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PMC:3317373 / 11672-12230 JSONTXT

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pmc-enju-pas

Id Subject Object Predicate Lexical cue
T4942 436-441 WDT denotes which
T4941 434-435 -COMMA- denotes ,
T4940 433-434 -RRB- denotes )
T4939 429-433 CD denotes 0.01
T4938 427-428 SYM denotes <
T4937 425-426 NN denotes P
T4936 424-425 -LRB- denotes (
T4935 422-423 -RRB- denotes )
T4934 417-422 CD denotes 0.098
T4933 415-416 CC denotes ±
T4932 410-414 CD denotes 0.98
T4931 409-410 -LRB- denotes (
T4930 405-408 NN denotes TBI
T4929 402-404 NN denotes WT
T4928 396-401 NN denotes group
T4927 393-395 IN denotes of
T4926 388-392 DT denotes that
T4925 383-387 IN denotes than
T4924 381-382 -RRB- denotes )
T4923 376-381 CD denotes 0.064
T4922 374-375 CC denotes ±
T4921 369-373 CD denotes 0.38
T4920 368-369 -LRB- denotes (
T4919 364-367 NN denotes TBI
T4918 361-363 NN denotes KO
T4917 355-360 NN denotes group
T4916 352-354 IN denotes of
T4915 346-351 NN denotes value
T4914 343-345 NN denotes OD
T4913 337-342 JJ denotes lower
T4912 330-336 VB denotes showed
T4911 325-329 RB denotes also
T4910 319-324 NN denotes assay
T4909 313-318 NN denotes CCK-8
T4908 308-311 NNP denotes TBI
T4907 305-307 NNP denotes KO
T4906 299-304 NN denotes group
T4905 296-298 IN denotes in
T4904 291-295 NN denotes line
T4903 283-290 NN denotes scratch
T4902 280-282 IN denotes of
T4901 274-279 NN denotes sides
T4900 269-273 DT denotes both
T4899 266-268 IN denotes in
T4898 257-265 VB denotes observed
T4897 250-256 RB denotes mostly
T4896 245-249 VB denotes were
T4895 234-244 NN denotes distortion
T4894 226-233 NN denotes process
T4893 214-225 JJ denotes cytoplasmic
T4892 210-213 CC denotes and
T4891 208-209 -COMMA- denotes ,
T4890 204-208 NN denotes lost
T4889 199-203 NN denotes cell
T4888 197-198 -COMMA- denotes ,
T4887 192-197 NN denotes plate
T4886 184-191 NN denotes culture
T4885 179-183 IN denotes from
T4884 168-178 NN denotes detachment
T4883 164-167 DT denotes The
T4882 155-162 NN denotes scratch
T4881 149-154 IN denotes after
T4880 147-148 NN denotes h
T4879 144-146 CD denotes 24
T4878 141-143 IN denotes at
T4877 132-140 NN denotes analysis
T4876 126-131 NN denotes CCK-8
T4875 122-125 CC denotes and
T4874 111-121 NN denotes microscope
T4873 108-110 IN denotes by
T4872 99-107 VB denotes detected
T4871 95-98 VB denotes was
T4870 89-94 NN denotes death
T4869 84-88 NN denotes Cell
T4868 77-83 NNP denotes Injury
T4867 69-76 NNP denotes Scratch
T4866 63-68 IN denotes after
T4865 52-62 NN denotes Astrocytes
T4864 49-51 IN denotes in
T4863 43-48 NNP denotes Death
T4862 38-42 NNP denotes Cell
T4861 34-37 DT denotes the
T4860 23-33 VB denotes Aggravated
T4859 18-22 NN denotes Nrf2
T4858 15-17 IN denotes of
T4857 5-14 NN denotes Depletion
T4856 0-3 CD denotes 3.1
T4963 551-557 NN denotes groups
T4962 537-550 JJ denotes sham-operated
T4961 533-536 CD denotes two
T4960 525-532 IN denotes between
T4959 514-524 NN denotes difference
T4958 511-513 DT denotes no
T4957 507-510 VB denotes was
T4956 501-506 EX denotes There
T4955 498-499 -RRB- denotes )
T4954 497-498 CD denotes 1
T4953 490-496 NN denotes Figure
T4952 489-490 -LRB- denotes (
T4951 485-488 NN denotes TBI
T4950 482-484 NN denotes KO
T4949 476-481 NN denotes group
T4948 473-475 IN denotes in
T4947 467-472 NN denotes death
T4946 462-466 NN denotes cell
T4945 457-461 JJ denotes more
T4944 447-456 VB denotes suggested
T4943 442-446 RB denotes also
R4123 T4857 T4858 arg1Of Depletion,of
R4124 T4857 T4860 arg1Of Depletion,Aggravated
R4125 T4859 T4858 arg2Of Nrf2,of
R4126 T4860 T4866 arg1Of Aggravated,after
R4127 T4863 T4860 arg2Of Death,Aggravated
R4128 T4863 T4861 arg1Of Death,the
R4129 T4863 T4862 arg1Of Death,Cell
R4130 T4863 T4864 arg1Of Death,in
R4131 T4865 T4864 arg2Of Astrocytes,in
R4132 T4868 T4866 arg2Of Injury,after
R4133 T4868 T4867 arg1Of Injury,Scratch
R4134 T4870 T4869 arg1Of death,Cell
R4135 T4870 T4871 arg1Of death,was
R4136 T4870 T4872 arg2Of death,detected
R4137 T4872 T4871 arg2Of detected,was
R4138 T4874 T4875 arg1Of microscope,and
R4139 T4876 T4875 arg2Of CCK-8,and
R4140 T4877 T4872 arg1Of analysis,detected
R4141 T4877 T4873 arg2Of analysis,by
R4142 T4877 T4874 arg1Of analysis,microscope
R4143 T4877 T4876 arg1Of analysis,CCK-8
R4144 T4877 T4878 arg1Of analysis,at
R4145 T4880 T4878 arg2Of h,at
R4146 T4880 T4879 arg1Of h,24
R4147 T4880 T4881 arg1Of h,after
R4148 T4882 T4881 arg2Of scratch,after
R4149 T4884 T4883 arg1Of detachment,The
R4150 T4884 T4885 arg1Of detachment,from
R4151 T4884 T4896 arg1Of detachment,were
R4152 T4884 T4898 arg2Of detachment,observed
R4153 T4887 T4886 arg1Of plate,culture
R4154 T4887 T4888 arg1Of plate,","
R4155 T4888 T4892 arg1Of ",",and
R4156 T4890 T4888 arg2Of lost,","
R4157 T4890 T4889 arg1Of lost,cell
R4158 T4892 T4885 arg2Of and,from
R4159 T4892 T4891 arg1Of and,","
R4160 T4895 T4892 arg2Of distortion,and
R4161 T4895 T4893 arg1Of distortion,cytoplasmic
R4162 T4895 T4894 arg1Of distortion,process
R4163 T4898 T4896 arg2Of observed,were
R4164 T4898 T4897 arg1Of observed,mostly
R4165 T4898 T4899 arg1Of observed,in
R4166 T4901 T4899 arg2Of sides,in
R4167 T4901 T4900 arg1Of sides,both
R4168 T4901 T4902 arg1Of sides,of
R4169 T4901 T4905 arg1Of sides,in
R4170 T4904 T4902 arg2Of line,of
R4171 T4904 T4903 arg1Of line,scratch
R4172 T4908 T4905 arg2Of TBI,in
R4173 T4908 T4906 arg1Of TBI,group
R4174 T4908 T4907 arg1Of TBI,KO
R4175 T4910 T4909 arg1Of assay,CCK-8
R4176 T4910 T4912 arg1Of assay,showed
R4177 T4912 T4911 arg1Of showed,also
R4178 T4915 T4912 arg2Of value,showed
R4179 T4915 T4913 arg1Of value,lower
R4180 T4915 T4914 arg1Of value,OD
R4181 T4915 T4916 arg1Of value,of
R4182 T4919 T4916 arg2Of TBI,of
R4183 T4919 T4917 arg1Of TBI,group
R4184 T4919 T4918 arg1Of TBI,KO
R4185 T4919 T4920 arg1Of TBI,(
R4186 T4919 T4925 arg1Of TBI,than
R4187 T4921 T4922 arg1Of 0.38,±
R4188 T4922 T4920 arg2Of ±,(
R4189 T4923 T4922 arg2Of 0.064,±
R4190 T4924 T4920 arg3Of ),(
R4191 T4926 T4925 arg2Of that,than
R4192 T4926 T4927 arg1Of that,of
R4193 T4930 T4927 arg2Of TBI,of
R4194 T4930 T4928 arg1Of TBI,group
R4195 T4930 T4929 arg1Of TBI,WT
R4196 T4930 T4931 arg1Of TBI,(
R4197 T4930 T4936 arg1Of TBI,(
R4198 T4930 T4941 arg1Of TBI,","
R4199 T4930 T4942 arg1Of TBI,which
R4200 T4930 T4944 arg1Of TBI,suggested
R4201 T4932 T4933 arg1Of 0.98,±
R4202 T4933 T4931 arg2Of ±,(
R4203 T4934 T4933 arg2Of 0.098,±
R4204 T4935 T4931 arg3Of ),(
R4205 T4937 T4936 arg2Of P,(
R4206 T4937 T4938 arg1Of P,<
R4207 T4937 T4939 arg1Of P,0.01
R4208 T4940 T4936 arg3Of ),(
R4209 T4944 T4943 arg1Of suggested,also
R4210 T4947 T4944 arg2Of death,suggested
R4211 T4947 T4945 arg1Of death,more
R4212 T4947 T4946 arg1Of death,cell
R4213 T4947 T4948 arg1Of death,in
R4214 T4951 T4948 arg2Of TBI,in
R4215 T4951 T4949 arg1Of TBI,group
R4216 T4951 T4950 arg1Of TBI,KO
R4217 T4951 T4952 arg1Of TBI,(
R4218 T4953 T4952 arg2Of Figure,(
R4219 T4953 T4954 arg1Of Figure,1
R4220 T4955 T4952 arg3Of ),(
R4221 T4956 T4957 arg1Of There,was
R4222 T4959 T4957 arg2Of difference,was
R4223 T4959 T4958 arg1Of difference,no
R4224 T4959 T4960 arg1Of difference,between
R4225 T4963 T4960 arg2Of groups,between
R4226 T4963 T4961 arg1Of groups,two
R4227 T4963 T4962 arg1Of groups,sham-operated

bionlp-st-ge-2016-uniprot

Id Subject Object Predicate Lexical cue
T4972 0-3 http://www.uniprot.org/uniprot/Q8IX06 denotes 3.1
T4971 0-3 http://www.uniprot.org/uniprot/Q5D1E8 denotes 3.1
T4970 0-3 http://www.uniprot.org/uniprot/Q5D1E7 denotes 3.1

GO-BP

Id Subject Object Predicate Lexical cue
T4855 467-472 http://purl.obolibrary.org/obo/GO_0016265 denotes death
T4854 89-94 http://purl.obolibrary.org/obo/GO_0016265 denotes death
T4853 43-48 http://purl.obolibrary.org/obo/GO_0016265 denotes Death
T4852 462-472 http://purl.obolibrary.org/obo/GO_0008219 denotes cell death
T4851 84-94 http://purl.obolibrary.org/obo/GO_0008219 denotes Cell death
T4850 38-48 http://purl.obolibrary.org/obo/GO_0008219 denotes Cell Death

GO-CC

Id Subject Object Predicate Lexical cue
T4969 192-203 http://purl.obolibrary.org/obo/GO_0009504 denotes plate, cell
T4968 462-466 http://purl.obolibrary.org/obo/GO_0005623 denotes cell
T4967 199-203 http://purl.obolibrary.org/obo/GO_0005623 denotes cell
T4966 84-88 http://purl.obolibrary.org/obo/GO_0005623 denotes Cell
T4965 38-42 http://purl.obolibrary.org/obo/GO_0005623 denotes Cell

sentences

Id Subject Object Predicate Lexical cue
T4849 501-558 Sentence denotes There was no difference between two sham-operated groups.
T4848 313-500 Sentence denotes CCK-8 assay also showed lower OD value of group KO TBI (0.38 ± 0.064) than that of group WT TBI (0.98 ± 0.098) (P < 0.01), which also suggested more cell death in group KO TBI (Figure 1).
T4847 164-312 Sentence denotes The detachment from culture plate, cell lost, and cytoplasmic process distortion were mostly observed in both sides of scratch line in group KO TBI.
T4846 84-163 Sentence denotes Cell death was detected by microscope and CCK-8 analysis at 24 h after scratch.
T4845 5-83 Sentence denotes Depletion of Nrf2 Aggravated the Cell Death in Astrocytes after Scratch Injury
T4844 0-4 Sentence denotes 3.1.
T124 0-4 Sentence denotes 3.1.
T125 5-83 Sentence denotes Depletion of Nrf2 Aggravated the Cell Death in Astrocytes after Scratch Injury
T126 84-163 Sentence denotes Cell death was detected by microscope and CCK-8 analysis at 24 h after scratch.
T127 164-312 Sentence denotes The detachment from culture plate, cell lost, and cytoplasmic process distortion were mostly observed in both sides of scratch line in group KO TBI.
T128 313-500 Sentence denotes CCK-8 assay also showed lower OD value of group KO TBI (0.38 ± 0.064) than that of group WT TBI (0.98 ± 0.098) (P < 0.01), which also suggested more cell death in group KO TBI (Figure 1).
T129 501-558 Sentence denotes There was no difference between two sham-operated groups.

ICD10

Id Subject Object Predicate Lexical cue
T4964 77-83 http://purl.bioontology.org/ontology/ICD10/T14.9 denotes Injury

bionlp-st-ge-2016-test-ihmc

Id Subject Object Predicate Lexical cue
T5101 34-83 Entity denotes the Cell Death in Astrocytes after Scratch Injury
T5100 52-62 Entity denotes Astrocytes
T5099 476-499 Protein denotes group KO TBI (Figure 1)
T5098 147-162 Entity denotes h after scratch
T5097 305-307 Protein denotes KO
T5096 199-208 Entity denotes cell lost
T5095 361-363 Protein denotes KO
T5094 126-131 Protein denotes CCK-8
T5093 38-42 Entity denotes Cell
T5092 537-550 Entity denotes sham-operated
T5091 15-22 Protein denotes of Nrf2
T5090 313-318 Protein denotes CCK-8

bionlp-st-ge-2016-spacy-parsed

Id Subject Object Predicate Lexical cue
T5087 557-558 . denotes .
T5086 551-557 NNS denotes groups
T5085 537-550 JJ denotes sham-operated
T5084 533-536 CD denotes two
T5083 525-532 IN denotes between
T5082 514-524 NN denotes difference
T5081 511-513 DT denotes no
T5080 507-510 VBD denotes was
T5079 501-506 EX denotes There
T5078 499-500 . denotes .
T5077 498-499 -RRB- denotes )
T5076 497-498 CD denotes 1
T5075 490-496 NNP denotes Figure
T5074 489-490 -LRB- denotes (
T5073 485-488 NNP denotes TBI
T5072 482-484 NNP denotes KO
T5071 476-481 NN denotes group
T5070 473-475 IN denotes in
T5069 467-472 NN denotes death
T5068 462-466 NN denotes cell
T5067 457-461 JJR denotes more
T5066 447-456 VBD denotes suggested
T5065 442-446 RB denotes also
T5062 436-441 WDT denotes which
T5025 299-304 NN denotes group
T5024 296-298 IN denotes in
T5023 291-295 NN denotes line
T5022 283-290 NN denotes scratch
T5021 280-282 IN denotes of
T5020 274-279 NNS denotes sides
T5019 269-273 DT denotes both
T5018 266-268 IN denotes in
T5017 257-265 VBN denotes observed
T5016 250-256 RB denotes mostly
T5015 245-249 VBD denotes were
T5014 234-244 NN denotes distortion
T5013 226-233 NN denotes process
T5012 214-225 JJ denotes cytoplasmic
T5011 210-213 CC denotes and
T5010 208-209 , denotes ,
T5009 204-208 VBD denotes lost
T5008 199-203 NN denotes cell
T5007 197-198 , denotes ,
T5006 192-197 NN denotes plate
T5005 184-191 NN denotes culture
T5004 179-183 IN denotes from
T5003 168-178 NN denotes detachment
T5002 164-167 DT denotes The
T5001 162-163 . denotes .
T5000 155-162 NN denotes scratch
T4999 149-154 IN denotes after
T4998 147-148 NN denotes h
T4997 144-146 CD denotes 24
T4980 38-42 NNP denotes Cell
T4979 34-37 DT denotes the
T4978 23-33 NNP denotes Aggravated
T4977 18-22 NNP denotes Nrf2
T4976 15-17 IN denotes of
T4975 5-14 NN denotes Depletion
T4974 3-4 . denotes .
T4973 0-3 CD denotes 3.1
T4996 141-143 IN denotes at
T4995 132-140 NN denotes analysis
T4994 126-131 NN denotes CCK-8
T4993 122-125 CC denotes and
T4992 111-121 NN denotes microscope
T4991 108-110 IN denotes by
T4990 99-107 VBN denotes detected
T4989 95-98 VBD denotes was
T4988 89-94 NN denotes death
T4987 84-88 NNP denotes Cell
T4986 77-83 NNP denotes Injury
T4985 69-76 NNP denotes Scratch
T4984 63-68 IN denotes after
T4983 52-62 NNP denotes Astrocytes
T4982 49-51 IN denotes in
T4981 43-48 NNP denotes Death
T5027 308-311 NNP denotes TBI
T5026 305-307 NNP denotes KO
T5061 434-435 , denotes ,
T5060 433-434 -RRB- denotes )
T5059 429-433 CD denotes 0.01
T5058 427-428 CD denotes <
T5057 425-426 NNP denotes P
T5056 424-425 -LRB- denotes (
T5055 422-423 -RRB- denotes )
T5054 417-422 CD denotes 0.098
T5053 415-416 CD denotes ±
T5052 410-414 CD denotes 0.98
T5051 409-410 -LRB- denotes (
T5050 405-408 NNP denotes TBI
T5049 402-404 NNP denotes WT
T5048 396-401 NN denotes group
T5047 393-395 IN denotes of
T5046 388-392 DT denotes that
T5045 383-387 IN denotes than
T5044 381-382 -RRB- denotes )
T5043 376-381 CD denotes 0.064
T5042 374-375 CD denotes ±
T5041 369-373 CD denotes 0.38
T5040 368-369 -LRB- denotes (
T5039 364-367 NNP denotes TBI
T5038 361-363 NNP denotes KO
T5037 355-360 NN denotes group
T5036 352-354 IN denotes of
T5035 346-351 NN denotes value
T5034 343-345 NNP denotes OD
T5033 337-342 JJR denotes lower
T5032 330-336 VBD denotes showed
T5031 325-329 RB denotes also
T5030 319-324 NN denotes assay
T5029 313-318 CD denotes CCK-8
T5028 311-312 . denotes .
R4267 T5012 T5014 amod cytoplasmic,distortion
R4268 T5013 T5014 compound process,distortion
R4269 T5014 T5017 nsubjpass distortion,observed
R4270 T5015 T5017 auxpass were,observed
R4271 T5016 T5017 advmod mostly,observed
R4272 T5017 T5009 conj observed,lost
R4273 T5018 T5017 prep in,observed
R4274 T5019 T5020 det both,sides
R4275 T5020 T5018 pobj sides,in
R4276 T5021 T5020 prep of,sides
R4277 T5022 T5023 compound scratch,line
R4278 T5023 T5021 pobj line,of
R4279 T5024 T5023 prep in,line
R4280 T5025 T5027 compound group,TBI
R4281 T5026 T5027 compound KO,TBI
R4282 T5027 T5024 pobj TBI,in
R4283 T5028 T5017 punct .,observed
R4284 T5029 T5030 nummod CCK-8,assay
R4285 T5030 T5032 nsubj assay,showed
R4286 T5031 T5032 advmod also,showed
R4287 T5032 T5032 ROOT showed,showed
R4288 T5033 T5035 amod lower,value
R4289 T5034 T5035 compound OD,value
R4290 T5035 T5032 dobj value,showed
R4291 T5036 T5035 prep of,value
R4292 T5037 T5039 compound group,TBI
R4293 T5038 T5039 compound KO,TBI
R4294 T5039 T5036 pobj TBI,of
R4295 T5040 T5041 punct (,0.38
R4296 T5041 T5043 nummod 0.38,0.064
R4297 T5042 T5043 nummod ±,0.064
R4298 T5043 T5039 appos 0.064,TBI
R4299 T5044 T5032 punct ),showed
R4300 T5045 T5032 prep than,showed
R4301 T5046 T5045 pobj that,than
R4302 T5047 T5046 prep of,that
R4303 T5048 T5047 pobj group,of
R4304 T5049 T5050 compound WT,TBI
R4305 T5050 T5048 appos TBI,group
R4306 T5051 T5052 punct (,0.98
R4307 T5052 T5054 nummod 0.98,0.098
R4308 T5053 T5054 nummod ±,0.098
R4309 T5054 T5048 appos 0.098,group
R4310 T5055 T5032 punct ),showed
R4311 T5056 T5032 punct (,showed
R4312 T5057 T5057 ROOT P,P
R4313 T5058 T5057 nummod <,P
R4314 T5059 T5057 nummod 0.01,P
R4315 T5060 T5057 punct ),P
R4316 T5061 T5057 punct ",",P
R4317 T5062 T5066 nsubj which,suggested
R4319 T5065 T5066 advmod also,suggested
R4320 T5066 T5057 relcl suggested,P
R4321 T5067 T5069 amod more,death
R4322 T5068 T5069 compound cell,death
R4323 T5069 T5066 dobj death,suggested
R4324 T5070 T5069 prep in,death
R4325 T5071 T5073 compound group,TBI
R4326 T5072 T5073 compound KO,TBI
R4327 T5073 T5070 pobj TBI,in
R4328 T5074 T5075 punct (,Figure
R4329 T5075 T5073 appos Figure,TBI
R4330 T5076 T5075 nummod 1,Figure
R4331 T5077 T5073 punct ),TBI
R4332 T5078 T5057 punct .,P
R4333 T5079 T5080 expl There,was
R4334 T5080 T5080 ROOT was,was
R4335 T5081 T5082 det no,difference
R4336 T5082 T5080 attr difference,was
R4337 T5083 T5082 prep between,difference
R4338 T5084 T5086 nummod two,groups
R4339 T5085 T5086 amod sham-operated,groups
R4340 T5086 T5083 pobj groups,between
R4341 T5087 T5080 punct .,was
R4228 T4973 T4973 ROOT 3.1,3.1
R4229 T4974 T4973 punct .,3.1
R4230 T4975 T4990 nsubjpass Depletion,detected
R4231 T4976 T4975 prep of,Depletion
R4232 T4977 T4978 compound Nrf2,Aggravated
R4233 T4978 T4976 pobj Aggravated,of
R4234 T4979 T4981 det the,Death
R4235 T4980 T4981 compound Cell,Death
R4236 T4981 T4978 dobj Death,Aggravated
R4237 T4982 T4981 prep in,Death
R4238 T4983 T4982 pobj Astrocytes,in
R4239 T4984 T4990 mark after,detected
R4240 T4985 T4988 compound Scratch,death
R4241 T4986 T4987 compound Injury,Cell
R4242 T4987 T4988 compound Cell,death
R4243 T4988 T4990 nsubjpass death,detected
R4244 T4989 T4990 auxpass was,detected
R4245 T4990 T4990 ROOT detected,detected
R4246 T4991 T4990 agent by,detected
R4247 T4992 T4991 pobj microscope,by
R4248 T4993 T4992 cc and,microscope
R4249 T4994 T4995 compound CCK-8,analysis
R4250 T4995 T4992 conj analysis,microscope
R4251 T4996 T4995 prep at,analysis
R4252 T4997 T4998 nummod 24,h
R4253 T4998 T4996 pobj h,at
R4254 T4999 T4990 prep after,detected
R4255 T5000 T4999 pobj scratch,after
R4256 T5001 T4990 punct .,detected
R4257 T5002 T5003 det The,detachment
R4258 T5003 T5009 nsubj detachment,lost
R4259 T5004 T5003 prep from,detachment
R4260 T5005 T5006 compound culture,plate
R4261 T5006 T5004 pobj plate,from
R4262 T5007 T5009 punct ",",lost
R4263 T5008 T5009 nsubj cell,lost
R4264 T5009 T5009 ROOT lost,lost
R4265 T5010 T5009 punct ",",lost
R4266 T5011 T5009 cc and,lost

bionlp-st-ge-2016-test-tees

Id Subject Object Predicate Lexical cue
T5064 5-14 Negative_regulation denotes Depletion
T5063 18-22 Protein denotes Nrf2
R4318 T5063 T5064 themeOf Nrf2,Depletion

testone

Id Subject Object Predicate Lexical cue
T4843 5-14 Negative_regulation denotes Depletion