |
T1184 |
1279-1288 |
VBP |
denotes |
culminate |
T1185 |
1289-1291 |
IN |
denotes |
in |
T1186 |
1292-1299 |
NNS |
denotes |
changes |
T1187 |
1300-1302 |
IN |
denotes |
in |
T1188 |
1303-1307 |
NN |
denotes |
gene |
T1189 |
1308-1318 |
NN |
denotes |
expression |
T1190 |
1318-1320 |
, |
denotes |
, |
T1191 |
1320-1326 |
NN |
denotes |
growth |
T1192 |
1326-1328 |
, |
denotes |
, |
T1193 |
1328-1331 |
CC |
denotes |
and |
T1194 |
1332-1347 |
NN |
denotes |
differentiation |
T1195 |
1348-1349 |
-LRB- |
denotes |
[ |
T1196 |
1349-1350 |
CD |
denotes |
2 |
T1197 |
1350-1351 |
-RRB- |
denotes |
] |
T1198 |
1351-1352 |
. |
denotes |
. |
T1199 |
1352-1525 |
sentence |
denotes |
How this constellation of signals collaborates in tailoring each budding process so that it executes a distinct morphogenetic program has yet to be comprehensively defined. |
T1200 |
1353-1356 |
WRB |
denotes |
How |
T1201 |
1387-1399 |
VBZ |
denotes |
collaborates |
T1202 |
1357-1361 |
DT |
denotes |
this |
T1203 |
1362-1375 |
NN |
denotes |
constellation |
T1204 |
1376-1378 |
IN |
denotes |
of |
T1205 |
1379-1386 |
NNS |
denotes |
signals |
T1206 |
1487-1490 |
VBZ |
denotes |
has |
T1207 |
1400-1402 |
IN |
denotes |
in |
T1208 |
1403-1412 |
VBG |
denotes |
tailoring |
T1209 |
1413-1417 |
DT |
denotes |
each |
T1210 |
1426-1433 |
NN |
denotes |
process |
T1211 |
1418-1425 |
NN |
denotes |
budding |
T1212 |
1434-1436 |
IN |
denotes |
so |
T1213 |
1445-1453 |
VBZ |
denotes |
executes |
T1214 |
1437-1441 |
IN |
denotes |
that |
T1215 |
1442-1444 |
PRP |
denotes |
it |
T1216 |
1454-1455 |
DT |
denotes |
a |
T1217 |
1479-1486 |
NN |
denotes |
program |
T1218 |
1456-1464 |
JJ |
denotes |
distinct |
T1219 |
1465-1478 |
JJ |
denotes |
morphogenetic |
T1220 |
1491-1494 |
RB |
denotes |
yet |
T1221 |
1517-1524 |
VBN |
denotes |
defined |
T1222 |
1495-1497 |
TO |
denotes |
to |
T1223 |
1498-1500 |
VB |
denotes |
be |
T1224 |
1501-1516 |
RB |
denotes |
comprehensively |
T1225 |
1524-1525 |
. |
denotes |
. |
T1226 |
1525-1751 |
sentence |
denotes |
However, the process appears to be patterned at the initial stages of bud formation, since the relative importance of these pathways and their downstream effectors differ as buds begin to develop and cell fates are specified. |
T1227 |
1526-1533 |
RB |
denotes |
However |
T1228 |
1547-1554 |
VBZ |
denotes |
appears |
T1229 |
1533-1535 |
, |
denotes |
, |
T1230 |
1535-1538 |
DT |
denotes |
the |
T1231 |
1539-1546 |
NN |
denotes |
process |
T1232 |
1555-1557 |
TO |
denotes |
to |
T1233 |
1561-1570 |
VBN |
denotes |
patterned |
T1234 |
1558-1560 |
VB |
denotes |
be |
T1235 |
1571-1573 |
IN |
denotes |
at |
T1236 |
1574-1577 |
DT |
denotes |
the |
T1237 |
1586-1592 |
NNS |
denotes |
stages |
T1238 |
1578-1585 |
JJ |
denotes |
initial |
T1239 |
1593-1595 |
IN |
denotes |
of |
T1240 |
1596-1599 |
NN |
denotes |
bud |
T1241 |
1600-1609 |
NN |
denotes |
formation |
T1242 |
1609-1611 |
, |
denotes |
, |
T1243 |
1611-1616 |
IN |
denotes |
since |
T1244 |
1690-1696 |
VBP |
denotes |
differ |
T1245 |
1617-1620 |
DT |
denotes |
the |
T1246 |
1630-1640 |
NN |
denotes |
importance |
T1247 |
1621-1629 |
JJ |
denotes |
relative |
T1248 |
1641-1643 |
IN |
denotes |
of |
T1249 |
1644-1649 |
DT |
denotes |
these |
T1250 |
1650-1658 |
NNS |
denotes |
pathways |
T1251 |
1659-1662 |
CC |
denotes |
and |
T1252 |
1663-1668 |
PRP$ |
denotes |
their |
T1253 |
1680-1689 |
NNS |
denotes |
effectors |
T1254 |
1669-1679 |
JJ |
denotes |
downstream |
T1255 |
1697-1699 |
IN |
denotes |
as |
T1256 |
1705-1710 |
VBP |
denotes |
begin |
T1257 |
1700-1704 |
NNS |
denotes |
buds |
T1258 |
1711-1713 |
TO |
denotes |
to |
T1259 |
1714-1721 |
VB |
denotes |
develop |
T1260 |
1722-1725 |
CC |
denotes |
and |
T1261 |
1726-1730 |
NN |
denotes |
cell |
T1262 |
1731-1736 |
NNS |
denotes |
fates |
T1263 |
1741-1750 |
VBN |
denotes |
specified |
T1264 |
1737-1740 |
VBP |
denotes |
are |
T1265 |
1750-1751 |
. |
denotes |
. |
T1266 |
1751-1883 |
sentence |
denotes |
The development of a bud requires a number of coordinated changes in the behavior of the targeted cells within an epithelial sheet. |
T1267 |
1752-1755 |
DT |
denotes |
The |
T1268 |
1756-1767 |
NN |
denotes |
development |
T1269 |
1777-1785 |
VBZ |
denotes |
requires |
T1270 |
1768-1770 |
IN |
denotes |
of |
T1271 |
1771-1772 |
DT |
denotes |
a |
T1272 |
1773-1776 |
NN |
denotes |
bud |
T1273 |
1786-1787 |
DT |
denotes |
a |
T1274 |
1788-1794 |
NN |
denotes |
number |
T1275 |
1795-1797 |
IN |
denotes |
of |
T1276 |
1798-1809 |
VBN |
denotes |
coordinated |
T1277 |
1810-1817 |
NNS |
denotes |
changes |
T1278 |
1818-1820 |
IN |
denotes |
in |
T1279 |
1821-1824 |
DT |
denotes |
the |
T1280 |
1825-1833 |
NN |
denotes |
behavior |
T1281 |
1834-1836 |
IN |
denotes |
of |
T1282 |
1837-1840 |
DT |
denotes |
the |
T1283 |
1850-1855 |
NNS |
denotes |
cells |
T1284 |
1841-1849 |
VBN |
denotes |
targeted |
T1285 |
1856-1862 |
IN |
denotes |
within |
T1286 |
1863-1865 |
DT |
denotes |
an |
T1287 |
1877-1882 |
NN |
denotes |
sheet |
T1288 |
1866-1876 |
JJ |
denotes |
epithelial |
T1289 |
1882-1883 |
. |
denotes |
. |
T1290 |
1883-2067 |
sentence |
denotes |
The process must be accompanied by alterations in the proliferation, polarity, shape, and adhesiveness of selected cells, as well as by modifications in their underlying basal lamina. |
T1291 |
1884-1887 |
DT |
denotes |
The |
T1292 |
1888-1895 |
NN |
denotes |
process |
T1293 |
1904-1915 |
VBN |
denotes |
accompanied |
T1294 |
1896-1900 |
MD |
denotes |
must |
T1295 |
1901-1903 |
VB |
denotes |
be |
T1296 |
1916-1918 |
IN |
denotes |
by |
T1297 |
1919-1930 |
NNS |
denotes |
alterations |
T1298 |
1931-1933 |
IN |
denotes |
in |
T1299 |
1934-1937 |
DT |
denotes |
the |
T1300 |
1938-1951 |
NN |
denotes |
proliferation |
T1301 |
1951-1953 |
, |
denotes |
, |
T1302 |
1953-1961 |
NN |
denotes |
polarity |
T1303 |
1961-1963 |
, |
denotes |
, |
T1304 |
1963-1968 |
NN |
denotes |
shape |
T1305 |
1968-1970 |
, |
denotes |
, |
T1306 |
1970-1973 |
CC |
denotes |
and |
T1307 |
1974-1986 |
NN |
denotes |
adhesiveness |
T1308 |
1987-1989 |
IN |
denotes |
of |
T1309 |
1990-1998 |
VBN |
denotes |
selected |
T1310 |
1999-2004 |
NNS |
denotes |
cells |
T1311 |
2004-2006 |
, |
denotes |
, |
T1312 |
2006-2008 |
RB |
denotes |
as |
T1313 |
2014-2016 |
IN |
denotes |
as |
T1314 |
2009-2013 |
RB |
denotes |
well |
T1315 |
2017-2019 |
IN |
denotes |
by |
T1316 |
2020-2033 |
NNS |
denotes |
modifications |
T1317 |
2034-2036 |
IN |
denotes |
in |
T1318 |
2037-2042 |
PRP$ |
denotes |
their |
T1319 |
2060-2066 |
NN |
denotes |
lamina |
T1320 |
2043-2053 |
VBG |
denotes |
underlying |
T1321 |
2054-2059 |
JJ |
denotes |
basal |
T1322 |
2066-2067 |
. |
denotes |
. |
T1324 |
2068-2072 |
RB |
denotes |
Thus |
T1325 |
2141-2153 |
VBN |
denotes |
orchestrated |
T1326 |
2072-2074 |
, |
denotes |
, |
T1327 |
2074-2087 |
JJ |
denotes |
extracellular |
T1328 |
2111-2120 |
NN |
denotes |
crosstalk |
T1329 |
2088-2098 |
JJ |
denotes |
epithelial |
T1330 |
2099-2110 |
JJ |
denotes |
mesenchymal |
T1331 |
2098-2099 |
HYPH |
denotes |
- |
T1332 |
2121-2125 |
MD |
denotes |
must |
T1333 |
2126-2128 |
VB |
denotes |
be |
T1334 |
2129-2140 |
RB |
denotes |
intricately |
T1335 |
2154-2156 |
TO |
denotes |
to |
T1336 |
2157-2163 |
VB |
denotes |
couple |
T1337 |
2164-2167 |
DT |
denotes |
the |
T1338 |
2168-2181 |
NN |
denotes |
determination |
T1339 |
2182-2184 |
IN |
denotes |
of |
T1340 |
2185-2193 |
JJ |
denotes |
distinct |
T1342 |
2194-2198 |
NN |
denotes |
cell |
R202 |
T364 |
T361 |
conj |
events,morphogens |
R203 |
T365 |
T364 |
amod |
transcriptional,events |
R211 |
T373 |
T371 |
pobj |
skin,within |
R212 |
T374 |
T373 |
amod |
developing,skin |
R227 |
T1019 |
T1020 |
amod |
Mammalian,development |
R228 |
T1020 |
T1021 |
nsubj |
development,involves |
R229 |
T1022 |
T1023 |
det |
the,morphogenesis |
R230 |
T1023 |
T1021 |
dobj |
morphogenesis,involves |
R231 |
T1024 |
T1023 |
prep |
of,morphogenesis |
R232 |
T1025 |
T1026 |
amod |
complex,structures |
R233 |
T1026 |
T1024 |
pobj |
structures,of |
R234 |
T1027 |
T1028 |
advmod |
three,dimensional |
R235 |
T1028 |
T1026 |
amod |
dimensional,structures |
R236 |
T1029 |
T1028 |
punct |
-,dimensional |
R237 |
T1030 |
T1023 |
prep |
from,morphogenesis |
R238 |
T1031 |
T1032 |
advmod |
seemingly,uniform |
R239 |
T1032 |
T1033 |
amod |
uniform,sheets |
R240 |
T1033 |
T1030 |
pobj |
sheets,from |
R241 |
T1034 |
T1033 |
cc |
or,sheets |
R242 |
T1035 |
T1033 |
conj |
masses,sheets |
R243 |
T1036 |
T1033 |
prep |
of,sheets |
R244 |
T1037 |
T1036 |
pobj |
cells,of |
R245 |
T1038 |
T1021 |
punct |
.,involves |
R246 |
T1040 |
T1041 |
det |
A,structure |
R247 |
T1041 |
T1046 |
nsubj |
structure,initiates |
R248 |
T1042 |
T1041 |
amod |
simple,structure |
R249 |
T1043 |
T1044 |
npadvmod |
bud,like |
R250 |
T1044 |
T1041 |
amod |
like,structure |
R251 |
T1045 |
T1044 |
punct |
-,like |
R252 |
T1047 |
T1048 |
det |
the,formation |
R253 |
T1048 |
T1046 |
dobj |
formation,initiates |
R254 |
T1049 |
T1048 |
prep |
of,formation |
R255 |
T1050 |
T1051 |
amod |
many,organs |
R256 |
T1051 |
T1049 |
pobj |
organs,of |
R257 |
T1052 |
T1051 |
punct |
", ",organs |
R258 |
T1053 |
T1051 |
prep |
including,organs |
R259 |
T1054 |
T1053 |
pobj |
lungs,including |
R260 |
T1055 |
T1054 |
punct |
", ",lungs |
R261 |
T1056 |
T1057 |
amod |
spinal,cord |
R262 |
T1057 |
T1054 |
conj |
cord,lungs |
R263 |
T1058 |
T1057 |
punct |
", ",cord |
R264 |
T1059 |
T1060 |
amod |
mammary,glands |
R265 |
T1060 |
T1057 |
conj |
glands,cord |
R266 |
T1061 |
T1060 |
punct |
", ",glands |
R267 |
T1062 |
T1060 |
cc |
and,glands |
R268 |
T1063 |
T1064 |
compound |
hair,follicles |
R269 |
T1064 |
T1060 |
conj |
follicles,glands |
R270 |
T1065 |
T1066 |
punct |
[,1 |
R271 |
T1066 |
T1046 |
parataxis |
1,initiates |
R272 |
T1067 |
T1066 |
punct |
],1 |
R273 |
T1068 |
T1046 |
punct |
.,initiates |
R274 |
T1070 |
T1071 |
det |
The,cells |
R275 |
T1071 |
T1076 |
nsubjpass |
cells,attached |
R276 |
T1072 |
T1071 |
amod |
multipotent,cells |
R277 |
T1073 |
T1071 |
punct |
", ",cells |
R278 |
T1074 |
T1071 |
amod |
adhering,cells |
R279 |
T1075 |
T1071 |
amod |
epithelial,cells |
R280 |
T1077 |
T1076 |
auxpass |
are,attached |
R281 |
T1078 |
T1076 |
advmod |
typically,attached |
R282 |
T1079 |
T1076 |
prep |
to,attached |
R283 |
T1080 |
T1081 |
det |
an,lamina |
R284 |
T1081 |
T1079 |
pobj |
lamina,to |
R285 |
T1082 |
T1081 |
amod |
underlying,lamina |
R286 |
T1083 |
T1081 |
amod |
basal,lamina |
R287 |
T1084 |
T1085 |
dep |
that,polarizes |
R288 |
T1085 |
T1081 |
relcl |
polarizes,lamina |
R289 |
T1086 |
T1087 |
det |
the,sheet |
R290 |
T1087 |
T1085 |
dobj |
sheet,polarizes |
R291 |
T1088 |
T1087 |
amod |
epithelial,sheet |
R292 |
T1089 |
T1085 |
cc |
and,polarizes |
R293 |
T1090 |
T1085 |
conj |
separates,polarizes |
R294 |
T1091 |
T1090 |
dobj |
it,separates |
R295 |
T1092 |
T1090 |
prep |
from,separates |
R296 |
T1093 |
T1094 |
amod |
surrounding,mesenchyme |
R297 |
T1094 |
T1092 |
pobj |
mesenchyme,from |
R298 |
T1095 |
T1076 |
punct |
.,attached |
R299 |
T1097 |
T1098 |
amod |
Budding,morphogenesis |
R300 |
T1098 |
T1099 |
nsubjpass |
morphogenesis,guided |
R301 |
T1100 |
T1099 |
auxpass |
is,guided |
R302 |
T1101 |
T1099 |
agent |
by,guided |
R303 |
T1102 |
T1103 |
det |
a,exchange |
R304 |
T1103 |
T1101 |
pobj |
exchange,by |
R305 |
T1104 |
T1103 |
amod |
reciprocal,exchange |
R306 |
T1105 |
T1103 |
prep |
of,exchange |
R307 |
T1106 |
T1105 |
pobj |
signals,of |
R308 |
T1107 |
T1103 |
prep |
between,exchange |
R309 |
T1108 |
T1107 |
pobj |
epithelium,between |
R310 |
T1109 |
T1108 |
cc |
and,epithelium |
R311 |
T1110 |
T1108 |
conj |
mesenchyme,epithelium |
R312 |
T1111 |
T1112 |
aux |
to,specify |
R313 |
T1112 |
T1103 |
advcl |
specify,exchange |
R314 |
T1113 |
T1114 |
det |
the,identity |
R315 |
T1114 |
T1112 |
dobj |
identity,specify |
R316 |
T1115 |
T1114 |
prep |
of,identity |
R317 |
T1116 |
T1117 |
det |
the,organ |
R318 |
T1117 |
T1115 |
pobj |
organ,of |
R319 |
T1118 |
T1119 |
dep |
that,form |
R320 |
T1119 |
T1117 |
relcl |
form,organ |
R321 |
T1120 |
T1119 |
aux |
will,form |
R322 |
T1121 |
T1112 |
cc |
and,specify |
R323 |
T1122 |
T1123 |
aux |
to,govern |
R324 |
T1123 |
T1112 |
conj |
govern,specify |
R325 |
T1124 |
T1125 |
poss |
its,growth |
R326 |
T1125 |
T1123 |
dobj |
growth,govern |
R327 |
T1126 |
T1099 |
punct |
.,guided |
R328 |
T1128 |
T1129 |
prep |
At,are |
R329 |
T1130 |
T1131 |
det |
the,helm |
R330 |
T1131 |
T1128 |
pobj |
helm,At |
R331 |
T1132 |
T1131 |
prep |
of,helm |
R332 |
T1133 |
T1134 |
det |
these,pathways |
R333 |
T1134 |
T1132 |
pobj |
pathways,of |
R334 |
T1135 |
T1136 |
amod |
molecular,communication |
R335 |
T1136 |
T1134 |
compound |
communication,pathways |
R336 |
T1137 |
T1129 |
nsubj |
Wnts,are |
R337 |
T1138 |
T1137 |
punct |
", ",Wnts |
R338 |
T1139 |
T1140 |
nmod |
bone,proteins |
R339 |
T1140 |
T1137 |
appos |
proteins,Wnts |
R340 |
T1141 |
T1140 |
amod |
morphogenic,proteins |
R341 |
T1142 |
T1140 |
punct |
(,proteins |
R342 |
T1143 |
T1140 |
appos |
BMPs,proteins |
R343 |
T1144 |
T1137 |
punct |
),Wnts |
R344 |
T1145 |
T1137 |
punct |
", ",Wnts |
R345 |
T1146 |
T1147 |
amod |
transforming,βs |
R346 |
T1147 |
T1137 |
appos |
βs,Wnts |
R347 |
T1148 |
T1147 |
compound |
growth,βs |
R348 |
T1149 |
T1147 |
compound |
factor,βs |
R349 |
T1150 |
T1147 |
punct |
(,βs |
R350 |
T1151 |
T1152 |
compound |
TGF,βs |
R351 |
T1152 |
T1147 |
appos |
βs,βs |
R352 |
T1153 |
T1152 |
punct |
-,βs |
R353 |
T1154 |
T1137 |
punct |
),Wnts |
R354 |
T1155 |
T1137 |
punct |
", ",Wnts |
R355 |
T1156 |
T1137 |
cc |
and,Wnts |
R356 |
T1157 |
T1158 |
compound |
fibroblast,factors |
R357 |
T1158 |
T1137 |
conj |
factors,Wnts |
R358 |
T1159 |
T1158 |
compound |
growth,factors |
R359 |
T1160 |
T1158 |
punct |
(,factors |
R360 |
T1161 |
T1158 |
appos |
FGFs,factors |
R361 |
T1162 |
T1129 |
punct |
),are |
R362 |
T1163 |
T1129 |
punct |
.,are |
R363 |
T1165 |
T1166 |
prep |
Through,trigger |
R364 |
T1167 |
T1165 |
pobj |
activation,Through |
R365 |
T1168 |
T1167 |
prep |
of,activation |
R366 |
T1169 |
T1170 |
compound |
cell,surface |
R367 |
T1170 |
T1171 |
compound |
surface,receptors |
R368 |
T1171 |
T1168 |
pobj |
receptors,of |
R369 |
T1172 |
T1171 |
compound |
transmembrane,receptors |
R370 |
T1173 |
T1166 |
punct |
", ",trigger |
R371 |
T1174 |
T1175 |
det |
these,molecules |
R372 |
T1175 |
T1166 |
nsubj |
molecules,trigger |
R373 |
T1176 |
T1175 |
amod |
external,molecules |
R374 |
T1177 |
T1175 |
compound |
signaling,molecules |
R375 |
T1178 |
T1179 |
amod |
distinct,cascades |
R376 |
T1179 |
T1166 |
dobj |
cascades,trigger |
R377 |
T1180 |
T1179 |
prep |
of,cascades |
R378 |
T1181 |
T1182 |
amod |
intracellular,events |
R379 |
T1182 |
T1180 |
pobj |
events,of |
R380 |
T1183 |
T1184 |
dep |
that,culminate |
R381 |
T1184 |
T1179 |
relcl |
culminate,cascades |
R382 |
T1185 |
T1184 |
prep |
in,culminate |
R383 |
T1186 |
T1185 |
pobj |
changes,in |
R384 |
T1187 |
T1186 |
prep |
in,changes |
R385 |
T1188 |
T1189 |
compound |
gene,expression |
R386 |
T1189 |
T1187 |
pobj |
expression,in |
R387 |
T1190 |
T1189 |
punct |
", ",expression |
R388 |
T1191 |
T1189 |
conj |
growth,expression |
R389 |
T1192 |
T1191 |
punct |
", ",growth |
R390 |
T1193 |
T1191 |
cc |
and,growth |
R391 |
T1194 |
T1191 |
conj |
differentiation,growth |
R392 |
T1195 |
T1196 |
punct |
[,2 |
R393 |
T1196 |
T1166 |
parataxis |
2,trigger |
R394 |
T1197 |
T1196 |
punct |
],2 |
R395 |
T1198 |
T1166 |
punct |
.,trigger |
R396 |
T1200 |
T1201 |
advmod |
How,collaborates |
R397 |
T1201 |
T1206 |
csubj |
collaborates,has |
R398 |
T1202 |
T1203 |
det |
this,constellation |
R399 |
T1203 |
T1201 |
nsubj |
constellation,collaborates |
R400 |
T1204 |
T1203 |
prep |
of,constellation |
R401 |
T1205 |
T1204 |
pobj |
signals,of |
R402 |
T1207 |
T1201 |
prep |
in,collaborates |
R403 |
T1208 |
T1207 |
pcomp |
tailoring,in |
R404 |
T1209 |
T1210 |
det |
each,process |
R405 |
T1210 |
T1208 |
dobj |
process,tailoring |
R406 |
T1211 |
T1210 |
compound |
budding,process |
R407 |
T1212 |
T1213 |
mark |
so,executes |
R408 |
T1213 |
T1208 |
advcl |
executes,tailoring |
R409 |
T1214 |
T1213 |
mark |
that,executes |
R410 |
T1215 |
T1213 |
nsubj |
it,executes |
R411 |
T1216 |
T1217 |
det |
a,program |
R412 |
T1217 |
T1213 |
dobj |
program,executes |
R413 |
T1218 |
T1217 |
amod |
distinct,program |
R414 |
T1219 |
T1217 |
amod |
morphogenetic,program |
R415 |
T1220 |
T1221 |
advmod |
yet,defined |
R416 |
T1221 |
T1206 |
xcomp |
defined,has |
R417 |
T1222 |
T1221 |
aux |
to,defined |
R418 |
T1223 |
T1221 |
auxpass |
be,defined |
R419 |
T1224 |
T1221 |
advmod |
comprehensively,defined |
R420 |
T1225 |
T1206 |
punct |
.,has |
R421 |
T1227 |
T1228 |
advmod |
However,appears |
R422 |
T1229 |
T1228 |
punct |
", ",appears |
R423 |
T1230 |
T1231 |
det |
the,process |
R424 |
T1231 |
T1228 |
nsubj |
process,appears |
R425 |
T1232 |
T1233 |
aux |
to,patterned |
R426 |
T1233 |
T1228 |
xcomp |
patterned,appears |
R427 |
T1234 |
T1233 |
auxpass |
be,patterned |
R428 |
T1235 |
T1233 |
prep |
at,patterned |
R429 |
T1236 |
T1237 |
det |
the,stages |
R430 |
T1237 |
T1235 |
pobj |
stages,at |
R431 |
T1238 |
T1237 |
amod |
initial,stages |
R432 |
T1239 |
T1237 |
prep |
of,stages |
R433 |
T1240 |
T1241 |
compound |
bud,formation |
R434 |
T1241 |
T1239 |
pobj |
formation,of |
R435 |
T1242 |
T1228 |
punct |
", ",appears |
R436 |
T1243 |
T1244 |
mark |
since,differ |
R437 |
T1244 |
T1228 |
advcl |
differ,appears |
R438 |
T1245 |
T1246 |
det |
the,importance |
R439 |
T1246 |
T1244 |
nsubj |
importance,differ |
R440 |
T1247 |
T1246 |
amod |
relative,importance |
R441 |
T1248 |
T1246 |
prep |
of,importance |
R442 |
T1249 |
T1250 |
det |
these,pathways |
R443 |
T1250 |
T1248 |
pobj |
pathways,of |
R444 |
T1251 |
T1250 |
cc |
and,pathways |
R445 |
T1252 |
T1253 |
poss |
their,effectors |
R446 |
T1253 |
T1250 |
conj |
effectors,pathways |
R447 |
T1254 |
T1253 |
amod |
downstream,effectors |
R448 |
T1255 |
T1256 |
mark |
as,begin |
R449 |
T1256 |
T1244 |
advcl |
begin,differ |
R450 |
T1257 |
T1256 |
nsubj |
buds,begin |
R451 |
T1258 |
T1259 |
aux |
to,develop |
R452 |
T1259 |
T1256 |
xcomp |
develop,begin |
R453 |
T1260 |
T1256 |
cc |
and,begin |
R454 |
T1261 |
T1262 |
compound |
cell,fates |
R455 |
T1262 |
T1263 |
nsubjpass |
fates,specified |
R456 |
T1263 |
T1256 |
conj |
specified,begin |
R457 |
T1264 |
T1263 |
auxpass |
are,specified |
R458 |
T1265 |
T1228 |
punct |
.,appears |
R459 |
T1267 |
T1268 |
det |
The,development |
R460 |
T1268 |
T1269 |
nsubj |
development,requires |
R461 |
T1270 |
T1268 |
prep |
of,development |
R462 |
T1271 |
T1272 |
det |
a,bud |
R463 |
T1272 |
T1270 |
pobj |
bud,of |
R464 |
T1273 |
T1274 |
det |
a,number |
R465 |
T1274 |
T1269 |
dobj |
number,requires |
R466 |
T1275 |
T1274 |
prep |
of,number |
R467 |
T1276 |
T1277 |
amod |
coordinated,changes |
R468 |
T1277 |
T1275 |
pobj |
changes,of |
R469 |
T1278 |
T1277 |
prep |
in,changes |
R470 |
T1279 |
T1280 |
det |
the,behavior |
R471 |
T1280 |
T1278 |
pobj |
behavior,in |
R472 |
T1281 |
T1280 |
prep |
of,behavior |
R473 |
T1282 |
T1283 |
det |
the,cells |
R474 |
T1283 |
T1281 |
pobj |
cells,of |
R475 |
T1284 |
T1283 |
amod |
targeted,cells |
R476 |
T1285 |
T1280 |
prep |
within,behavior |
R477 |
T1286 |
T1287 |
det |
an,sheet |
R478 |
T1287 |
T1285 |
pobj |
sheet,within |
R479 |
T1288 |
T1287 |
amod |
epithelial,sheet |
R480 |
T1289 |
T1269 |
punct |
.,requires |
R481 |
T1291 |
T1292 |
det |
The,process |
R482 |
T1292 |
T1293 |
nsubjpass |
process,accompanied |
R483 |
T1294 |
T1293 |
aux |
must,accompanied |
R484 |
T1295 |
T1293 |
auxpass |
be,accompanied |
R485 |
T1296 |
T1293 |
prep |
by,accompanied |
R486 |
T1297 |
T1296 |
pobj |
alterations,by |
R487 |
T1298 |
T1297 |
prep |
in,alterations |
R488 |
T1299 |
T1300 |
det |
the,proliferation |
R489 |
T1300 |
T1298 |
pobj |
proliferation,in |
R490 |
T1301 |
T1300 |
punct |
", ",proliferation |
R491 |
T1302 |
T1300 |
conj |
polarity,proliferation |
R492 |
T1303 |
T1302 |
punct |
", ",polarity |
R493 |
T1304 |
T1302 |
conj |
shape,polarity |
R494 |
T1305 |
T1304 |
punct |
", ",shape |
R495 |
T1306 |
T1304 |
cc |
and,shape |
R496 |
T1307 |
T1304 |
conj |
adhesiveness,shape |
R497 |
T1308 |
T1300 |
prep |
of,proliferation |
R498 |
T1309 |
T1310 |
amod |
selected,cells |
R499 |
T1310 |
T1308 |
pobj |
cells,of |
R500 |
T1311 |
T1296 |
punct |
", ",by |
R501 |
T1312 |
T1313 |
advmod |
as,as |
R502 |
T1313 |
T1296 |
cc |
as,by |
R503 |
T1314 |
T1313 |
advmod |
well,as |
R504 |
T1315 |
T1296 |
conj |
by,by |
R505 |
T1316 |
T1315 |
pobj |
modifications,by |
R506 |
T1317 |
T1316 |
prep |
in,modifications |
R507 |
T1318 |
T1319 |
poss |
their,lamina |
R508 |
T1319 |
T1317 |
pobj |
lamina,in |
R509 |
T1320 |
T1319 |
amod |
underlying,lamina |
R510 |
T1321 |
T1319 |
amod |
basal,lamina |
R511 |
T1322 |
T1293 |
punct |
.,accompanied |
R512 |
T1324 |
T1325 |
advmod |
Thus,orchestrated |
R513 |
T1326 |
T1325 |
punct |
", ",orchestrated |
R514 |
T1327 |
T1328 |
amod |
extracellular,crosstalk |
R515 |
T1328 |
T1325 |
nsubjpass |
crosstalk,orchestrated |
R516 |
T1329 |
T1330 |
amod |
epithelial,mesenchymal |
R517 |
T1330 |
T1328 |
amod |
mesenchymal,crosstalk |
R518 |
T1331 |
T1330 |
punct |
-,mesenchymal |
R519 |
T1332 |
T1325 |
aux |
must,orchestrated |
R520 |
T1333 |
T1325 |
auxpass |
be,orchestrated |
R521 |
T1334 |
T1325 |
advmod |
intricately,orchestrated |
R522 |
T1335 |
T1336 |
aux |
to,couple |
R523 |
T1336 |
T1325 |
advcl |
couple,orchestrated |
R524 |
T1337 |
T1338 |
det |
the,determination |
R525 |
T1338 |
T1336 |
dobj |
determination,couple |
R526 |
T1339 |
T1338 |
prep |
of,determination |
R198 |
T360 |
T361 |
amod |
different,morphogens |
R199 |
T361 |
T359 |
pobj |
morphogens,of |
R200 |
T362 |
T361 |
cc |
and,morphogens |
R201 |
T363 |
T364 |
amod |
downstream,events |
R204 |
T366 |
T367 |
dep |
that,guide |
R206 |
T368 |
T369 |
compound |
hair,bud |
R207 |
T369 |
T370 |
compound |
bud,formation |
R208 |
T370 |
T367 |
dobj |
formation,guide |
R209 |
T371 |
T367 |
prep |
within,guide |
R210 |
T372 |
T373 |
det |
the,skin |