Id |
Subject |
Object |
Predicate |
Lexical cue |
T1953 |
0-1 |
DT |
denotes |
A |
T1954 |
10-16 |
NN |
denotes |
system |
T1955 |
2-9 |
JJ |
denotes |
genetic |
T1956 |
90-92 |
VB |
denotes |
be |
T1957 |
17-20 |
IN |
denotes |
for |
T1958 |
21-31 |
VBG |
denotes |
activating |
T1959 |
32-34 |
CC |
denotes |
or |
T1960 |
35-47 |
VBG |
denotes |
inactivating |
T1961 |
48-53 |
NNS |
denotes |
genes |
T1962 |
54-66 |
RB |
denotes |
specifically |
T1963 |
67-69 |
IN |
denotes |
in |
T1964 |
70-75 |
NN |
denotes |
joint |
T1965 |
76-83 |
NNS |
denotes |
tissues |
T1966 |
84-89 |
MD |
denotes |
would |
T1967 |
93-105 |
RB |
denotes |
particularly |
T1968 |
106-112 |
JJ |
denotes |
useful |
T1969 |
113-116 |
IN |
denotes |
for |
T1970 |
117-124 |
JJ |
denotes |
further |
T1971 |
125-132 |
NNS |
denotes |
studies |
T1972 |
133-135 |
IN |
denotes |
of |
T1973 |
136-141 |
NN |
denotes |
joint |
T1974 |
142-151 |
NN |
denotes |
formation |
T1975 |
152-155 |
CC |
denotes |
and |
T1976 |
156-167 |
NN |
denotes |
maintenance |
T1977 |
167-168 |
. |
denotes |
. |
T1978 |
168-370 |
sentence |
denotes |
Here we take advantage of the tissue-specific expression pattern of the Gdf5 gene to engineer a Cre/loxP system (Nagy 2000), Gdf5-Cre, that can be used to remove or ectopically express genes in joints. |
T1979 |
169-173 |
RB |
denotes |
Here |
T1980 |
177-181 |
VBP |
denotes |
take |
T1981 |
174-176 |
PRP |
denotes |
we |
T1982 |
182-191 |
NN |
denotes |
advantage |
T1983 |
192-194 |
IN |
denotes |
of |
T1984 |
195-198 |
DT |
denotes |
the |
T1985 |
226-233 |
NN |
denotes |
pattern |
T1986 |
199-205 |
NN |
denotes |
tissue |
T1987 |
206-214 |
JJ |
denotes |
specific |
T1988 |
205-206 |
HYPH |
denotes |
- |
T1989 |
215-225 |
NN |
denotes |
expression |
T1990 |
234-236 |
IN |
denotes |
of |
T1991 |
237-240 |
DT |
denotes |
the |
T1992 |
246-250 |
NN |
denotes |
gene |
T1993 |
241-245 |
NN |
denotes |
Gdf5 |
T1994 |
251-253 |
TO |
denotes |
to |
T1995 |
254-262 |
VB |
denotes |
engineer |
T1996 |
263-264 |
DT |
denotes |
a |
T1997 |
274-280 |
NN |
denotes |
system |
T1998 |
265-268 |
NN |
denotes |
Cre |
T1999 |
269-273 |
NN |
denotes |
loxP |
T2000 |
268-269 |
HYPH |
denotes |
/ |
T2001 |
281-282 |
-LRB- |
denotes |
( |
T2002 |
282-286 |
NNP |
denotes |
Nagy |
T2003 |
287-291 |
CD |
denotes |
2000 |
T2004 |
291-292 |
-RRB- |
denotes |
) |
T2005 |
292-294 |
, |
denotes |
, |
T2006 |
294-298 |
NN |
denotes |
Gdf5 |
T2007 |
299-302 |
NN |
denotes |
Cre |
T2008 |
298-299 |
HYPH |
denotes |
- |
T2009 |
302-304 |
, |
denotes |
, |
T2010 |
304-308 |
WDT |
denotes |
that |
T2011 |
316-320 |
VBN |
denotes |
used |
T2012 |
309-312 |
MD |
denotes |
can |
T2013 |
313-315 |
VB |
denotes |
be |
T2014 |
321-323 |
TO |
denotes |
to |
T2015 |
324-330 |
VB |
denotes |
remove |
T2016 |
331-333 |
CC |
denotes |
or |
T2017 |
334-345 |
RB |
denotes |
ectopically |
T2018 |
346-353 |
VB |
denotes |
express |
T2019 |
354-359 |
NNS |
denotes |
genes |
T2020 |
360-362 |
IN |
denotes |
in |
T2021 |
363-369 |
NNS |
denotes |
joints |
T2022 |
369-370 |
. |
denotes |
. |
T2023 |
370-600 |
sentence |
denotes |
Tests with reporter mice show that this system is capable of modifying genes in all of the structures of the mature synovial joint, including the ligaments of the joint capsule, the synovial membrane, and the articular cartilage. |
T2024 |
371-376 |
NNS |
denotes |
Tests |
T2025 |
396-400 |
VBP |
denotes |
show |
T2026 |
377-381 |
IN |
denotes |
with |
T2027 |
382-390 |
NN |
denotes |
reporter |
T2028 |
391-395 |
NNS |
denotes |
mice |
T2029 |
401-405 |
IN |
denotes |
that |
T2030 |
418-420 |
VBZ |
denotes |
is |
T2031 |
406-410 |
DT |
denotes |
this |
T2032 |
411-417 |
NN |
denotes |
system |
T2033 |
421-428 |
JJ |
denotes |
capable |
T2034 |
429-431 |
IN |
denotes |
of |
T2035 |
432-441 |
VBG |
denotes |
modifying |
T2036 |
442-447 |
NNS |
denotes |
genes |
T2037 |
448-450 |
IN |
denotes |
in |
T2038 |
451-454 |
DT |
denotes |
all |
T2039 |
455-457 |
IN |
denotes |
of |
T2040 |
458-461 |
DT |
denotes |
the |
T2041 |
462-472 |
NNS |
denotes |
structures |
T2042 |
473-475 |
IN |
denotes |
of |
T2043 |
476-479 |
DT |
denotes |
the |
T2044 |
496-501 |
NN |
denotes |
joint |
T2045 |
480-486 |
JJ |
denotes |
mature |
T2046 |
487-495 |
JJ |
denotes |
synovial |
T2047 |
501-503 |
, |
denotes |
, |
T2048 |
503-512 |
VBG |
denotes |
including |
T2049 |
513-516 |
DT |
denotes |
the |
T2050 |
517-526 |
NNS |
denotes |
ligaments |
T2051 |
527-529 |
IN |
denotes |
of |
T2052 |
530-533 |
DT |
denotes |
the |
T2053 |
540-547 |
NN |
denotes |
capsule |
T2054 |
534-539 |
NN |
denotes |
joint |
T2055 |
547-549 |
, |
denotes |
, |
T2056 |
549-552 |
DT |
denotes |
the |
T2057 |
562-570 |
NN |
denotes |
membrane |
T2058 |
553-561 |
JJ |
denotes |
synovial |
T2059 |
570-572 |
, |
denotes |
, |
T2060 |
572-575 |
CC |
denotes |
and |
T2061 |
576-579 |
DT |
denotes |
the |
T2062 |
590-599 |
NN |
denotes |
cartilage |
T2063 |
580-589 |
JJ |
denotes |
articular |
T2064 |
599-600 |
. |
denotes |
. |
T2065 |
600-847 |
sentence |
denotes |
Gdf5-Cre recombination bypasses the early embryonic lethality of null mutations in Bmpr1a, and shows that this receptor is required for early joint formation at some locations and for initiation of programmed cell death in webbing between digits. |
T2066 |
601-605 |
NN |
denotes |
Gdf5 |
T2067 |
606-609 |
NN |
denotes |
Cre |
T2068 |
605-606 |
HYPH |
denotes |
- |
T2069 |
610-623 |
NN |
denotes |
recombination |
T2070 |
624-632 |
VBZ |
denotes |
bypasses |
T2071 |
633-636 |
DT |
denotes |
the |
T2072 |
653-662 |
NN |
denotes |
lethality |
T2073 |
637-642 |
JJ |
denotes |
early |
T2074 |
643-652 |
JJ |
denotes |
embryonic |
T2075 |
663-665 |
IN |
denotes |
of |
T2076 |
666-670 |
JJ |
denotes |
null |
T2077 |
671-680 |
NNS |
denotes |
mutations |
T2078 |
681-683 |
IN |
denotes |
in |
T2079 |
684-690 |
NN |
denotes |
Bmpr1a |
T2080 |
690-692 |
, |
denotes |
, |
T2081 |
692-695 |
CC |
denotes |
and |
T2082 |
696-701 |
VBZ |
denotes |
shows |
T2083 |
702-706 |
IN |
denotes |
that |
T2084 |
724-732 |
VBN |
denotes |
required |
T2085 |
707-711 |
DT |
denotes |
this |
T2086 |
712-720 |
NN |
denotes |
receptor |
T2087 |
721-723 |
VBZ |
denotes |
is |
T2088 |
733-736 |
IN |
denotes |
for |
T2089 |
737-742 |
JJ |
denotes |
early |
T2090 |
749-758 |
NN |
denotes |
formation |
T2091 |
743-748 |
NN |
denotes |
joint |
T2092 |
759-761 |
IN |
denotes |
at |
T2093 |
762-766 |
DT |
denotes |
some |
T2094 |
767-776 |
NNS |
denotes |
locations |
T2095 |
777-780 |
CC |
denotes |
and |
T2096 |
781-784 |
IN |
denotes |
for |
T2097 |
785-795 |
NN |
denotes |
initiation |
T2098 |
796-798 |
IN |
denotes |
of |
T2099 |
799-809 |
VBN |
denotes |
programmed |
T2100 |
815-820 |
NN |
denotes |
death |
T2101 |
810-814 |
NN |
denotes |
cell |
T2102 |
821-823 |
IN |
denotes |
in |
T2103 |
824-831 |
VBG |
denotes |
webbing |
T2104 |
832-839 |
IN |
denotes |
between |
T2105 |
840-846 |
NNS |
denotes |
digits |
T2106 |
846-847 |
. |
denotes |
. |
T2107 |
847-968 |
sentence |
denotes |
Interestingly, Bmpr1a is also required for postnatal maintenance of articular cartilage throughout most of the skeleton. |
T2108 |
848-861 |
RB |
denotes |
Interestingly |
T2109 |
870-872 |
VBZ |
denotes |
is |
T2110 |
861-863 |
, |
denotes |
, |
T2111 |
863-869 |
NN |
denotes |
Bmpr1a |
T2112 |
873-877 |
RB |
denotes |
also |
T2113 |
878-886 |
VBN |
denotes |
required |
T2114 |
887-890 |
IN |
denotes |
for |
T2115 |
891-900 |
JJ |
denotes |
postnatal |
T2116 |
901-912 |
NN |
denotes |
maintenance |
T2117 |
913-915 |
IN |
denotes |
of |
T2118 |
916-925 |
JJ |
denotes |
articular |
T2119 |
926-935 |
NN |
denotes |
cartilage |
T2120 |
936-946 |
IN |
denotes |
throughout |
T2121 |
947-951 |
JJS |
denotes |
most |
T2122 |
952-954 |
IN |
denotes |
of |
T2123 |
955-958 |
DT |
denotes |
the |
T2124 |
959-967 |
NN |
denotes |
skeleton |
T2125 |
967-968 |
. |
denotes |
. |
T2126 |
968-1124 |
sentence |
denotes |
In Gdf5-Cre/Bmpr1afloxP mice, articular cartilage initially forms normally, but subsequently loses expression of several key cartilage markers after birth. |
T2127 |
969-971 |
IN |
denotes |
In |
T2128 |
1029-1034 |
VBZ |
denotes |
forms |
T2129 |
972-976 |
NN |
denotes |
Gdf5 |
T2130 |
977-980 |
NN |
denotes |
Cre |
T2131 |
976-977 |
HYPH |
denotes |
- |
T2132 |
993-997 |
NNS |
denotes |
mice |
T2133 |
980-981 |
HYPH |
denotes |
/ |
T2134 |
981-992 |
NN |
denotes |
Bmpr1afloxP |
T2135 |
997-999 |
, |
denotes |
, |
T2136 |
999-1008 |
JJ |
denotes |
articular |
T2137 |
1009-1018 |
NN |
denotes |
cartilage |
T2138 |
1019-1028 |
RB |
denotes |
initially |
T2139 |
1035-1043 |
RB |
denotes |
normally |
T2140 |
1043-1045 |
, |
denotes |
, |
T2141 |
1045-1048 |
CC |
denotes |
but |
T2142 |
1049-1061 |
RB |
denotes |
subsequently |
T2143 |
1062-1067 |
VBZ |
denotes |
loses |
T2144 |
1068-1078 |
NN |
denotes |
expression |
T2145 |
1079-1081 |
IN |
denotes |
of |
T2146 |
1082-1089 |
JJ |
denotes |
several |
T2147 |
1104-1111 |
NNS |
denotes |
markers |
T2148 |
1090-1093 |
JJ |
denotes |
key |
T2149 |
1094-1103 |
NN |
denotes |
cartilage |
T2150 |
1112-1117 |
IN |
denotes |
after |
T2151 |
1118-1123 |
NN |
denotes |
birth |
T2152 |
1123-1124 |
. |
denotes |
. |
T2153 |
1124-1224 |
sentence |
denotes |
It ultimately fibrillates and degenerates, resulting in severe osteoarthritis and loss of mobility. |
T2154 |
1125-1127 |
PRP |
denotes |
It |
T2155 |
1139-1150 |
VBZ |
denotes |
fibrillates |
T2156 |
1128-1138 |
RB |
denotes |
ultimately |
T2157 |
1151-1154 |
CC |
denotes |
and |
T2158 |
1155-1166 |
VBZ |
denotes |
degenerates |
T2159 |
1166-1168 |
, |
denotes |
, |
T2160 |
1168-1177 |
VBG |
denotes |
resulting |
T2161 |
1178-1180 |
IN |
denotes |
in |
T2162 |
1181-1187 |
JJ |
denotes |
severe |
T2163 |
1188-1202 |
NN |
denotes |
osteoarthritis |
T2164 |
1203-1206 |
CC |
denotes |
and |
T2165 |
1207-1211 |
NN |
denotes |
loss |
T2166 |
1212-1214 |
IN |
denotes |
of |
T2167 |
1215-1223 |
NN |
denotes |
mobility |
T2168 |
1223-1224 |
. |
denotes |
. |
T2169 |
1224-1432 |
sentence |
denotes |
These experiments suggest that BMP signaling is required for normal maintenance of postnatal articular cartilage, and that modulation of the BMP signaling pathway may play an important role in joint disease. |
T2170 |
1225-1230 |
DT |
denotes |
These |
T2171 |
1231-1242 |
NNS |
denotes |
experiments |
T2172 |
1243-1250 |
VBP |
denotes |
suggest |
T2173 |
1251-1255 |
IN |
denotes |
that |
T2174 |
1273-1281 |
VBN |
denotes |
required |
T2175 |
1256-1259 |
NN |
denotes |
BMP |
T2176 |
1260-1269 |
NN |
denotes |
signaling |
T2177 |
1270-1272 |
VBZ |
denotes |
is |
T2178 |
1282-1285 |
IN |
denotes |
for |
T2179 |
1286-1292 |
JJ |
denotes |
normal |
T2180 |
1293-1304 |
NN |
denotes |
maintenance |
T2181 |
1305-1307 |
IN |
denotes |
of |
T2182 |
1308-1317 |
JJ |
denotes |
postnatal |
T2183 |
1328-1337 |
NN |
denotes |
cartilage |
T2184 |
1318-1327 |
JJ |
denotes |
articular |
T2185 |
1337-1339 |
, |
denotes |
, |
T2186 |
1339-1342 |
CC |
denotes |
and |
T2187 |
1343-1347 |
IN |
denotes |
that |
T2188 |
1392-1396 |
VB |
denotes |
play |
T2189 |
1348-1358 |
NN |
denotes |
modulation |
T2190 |
1359-1361 |
IN |
denotes |
of |
T2191 |
1362-1365 |
DT |
denotes |
the |
T2192 |
1380-1387 |
NN |
denotes |
pathway |
T2193 |
1366-1369 |
NN |
denotes |
BMP |
T2194 |
1370-1379 |
NN |
denotes |
signaling |
T2195 |
1388-1391 |
MD |
denotes |
may |
T2196 |
1397-1399 |
DT |
denotes |
an |
T2197 |
1410-1414 |
NN |
denotes |
role |
T2198 |
1400-1409 |
JJ |
denotes |
important |
T2199 |
1415-1417 |
IN |
denotes |
in |
T2200 |
1418-1423 |
NN |
denotes |
joint |
T2201 |
1424-1431 |
NN |
denotes |
disease |
T2202 |
1431-1432 |
. |
denotes |
. |
R1140 |
T2014 |
T2015 |
aux |
to,remove |
R1141 |
T2015 |
T2011 |
advcl |
remove,used |
R1143 |
T2016 |
T2015 |
cc |
or,remove |
R1144 |
T2017 |
T2018 |
advmod |
ectopically,express |
R1145 |
T2018 |
T2015 |
conj |
express,remove |
R1147 |
T2019 |
T2018 |
dobj |
genes,express |
R1148 |
T2020 |
T2019 |
prep |
in,genes |
R1150 |
T2021 |
T2020 |
pobj |
joints,in |
R1151 |
T2022 |
T1980 |
punct |
.,take |
R1153 |
T2024 |
T2025 |
nsubj |
Tests,show |
R1155 |
T2026 |
T2024 |
prep |
with,Tests |
R1156 |
T2027 |
T2028 |
compound |
reporter,mice |
R1157 |
T2028 |
T2026 |
pobj |
mice,with |
R1159 |
T2029 |
T2030 |
mark |
that,is |
R1160 |
T2030 |
T2025 |
ccomp |
is,show |
R1161 |
T2031 |
T2032 |
det |
this,system |
R1163 |
T2032 |
T2030 |
nsubj |
system,is |
R1164 |
T2033 |
T2030 |
acomp |
capable,is |
R1165 |
T2034 |
T2033 |
prep |
of,capable |
R1166 |
T2035 |
T2034 |
pcomp |
modifying,of |
R1167 |
T2036 |
T2035 |
dobj |
genes,modifying |
R1169 |
T2037 |
T2035 |
prep |
in,modifying |
R1170 |
T2038 |
T2037 |
pobj |
all,in |
R1172 |
T2039 |
T2038 |
prep |
of,all |
R1173 |
T2040 |
T2041 |
det |
the,structures |
R1174 |
T2041 |
T2039 |
pobj |
structures,of |
R1176 |
T2042 |
T2041 |
prep |
of,structures |
R1177 |
T2043 |
T2044 |
det |
the,joint |
R1178 |
T2044 |
T2042 |
pobj |
joint,of |
R1180 |
T2045 |
T2044 |
amod |
mature,joint |
R1181 |
T2046 |
T2044 |
amod |
synovial,joint |
R1183 |
T2047 |
T2038 |
punct |
", ",all |
R1185 |
T2048 |
T2038 |
prep |
including,all |
R1187 |
T2049 |
T2050 |
det |
the,ligaments |
R1188 |
T2050 |
T2048 |
pobj |
ligaments,including |
R1189 |
T2051 |
T2050 |
prep |
of,ligaments |
R1190 |
T2052 |
T2053 |
det |
the,capsule |
R1193 |
T2053 |
T2051 |
pobj |
capsule,of |
R1194 |
T2054 |
T2053 |
compound |
joint,capsule |
R1196 |
T2055 |
T2050 |
punct |
", ",ligaments |
R1197 |
T2056 |
T2057 |
det |
the,membrane |
R1199 |
T2057 |
T2050 |
conj |
membrane,ligaments |
R1200 |
T2058 |
T2057 |
amod |
synovial,membrane |
R1202 |
T2059 |
T2057 |
punct |
", ",membrane |
R1203 |
T2060 |
T2057 |
cc |
and,membrane |
R1204 |
T2061 |
T2062 |
det |
the,cartilage |
R1206 |
T2062 |
T2057 |
conj |
cartilage,membrane |
R1207 |
T2063 |
T2062 |
amod |
articular,cartilage |
R1209 |
T2064 |
T2025 |
punct |
.,show |
R1210 |
T2066 |
T2067 |
compound |
Gdf5,Cre |
R1212 |
T2067 |
T2069 |
compound |
Cre,recombination |
R1213 |
T2068 |
T2067 |
punct |
-,Cre |
R1214 |
T2069 |
T2070 |
nsubj |
recombination,bypasses |
R1216 |
T2071 |
T2072 |
det |
the,lethality |
R1218 |
T2072 |
T2070 |
dobj |
lethality,bypasses |
R1219 |
T2073 |
T2074 |
amod |
early,embryonic |
R1220 |
T2074 |
T2072 |
amod |
embryonic,lethality |
R1222 |
T2075 |
T2072 |
prep |
of,lethality |
R1223 |
T2076 |
T2077 |
amod |
null,mutations |
R1225 |
T2077 |
T2075 |
pobj |
mutations,of |
R1226 |
T2078 |
T2070 |
prep |
in,bypasses |
R1227 |
T2079 |
T2078 |
pobj |
Bmpr1a,in |
R1229 |
T2080 |
T2070 |
punct |
", ",bypasses |
R1230 |
T2081 |
T2070 |
cc |
and,bypasses |
R1231 |
T2082 |
T2070 |
conj |
shows,bypasses |
R1233 |
T2083 |
T2084 |
mark |
that,required |
R1234 |
T2084 |
T2082 |
ccomp |
required,shows |
R1236 |
T2085 |
T2086 |
det |
this,receptor |
R1237 |
T2086 |
T2084 |
nsubjpass |
receptor,required |
R1238 |
T2087 |
T2084 |
auxpass |
is,required |
R1240 |
T2088 |
T2084 |
prep |
for,required |
R1241 |
T2089 |
T2090 |
amod |
early,formation |
R1242 |
T2090 |
T2088 |
pobj |
formation,for |
R1243 |
T2091 |
T2090 |
compound |
joint,formation |
R1244 |
T2092 |
T2090 |
prep |
at,formation |
R1245 |
T2093 |
T2094 |
det |
some,locations |
R1247 |
T2094 |
T2092 |
pobj |
locations,at |
R1248 |
T2095 |
T2088 |
cc |
and,for |
R1249 |
T2096 |
T2088 |
conj |
for,for |
R1251 |
T2097 |
T2096 |
pobj |
initiation,for |
R1252 |
T2098 |
T2097 |
prep |
of,initiation |
R1253 |
T2099 |
T2100 |
amod |
programmed,death |
R1254 |
T2100 |
T2098 |
pobj |
death,of |
R1256 |
T2101 |
T2100 |
compound |
cell,death |
R1257 |
T2102 |
T2100 |
prep |
in,death |
R1259 |
T2103 |
T2102 |
pobj |
webbing,in |
R1261 |
T2104 |
T2103 |
prep |
between,webbing |
R1262 |
T2105 |
T2104 |
pobj |
digits,between |
R1263 |
T2106 |
T2070 |
punct |
.,bypasses |
R1265 |
T2108 |
T2109 |
advmod |
Interestingly,is |
R1267 |
T2110 |
T2109 |
punct |
", ",is |
R1268 |
T2111 |
T2109 |
nsubj |
Bmpr1a,is |
R1269 |
T2112 |
T2109 |
advmod |
also,is |
R1271 |
T2113 |
T2109 |
acomp |
required,is |
R1272 |
T2114 |
T2109 |
prep |
for,is |
R1273 |
T2115 |
T2116 |
amod |
postnatal,maintenance |
R1274 |
T1953 |
T1954 |
det |
A,system |
R1275 |
T2116 |
T2114 |
pobj |
maintenance,for |
R1276 |
T2117 |
T2116 |
prep |
of,maintenance |
R1277 |
T2118 |
T2119 |
amod |
articular,cartilage |
R1278 |
T2119 |
T2117 |
pobj |
cartilage,of |
R1279 |
T1954 |
T1956 |
nsubj |
system,be |
R1280 |
T2120 |
T2109 |
prep |
throughout,is |
R1281 |
T1955 |
T1954 |
amod |
genetic,system |
R1282 |
T1957 |
T1954 |
prep |
for,system |
R1283 |
T2121 |
T2120 |
pobj |
most,throughout |
R1284 |
T1958 |
T1957 |
pcomp |
activating,for |
R1285 |
T2122 |
T2121 |
prep |
of,most |
R1286 |
T2123 |
T2124 |
det |
the,skeleton |
R1287 |
T2124 |
T2122 |
pobj |
skeleton,of |
R1288 |
T1959 |
T1958 |
cc |
or,activating |
R1289 |
T2125 |
T2109 |
punct |
.,is |
R1290 |
T2127 |
T2128 |
prep |
In,forms |
R1291 |
T1960 |
T1958 |
conj |
inactivating,activating |
R1292 |
T2129 |
T2130 |
nmod |
Gdf5,Cre |
R1293 |
T2130 |
T2132 |
nmod |
Cre,mice |
R1294 |
T2131 |
T2130 |
punct |
-,Cre |
R1295 |
T2132 |
T2127 |
pobj |
mice,In |
R1296 |
T2133 |
T2130 |
punct |
/,Cre |
R1297 |
T1961 |
T1960 |
dobj |
genes,inactivating |
R1298 |
T2134 |
T2130 |
appos |
Bmpr1afloxP,Cre |
R1299 |
T1962 |
T1963 |
advmod |
specifically,in |
R1300 |
T2135 |
T2128 |
punct |
", ",forms |
R1301 |
T2136 |
T2137 |
amod |
articular,cartilage |
R1302 |
T1963 |
T1960 |
prep |
in,inactivating |
R1303 |
T2137 |
T2128 |
nsubj |
cartilage,forms |
R1304 |
T2138 |
T2128 |
advmod |
initially,forms |
R1305 |
T2139 |
T2128 |
advmod |
normally,forms |
R1306 |
T1964 |
T1965 |
compound |
joint,tissues |
R1307 |
T2140 |
T2128 |
punct |
", ",forms |
R1308 |
T2141 |
T2128 |
cc |
but,forms |
R1309 |
T1965 |
T1963 |
pobj |
tissues,in |
R1310 |
T2142 |
T2143 |
advmod |
subsequently,loses |
R1311 |
T2143 |
T2128 |
conj |
loses,forms |
R1312 |
T1966 |
T1956 |
aux |
would,be |
R1313 |
T2144 |
T2143 |
dobj |
expression,loses |
R1314 |
T2145 |
T2144 |
prep |
of,expression |
R1315 |
T2146 |
T2147 |
amod |
several,markers |
R1316 |
T1967 |
T1968 |
advmod |
particularly,useful |
R1317 |
T2147 |
T2145 |
pobj |
markers,of |
R1318 |
T2148 |
T2147 |
amod |
key,markers |
R1319 |
T1968 |
T1956 |
acomp |
useful,be |
R1320 |
T2149 |
T2147 |
compound |
cartilage,markers |
R1321 |
T2150 |
T2143 |
prep |
after,loses |
R1322 |
T2151 |
T2150 |
pobj |
birth,after |
R1323 |
T1969 |
T1968 |
prep |
for,useful |
R1324 |
T2152 |
T2128 |
punct |
.,forms |
R1325 |
T1970 |
T1971 |
amod |
further,studies |
R1326 |
T2154 |
T2155 |
nsubj |
It,fibrillates |
R1327 |
T1971 |
T1969 |
pobj |
studies,for |
R1328 |
T2156 |
T2155 |
advmod |
ultimately,fibrillates |
R1329 |
T2157 |
T2155 |
cc |
and,fibrillates |
R1330 |
T2158 |
T2155 |
conj |
degenerates,fibrillates |
R1331 |
T1972 |
T1971 |
prep |
of,studies |
R1332 |
T2159 |
T2158 |
punct |
", ",degenerates |
R1333 |
T2160 |
T2158 |
conj |
resulting,degenerates |
R1334 |
T2161 |
T2160 |
prep |
in,resulting |
R1335 |
T1973 |
T1974 |
compound |
joint,formation |
R1336 |
T2162 |
T2163 |
amod |
severe,osteoarthritis |
R1337 |
T2163 |
T2161 |
pobj |
osteoarthritis,in |
R1338 |
T2164 |
T2163 |
cc |
and,osteoarthritis |
R1339 |
T1974 |
T1972 |
pobj |
formation,of |
R1340 |
T2165 |
T2163 |
conj |
loss,osteoarthritis |
R1341 |
T2166 |
T2165 |
prep |
of,loss |
R1342 |
T1975 |
T1974 |
cc |
and,formation |
R1343 |
T2167 |
T2166 |
pobj |
mobility,of |
R1344 |
T2168 |
T2155 |
punct |
.,fibrillates |
R1345 |
T2170 |
T2171 |
det |
These,experiments |
R1346 |
T2171 |
T2172 |
nsubj |
experiments,suggest |
R1347 |
T1976 |
T1974 |
conj |
maintenance,formation |
R1348 |
T2173 |
T2174 |
mark |
that,required |
R1349 |
T2174 |
T2172 |
advcl |
required,suggest |
R1350 |
T1977 |
T1956 |
punct |
.,be |
R1351 |
T2175 |
T2176 |
compound |
BMP,signaling |
R1352 |
T2176 |
T2174 |
nsubjpass |
signaling,required |
R1353 |
T2177 |
T2174 |
auxpass |
is,required |
R1354 |
T1979 |
T1980 |
advmod |
Here,take |
R1355 |
T2178 |
T2174 |
prep |
for,required |
R1356 |
T2179 |
T2180 |
amod |
normal,maintenance |
R1357 |
T2180 |
T2178 |
pobj |
maintenance,for |
R1358 |
T2181 |
T2180 |
prep |
of,maintenance |
R1359 |
T1981 |
T1980 |
nsubj |
we,take |
R1360 |
T2182 |
T2183 |
amod |
postnatal,cartilage |
R1361 |
T2183 |
T2181 |
pobj |
cartilage,of |
R1362 |
T2184 |
T2183 |
amod |
articular,cartilage |
R1363 |
T1982 |
T1980 |
dobj |
advantage,take |
R1364 |
T2185 |
T2174 |
punct |
", ",required |
R1365 |
T2186 |
T2174 |
cc |
and,required |
R1366 |
T2187 |
T2188 |
mark |
that,play |
R1367 |
T1983 |
T1980 |
prep |
of,take |
R1368 |
T2188 |
T2174 |
conj |
play,required |
R1369 |
T2189 |
T2188 |
nsubj |
modulation,play |
R1370 |
T1984 |
T1985 |
det |
the,pattern |
R1371 |
T2190 |
T2189 |
prep |
of,modulation |
R1372 |
T2191 |
T2192 |
det |
the,pathway |
R1373 |
T2192 |
T2190 |
pobj |
pathway,of |
R1374 |
T1985 |
T1983 |
pobj |
pattern,of |
R1375 |
T2193 |
T2194 |
compound |
BMP,signaling |
R1376 |
T2194 |
T2192 |
compound |
signaling,pathway |
R1377 |
T2195 |
T2188 |
aux |
may,play |
R1378 |
T1986 |
T1987 |
npadvmod |
tissue,specific |
R1379 |
T2196 |
T2197 |
det |
an,role |
R1380 |
T2197 |
T2188 |
dobj |
role,play |
R1381 |
T2198 |
T2197 |
amod |
important,role |
R1382 |
T2199 |
T2188 |
prep |
in,play |
R1383 |
T1987 |
T1985 |
amod |
specific,pattern |
R1384 |
T2200 |
T2201 |
compound |
joint,disease |
R1385 |
T2201 |
T2199 |
pobj |
disease,in |
R1386 |
T1988 |
T1987 |
punct |
-,specific |
R1387 |
T2202 |
T2172 |
punct |
.,suggest |
R1388 |
T1989 |
T1985 |
compound |
expression,pattern |
R1389 |
T1990 |
T1985 |
prep |
of,pattern |
R1390 |
T1991 |
T1992 |
det |
the,gene |
R1391 |
T1992 |
T1990 |
pobj |
gene,of |
R1392 |
T1993 |
T1992 |
compound |
Gdf5,gene |
R1393 |
T1994 |
T1995 |
aux |
to,engineer |
R1394 |
T1995 |
T1980 |
advcl |
engineer,take |
R1395 |
T1996 |
T1997 |
det |
a,system |
R1396 |
T1997 |
T1995 |
dobj |
system,engineer |
R1397 |
T1998 |
T1999 |
compound |
Cre,loxP |
R1398 |
T1999 |
T1997 |
compound |
loxP,system |
R1399 |
T2000 |
T1999 |
punct |
/,loxP |
R1400 |
T2001 |
T2002 |
punct |
(,Nagy |
R1401 |
T2002 |
T1997 |
meta |
Nagy,system |
R1402 |
T2003 |
T2002 |
nummod |
2000,Nagy |
R1403 |
T2004 |
T2002 |
punct |
),Nagy |
R1404 |
T2005 |
T1997 |
punct |
", ",system |
R1405 |
T2006 |
T2007 |
compound |
Gdf5,Cre |
R1406 |
T2007 |
T1997 |
appos |
Cre,system |
R1407 |
T2008 |
T2007 |
punct |
-,Cre |
R1408 |
T2009 |
T1997 |
punct |
", ",system |
R1409 |
T2010 |
T2011 |
dep |
that,used |
R1410 |
T2011 |
T1997 |
relcl |
used,system |
R1411 |
T2012 |
T2011 |
aux |
can,used |
R1412 |
T2013 |
T2011 |
auxpass |
be,used |