| Id |
Subject |
Object |
Predicate |
Lexical cue |
| T181 |
0-10 |
sentence |
denotes |
Background |
| T182 |
0-10 |
NN |
denotes |
Background |
| T183 |
10-125 |
sentence |
denotes |
Tripeptidyl-peptidase I, also known as CLN2, is a member of the family of sedolisins (serine-carboxyl peptidases). |
| T184 |
11-22 |
NN |
denotes |
Tripeptidyl |
| T185 |
23-32 |
NN |
denotes |
peptidase |
| T186 |
22-23 |
HYPH |
denotes |
- |
| T187 |
56-58 |
VBZ |
denotes |
is |
| T188 |
33-34 |
CD |
denotes |
I |
| T189 |
34-36 |
, |
denotes |
, |
| T190 |
36-40 |
RB |
denotes |
also |
| T191 |
41-46 |
VBN |
denotes |
known |
| T192 |
47-49 |
IN |
denotes |
as |
| T193 |
50-54 |
NN |
denotes |
CLN2 |
| T194 |
54-56 |
, |
denotes |
, |
| T195 |
59-60 |
DT |
denotes |
a |
| T196 |
61-67 |
NN |
denotes |
member |
| T197 |
68-70 |
IN |
denotes |
of |
| T198 |
71-74 |
DT |
denotes |
the |
| T199 |
75-81 |
NN |
denotes |
family |
| T200 |
82-84 |
IN |
denotes |
of |
| T201 |
85-95 |
NNS |
denotes |
sedolisins |
| T202 |
96-97 |
-LRB- |
denotes |
( |
| T203 |
97-103 |
NN |
denotes |
serine |
| T204 |
104-112 |
NN |
denotes |
carboxyl |
| T205 |
103-104 |
HYPH |
denotes |
- |
| T206 |
113-123 |
NNS |
denotes |
peptidases |
| T207 |
123-124 |
-RRB- |
denotes |
) |
| T208 |
124-125 |
. |
denotes |
. |
| T209 |
125-273 |
sentence |
denotes |
In humans, defects in expression of this enzyme lead to a fatal neurodegenerative disease, classical late-infantile neuronal ceroid lipofuscinosis. |
| T210 |
126-128 |
IN |
denotes |
In |
| T211 |
174-178 |
VBP |
denotes |
lead |
| T212 |
129-135 |
NNS |
denotes |
humans |
| T213 |
135-137 |
, |
denotes |
, |
| T214 |
137-144 |
VBZ |
denotes |
defects |
| T215 |
145-147 |
IN |
denotes |
in |
| T216 |
148-158 |
NN |
denotes |
expression |
| T217 |
159-161 |
IN |
denotes |
of |
| T218 |
162-166 |
DT |
denotes |
this |
| T219 |
167-173 |
NN |
denotes |
enzyme |
| T220 |
179-181 |
IN |
denotes |
to |
| T221 |
182-183 |
DT |
denotes |
a |
| T222 |
208-215 |
NN |
denotes |
disease |
| T223 |
184-189 |
JJ |
denotes |
fatal |
| T224 |
190-207 |
JJ |
denotes |
neurodegenerative |
| T225 |
215-217 |
, |
denotes |
, |
| T226 |
217-226 |
JJ |
denotes |
classical |
| T227 |
258-272 |
NN |
denotes |
lipofuscinosis |
| T228 |
227-231 |
JJ |
denotes |
late |
| T229 |
232-241 |
JJ |
denotes |
infantile |
| T230 |
231-232 |
HYPH |
denotes |
- |
| T231 |
242-250 |
JJ |
denotes |
neuronal |
| T232 |
251-257 |
JJ |
denotes |
ceroid |
| T233 |
272-273 |
. |
denotes |
. |
| T234 |
273-469 |
sentence |
denotes |
Similar enzymes have been found in the genomic sequences of several species, but neither systematic analyses of their distribution nor modeling of their structures have been previously attempted. |
| T235 |
274-281 |
JJ |
denotes |
Similar |
| T236 |
282-289 |
NNS |
denotes |
enzymes |
| T237 |
300-305 |
VBN |
denotes |
found |
| T238 |
290-294 |
VBP |
denotes |
have |
| T239 |
295-299 |
VBN |
denotes |
been |
| T240 |
306-308 |
IN |
denotes |
in |
| T241 |
309-312 |
DT |
denotes |
the |
| T242 |
321-330 |
NNS |
denotes |
sequences |
| T243 |
313-320 |
JJ |
denotes |
genomic |
| T244 |
331-333 |
IN |
denotes |
of |
| T245 |
334-341 |
JJ |
denotes |
several |
| T246 |
342-349 |
NNS |
denotes |
species |
| T247 |
349-351 |
, |
denotes |
, |
| T248 |
351-354 |
CC |
denotes |
but |
| T249 |
355-362 |
CC |
denotes |
neither |
| T250 |
374-382 |
NNS |
denotes |
analyses |
| T251 |
363-373 |
JJ |
denotes |
systematic |
| T252 |
459-468 |
VBN |
denotes |
attempted |
| T253 |
383-385 |
IN |
denotes |
of |
| T254 |
386-391 |
PRP$ |
denotes |
their |
| T255 |
392-404 |
NN |
denotes |
distribution |
| T256 |
405-408 |
CC |
denotes |
nor |
| T257 |
409-417 |
NN |
denotes |
modeling |
| T258 |
418-420 |
IN |
denotes |
of |
| T259 |
421-426 |
PRP$ |
denotes |
their |
| T260 |
427-437 |
NNS |
denotes |
structures |
| T261 |
438-442 |
VBP |
denotes |
have |
| T262 |
443-447 |
VBN |
denotes |
been |
| T263 |
448-458 |
RB |
denotes |
previously |
| T264 |
468-469 |
. |
denotes |
. |
| T265 |
469-478 |
sentence |
denotes |
Results |
| T266 |
471-478 |
NNS |
denotes |
Results |
| T267 |
478-595 |
sentence |
denotes |
We have analyzed the presence of orthologs of human CLN2 in the genomic sequences of a number of eukaryotic species. |
| T268 |
479-481 |
PRP |
denotes |
We |
| T269 |
487-495 |
VBN |
denotes |
analyzed |
| T270 |
482-486 |
VBP |
denotes |
have |
| T271 |
496-499 |
DT |
denotes |
the |
| T272 |
500-508 |
NN |
denotes |
presence |
| T273 |
509-511 |
IN |
denotes |
of |
| T274 |
512-521 |
NNS |
denotes |
orthologs |
| T275 |
522-524 |
IN |
denotes |
of |
| T276 |
525-530 |
JJ |
denotes |
human |
| T277 |
531-535 |
NN |
denotes |
CLN2 |
| T278 |
536-538 |
IN |
denotes |
in |
| T279 |
539-542 |
DT |
denotes |
the |
| T280 |
551-560 |
NNS |
denotes |
sequences |
| T281 |
543-550 |
JJ |
denotes |
genomic |
| T282 |
561-563 |
IN |
denotes |
of |
| T283 |
564-565 |
DT |
denotes |
a |
| T284 |
566-572 |
NN |
denotes |
number |
| T285 |
573-575 |
IN |
denotes |
of |
| T286 |
576-586 |
JJ |
denotes |
eukaryotic |
| T287 |
587-594 |
NNS |
denotes |
species |
| T288 |
594-595 |
. |
denotes |
. |
| T289 |
595-713 |
sentence |
denotes |
Enzymes with sequences sharing over 80% identity have been found in the genomes of macaque, mouse, rat, dog, and cow. |
| T290 |
596-603 |
NNS |
denotes |
Enzymes |
| T291 |
655-660 |
VBN |
denotes |
found |
| T292 |
604-608 |
IN |
denotes |
with |
| T293 |
609-618 |
NNS |
denotes |
sequences |
| T294 |
619-626 |
VBG |
denotes |
sharing |
| T295 |
627-631 |
IN |
denotes |
over |
| T296 |
632-634 |
CD |
denotes |
80 |
| T297 |
634-635 |
NN |
denotes |
% |
| T298 |
636-644 |
NN |
denotes |
identity |
| T299 |
645-649 |
VBP |
denotes |
have |
| T300 |
650-654 |
VBN |
denotes |
been |
| T301 |
661-663 |
IN |
denotes |
in |
| T302 |
664-667 |
DT |
denotes |
the |
| T303 |
668-675 |
NNS |
denotes |
genomes |
| T304 |
676-678 |
IN |
denotes |
of |
| T305 |
679-686 |
NN |
denotes |
macaque |
| T306 |
686-688 |
, |
denotes |
, |
| T307 |
688-693 |
NN |
denotes |
mouse |
| T308 |
693-695 |
, |
denotes |
, |
| T309 |
695-698 |
NN |
denotes |
rat |
| T310 |
698-700 |
, |
denotes |
, |
| T311 |
700-703 |
NN |
denotes |
dog |
| T312 |
703-705 |
, |
denotes |
, |
| T313 |
705-708 |
CC |
denotes |
and |
| T314 |
709-712 |
NN |
denotes |
cow |
| T315 |
712-713 |
. |
denotes |
. |
| T316 |
713-845 |
sentence |
denotes |
Closely related, although clearly distinct, enzymes are present in fish (fugu and zebra), as well as in frogs (Xenopus tropicalis). |
| T317 |
714-721 |
RB |
denotes |
Closely |
| T318 |
722-729 |
VBN |
denotes |
related |
| T319 |
758-765 |
NNS |
denotes |
enzymes |
| T320 |
729-731 |
, |
denotes |
, |
| T321 |
748-756 |
JJ |
denotes |
distinct |
| T322 |
731-739 |
IN |
denotes |
although |
| T323 |
740-747 |
RB |
denotes |
clearly |
| T324 |
756-758 |
, |
denotes |
, |
| T325 |
766-769 |
VBP |
denotes |
are |
| T326 |
770-777 |
JJ |
denotes |
present |
| T327 |
778-780 |
IN |
denotes |
in |
| T328 |
781-785 |
NN |
denotes |
fish |
| T329 |
786-787 |
-LRB- |
denotes |
( |
| T330 |
787-791 |
NN |
denotes |
fugu |
| T331 |
792-795 |
CC |
denotes |
and |
| T332 |
796-801 |
NN |
denotes |
zebra |
| T333 |
801-802 |
-RRB- |
denotes |
) |
| T334 |
802-804 |
, |
denotes |
, |
| T335 |
804-806 |
RB |
denotes |
as |
| T336 |
812-814 |
IN |
denotes |
as |
| T337 |
807-811 |
RB |
denotes |
well |
| T338 |
815-817 |
IN |
denotes |
in |
| T339 |
818-823 |
NNS |
denotes |
frogs |
| T340 |
824-825 |
-LRB- |
denotes |
( |
| T341 |
825-832 |
NNP |
denotes |
Xenopus |
| T342 |
833-843 |
NNP |
denotes |
tropicalis |
| T343 |
843-844 |
-RRB- |
denotes |
) |
| T344 |
844-845 |
. |
denotes |
. |
| T345 |
845-945 |
sentence |
denotes |
A three-dimensional model of human CLN2 was built based mainly on the homology with Pseudomonas sp. |
| T346 |
846-847 |
DT |
denotes |
A |
| T347 |
866-871 |
NN |
denotes |
model |
| T348 |
848-853 |
CD |
denotes |
three |
| T349 |
854-865 |
JJ |
denotes |
dimensional |
| T350 |
853-854 |
HYPH |
denotes |
- |
| T351 |
890-895 |
VBN |
denotes |
built |
| T352 |
872-874 |
IN |
denotes |
of |
| T353 |
875-880 |
JJ |
denotes |
human |
| T354 |
881-885 |
NN |
denotes |
CLN2 |
| T355 |
886-889 |
VBD |
denotes |
was |
| T356 |
896-901 |
VBN |
denotes |
based |
| T357 |
902-908 |
RB |
denotes |
mainly |
| T358 |
909-911 |
IN |
denotes |
on |
| T359 |
912-915 |
DT |
denotes |
the |
| T360 |
916-924 |
NN |
denotes |
homology |
| T361 |
925-929 |
IN |
denotes |
with |
| T362 |
930-941 |
NNP |
denotes |
Pseudomonas |
| T363 |
942-944 |
NNP |
denotes |
sp |
| T364 |
944-945 |
. |
denotes |
. |
| T365 |
945-960 |
sentence |
denotes |
101 sedolisin. |
| T366 |
946-949 |
CD |
denotes |
101 |
| T367 |
950-959 |
NN |
denotes |
sedolisin |
| T368 |
959-960 |
. |
denotes |
. |
| T369 |
960-972 |
sentence |
denotes |
Conclusion |
| T370 |
962-972 |
NN |
denotes |
Conclusion |
| T371 |
972-1101 |
sentence |
denotes |
CLN2 is very highly conserved and widely distributed among higher organisms and may play an important role in their life cycles. |
| T372 |
973-977 |
NN |
denotes |
CLN2 |
| T373 |
993-1002 |
VBN |
denotes |
conserved |
| T374 |
978-980 |
VBZ |
denotes |
is |
| T375 |
981-985 |
RB |
denotes |
very |
| T376 |
986-992 |
RB |
denotes |
highly |
| T377 |
1003-1006 |
CC |
denotes |
and |
| T378 |
1007-1013 |
RB |
denotes |
widely |
| T379 |
1014-1025 |
VBN |
denotes |
distributed |
| T380 |
1026-1031 |
IN |
denotes |
among |
| T381 |
1032-1038 |
JJR |
denotes |
higher |
| T382 |
1039-1048 |
NNS |
denotes |
organisms |
| T383 |
1049-1052 |
CC |
denotes |
and |
| T384 |
1053-1056 |
MD |
denotes |
may |
| T385 |
1057-1061 |
VB |
denotes |
play |
| T386 |
1062-1064 |
DT |
denotes |
an |
| T387 |
1075-1079 |
NN |
denotes |
role |
| T388 |
1065-1074 |
JJ |
denotes |
important |
| T389 |
1080-1082 |
IN |
denotes |
in |
| T390 |
1083-1088 |
PRP$ |
denotes |
their |
| T391 |
1094-1100 |
NNS |
denotes |
cycles |
| T392 |
1089-1093 |
NN |
denotes |
life |
| T393 |
1100-1101 |
. |
denotes |
. |
| T394 |
1101-1241 |
sentence |
denotes |
The model presented here indicates a very open and accessible active site that is almost completely conserved among all known CLN2 enzymes. |
| T395 |
1102-1105 |
DT |
denotes |
The |
| T396 |
1106-1111 |
NN |
denotes |
model |
| T397 |
1127-1136 |
VBZ |
denotes |
indicates |
| T398 |
1112-1121 |
VBN |
denotes |
presented |
| T399 |
1122-1126 |
RB |
denotes |
here |
| T400 |
1137-1138 |
DT |
denotes |
a |
| T401 |
1171-1175 |
NN |
denotes |
site |
| T402 |
1139-1143 |
RB |
denotes |
very |
| T403 |
1144-1148 |
JJ |
denotes |
open |
| T404 |
1149-1152 |
CC |
denotes |
and |
| T405 |
1153-1163 |
JJ |
denotes |
accessible |
| T406 |
1164-1170 |
JJ |
denotes |
active |
| T407 |
1176-1180 |
WDT |
denotes |
that |
| T408 |
1202-1211 |
VBN |
denotes |
conserved |
| T409 |
1181-1183 |
VBZ |
denotes |
is |
| T410 |
1184-1190 |
RB |
denotes |
almost |
| T411 |
1191-1201 |
RB |
denotes |
completely |
| T412 |
1212-1217 |
IN |
denotes |
among |
| T413 |
1218-1221 |
DT |
denotes |
all |
| T414 |
1233-1240 |
NNS |
denotes |
enzymes |
| T415 |
1222-1227 |
VBN |
denotes |
known |
| T416 |
1228-1232 |
NN |
denotes |
CLN2 |
| T417 |
1240-1241 |
. |
denotes |
. |
| T418 |
1241-1367 |
sentence |
denotes |
This result is somehow surprising for a tripeptidase where the presence of a more constrained binding pocket was anticipated. |
| T419 |
1242-1246 |
DT |
denotes |
This |
| T420 |
1247-1253 |
NN |
denotes |
result |
| T421 |
1254-1256 |
VBZ |
denotes |
is |
| T422 |
1257-1264 |
RB |
denotes |
somehow |
| T423 |
1265-1275 |
JJ |
denotes |
surprising |
| T424 |
1276-1279 |
IN |
denotes |
for |
| T425 |
1280-1281 |
DT |
denotes |
a |
| T426 |
1282-1294 |
NN |
denotes |
tripeptidase |
| T427 |
1295-1300 |
WRB |
denotes |
where |
| T428 |
1355-1366 |
VBN |
denotes |
anticipated |
| T429 |
1301-1304 |
DT |
denotes |
the |
| T430 |
1305-1313 |
NN |
denotes |
presence |
| T431 |
1314-1316 |
IN |
denotes |
of |
| T432 |
1317-1318 |
DT |
denotes |
a |
| T433 |
1344-1350 |
NN |
denotes |
pocket |
| T434 |
1319-1323 |
RBR |
denotes |
more |
| T435 |
1324-1335 |
JJ |
denotes |
constrained |
| T436 |
1336-1343 |
NN |
denotes |
binding |
| T437 |
1351-1354 |
VBD |
denotes |
was |
| T438 |
1366-1367 |
. |
denotes |
. |
| T439 |
1367-1525 |
sentence |
denotes |
This structural model should be useful in the search for the physiological substrates of these enzymes and in the design of more specific inhibitors of CLN2. |
| T440 |
1368-1372 |
DT |
denotes |
This |
| T441 |
1384-1389 |
NN |
denotes |
model |
| T442 |
1373-1383 |
JJ |
denotes |
structural |
| T443 |
1397-1399 |
VB |
denotes |
be |
| T444 |
1390-1396 |
MD |
denotes |
should |
| T445 |
1400-1406 |
JJ |
denotes |
useful |
| T446 |
1407-1409 |
IN |
denotes |
in |
| T447 |
1410-1413 |
DT |
denotes |
the |
| T448 |
1414-1420 |
NN |
denotes |
search |
| T449 |
1421-1424 |
IN |
denotes |
for |
| T450 |
1425-1428 |
DT |
denotes |
the |
| T451 |
1443-1453 |
NNS |
denotes |
substrates |
| T452 |
1429-1442 |
JJ |
denotes |
physiological |
| T453 |
1454-1456 |
IN |
denotes |
of |
| T454 |
1457-1462 |
DT |
denotes |
these |
| T455 |
1463-1470 |
NNS |
denotes |
enzymes |
| T456 |
1471-1474 |
CC |
denotes |
and |
| T457 |
1475-1477 |
IN |
denotes |
in |
| T458 |
1478-1481 |
DT |
denotes |
the |
| T459 |
1482-1488 |
NN |
denotes |
design |
| T460 |
1489-1491 |
IN |
denotes |
of |
| T461 |
1492-1496 |
JJR |
denotes |
more |
| T462 |
1506-1516 |
NNS |
denotes |
inhibitors |
| T463 |
1497-1505 |
JJ |
denotes |
specific |
| T464 |
1517-1519 |
IN |
denotes |
of |
| T465 |
1520-1524 |
NN |
denotes |
CLN2 |
| T466 |
1524-1525 |
. |
denotes |
. |
| R25 |
T184 |
T185 |
compound |
Tripeptidyl,peptidase |
| R26 |
T185 |
T187 |
nsubj |
peptidase,is |
| R28 |
T188 |
T185 |
nummod |
I,peptidase |
| R29 |
T189 |
T185 |
punct |
", ",peptidase |
| R30 |
T190 |
T191 |
advmod |
also,known |
| R31 |
T191 |
T185 |
acl |
known,peptidase |
| R32 |
T192 |
T191 |
prep |
as,known |
| R33 |
T193 |
T192 |
pobj |
CLN2,as |
| R34 |
T194 |
T187 |
punct |
", ",is |
| R35 |
T195 |
T196 |
det |
a,member |
| R36 |
T196 |
T187 |
attr |
member,is |
| R37 |
T197 |
T196 |
prep |
of,member |
| R38 |
T198 |
T199 |
det |
the,family |
| R39 |
T199 |
T197 |
pobj |
family,of |
| R40 |
T200 |
T199 |
prep |
of,family |
| R41 |
T201 |
T200 |
pobj |
sedolisins,of |
| R42 |
T202 |
T201 |
punct |
(,sedolisins |
| R43 |
T203 |
T204 |
compound |
serine,carboxyl |
| R44 |
T204 |
T206 |
compound |
carboxyl,peptidases |
| R46 |
T206 |
T201 |
appos |
peptidases,sedolisins |
| R47 |
T207 |
T187 |
punct |
),is |
| R48 |
T208 |
T187 |
punct |
.,is |
| R49 |
T210 |
T211 |
prep |
In,lead |
| R50 |
T212 |
T210 |
pobj |
humans,In |
| R51 |
T213 |
T211 |
punct |
", ",lead |
| R52 |
T214 |
T211 |
nsubj |
defects,lead |
| R53 |
T215 |
T214 |
prep |
in,defects |
| R54 |
T216 |
T215 |
pobj |
expression,in |
| R55 |
T217 |
T216 |
prep |
of,expression |
| R56 |
T218 |
T219 |
det |
this,enzyme |
| R57 |
T219 |
T217 |
pobj |
enzyme,of |
| R58 |
T220 |
T211 |
prep |
to,lead |
| R59 |
T221 |
T222 |
det |
a,disease |
| R60 |
T222 |
T220 |
pobj |
disease,to |
| R63 |
T225 |
T222 |
punct |
", ",disease |
| R64 |
T226 |
T227 |
amod |
classical,lipofuscinosis |
| R65 |
T227 |
T222 |
appos |
lipofuscinosis,disease |
| R67 |
T229 |
T227 |
amod |
infantile,lipofuscinosis |
| R68 |
T230 |
T229 |
punct |
-,infantile |
| R71 |
T233 |
T211 |
punct |
.,lead |
| R72 |
T235 |
T236 |
amod |
Similar,enzymes |
| R73 |
T236 |
T237 |
nsubjpass |
enzymes,found |
| R74 |
T238 |
T237 |
aux |
have,found |
| R75 |
T239 |
T237 |
auxpass |
been,found |
| R76 |
T240 |
T237 |
prep |
in,found |
| R77 |
T241 |
T242 |
det |
the,sequences |
| R78 |
T242 |
T240 |
pobj |
sequences,in |
| R80 |
T244 |
T242 |
prep |
of,sequences |
| R81 |
T245 |
T246 |
amod |
several,species |
| R82 |
T246 |
T244 |
pobj |
species,of |
| R83 |
T247 |
T237 |
punct |
", ",found |
| R84 |
T248 |
T237 |
cc |
but,found |
| R85 |
T249 |
T250 |
preconj |
neither,analyses |
| R86 |
T250 |
T252 |
nsubjpass |
analyses,attempted |
| R88 |
T252 |
T237 |
conj |
attempted,found |
| R100 |
T264 |
T252 |
punct |
.,attempted |
| R101 |
T268 |
T269 |
nsubj |
We,analyzed |
| R102 |
T270 |
T269 |
aux |
have,analyzed |
| R103 |
T271 |
T272 |
det |
the,presence |
| R104 |
T272 |
T269 |
dobj |
presence,analyzed |
| R105 |
T273 |
T272 |
prep |
of,presence |
| R106 |
T274 |
T273 |
pobj |
orthologs,of |
| R107 |
T275 |
T274 |
prep |
of,orthologs |
| R108 |
T276 |
T277 |
amod |
human,CLN2 |
| R109 |
T277 |
T275 |
pobj |
CLN2,of |
| R110 |
T278 |
T269 |
prep |
in,analyzed |
| R111 |
T279 |
T280 |
det |
the,sequences |
| R112 |
T280 |
T278 |
pobj |
sequences,in |
| R114 |
T282 |
T280 |
prep |
of,sequences |
| R115 |
T283 |
T284 |
det |
a,number |
| R116 |
T284 |
T282 |
pobj |
number,of |
| R117 |
T285 |
T284 |
prep |
of,number |
| R118 |
T286 |
T287 |
amod |
eukaryotic,species |
| R119 |
T287 |
T285 |
pobj |
species,of |
| R120 |
T288 |
T269 |
punct |
.,analyzed |
| R121 |
T290 |
T291 |
nsubjpass |
Enzymes,found |
| R122 |
T292 |
T290 |
prep |
with,Enzymes |
| R123 |
T293 |
T292 |
pobj |
sequences,with |
| R124 |
T294 |
T293 |
acl |
sharing,sequences |
| R125 |
T295 |
T296 |
quantmod |
over,80 |
| R126 |
T296 |
T297 |
nummod |
80,% |
| R127 |
T297 |
T298 |
compound |
%,identity |
| R128 |
T298 |
T294 |
dobj |
identity,sharing |
| R129 |
T299 |
T291 |
aux |
have,found |
| R130 |
T300 |
T291 |
auxpass |
been,found |
| R131 |
T301 |
T291 |
prep |
in,found |
| R132 |
T302 |
T303 |
det |
the,genomes |
| R133 |
T303 |
T301 |
pobj |
genomes,in |
| R134 |
T304 |
T303 |
prep |
of,genomes |
| R135 |
T305 |
T304 |
pobj |
macaque,of |
| R136 |
T306 |
T305 |
punct |
", ",macaque |
| R137 |
T307 |
T305 |
conj |
mouse,macaque |
| R138 |
T308 |
T307 |
punct |
", ",mouse |
| R139 |
T309 |
T307 |
conj |
rat,mouse |
| R140 |
T310 |
T309 |
punct |
", ",rat |
| R141 |
T311 |
T309 |
conj |
dog,rat |
| R142 |
T312 |
T311 |
punct |
", ",dog |
| R143 |
T313 |
T311 |
cc |
and,dog |
| R144 |
T314 |
T311 |
conj |
cow,dog |
| R145 |
T315 |
T291 |
punct |
.,found |
| R146 |
T317 |
T318 |
advmod |
Closely,related |
| R147 |
T318 |
T319 |
amod |
related,enzymes |
| R148 |
T319 |
T325 |
nsubj |
enzymes,are |
| R150 |
T321 |
T318 |
parataxis |
distinct,related |
| R151 |
T322 |
T321 |
mark |
although,distinct |
| R152 |
T323 |
T321 |
advmod |
clearly,distinct |
| R154 |
T326 |
T325 |
acomp |
present,are |
| R155 |
T327 |
T325 |
prep |
in,are |
| R156 |
T328 |
T327 |
pobj |
fish,in |
| R157 |
T329 |
T328 |
punct |
(,fish |
| R158 |
T330 |
T328 |
appos |
fugu,fish |
| R159 |
T331 |
T330 |
cc |
and,fugu |
| R160 |
T332 |
T330 |
conj |
zebra,fugu |
| R161 |
T333 |
T327 |
punct |
),in |
| R162 |
T334 |
T327 |
punct |
", ",in |
| R163 |
T335 |
T336 |
advmod |
as,as |
| R164 |
T336 |
T327 |
cc |
as,in |
| R166 |
T338 |
T327 |
conj |
in,in |
| R167 |
T339 |
T338 |
pobj |
frogs,in |
| R168 |
T340 |
T339 |
punct |
(,frogs |
| R169 |
T341 |
T342 |
compound |
Xenopus,tropicalis |
| R170 |
T342 |
T339 |
appos |
tropicalis,frogs |
| R171 |
T343 |
T325 |
punct |
),are |
| R172 |
T344 |
T325 |
punct |
.,are |
| R173 |
T346 |
T347 |
det |
A,model |
| R174 |
T347 |
T351 |
nsubjpass |
model,built |
| R176 |
T349 |
T347 |
amod |
dimensional,model |
| R177 |
T350 |
T349 |
punct |
-,dimensional |
| R178 |
T352 |
T347 |
prep |
of,model |
| R179 |
T353 |
T354 |
amod |
human,CLN2 |
| R180 |
T354 |
T352 |
pobj |
CLN2,of |
| R181 |
T355 |
T351 |
auxpass |
was,built |
| R182 |
T356 |
T351 |
prep |
based,built |
| R183 |
T357 |
T358 |
advmod |
mainly,on |
| R184 |
T358 |
T356 |
prep |
on,based |
| R185 |
T359 |
T360 |
det |
the,homology |
| R186 |
T360 |
T358 |
pobj |
homology,on |
| R187 |
T361 |
T360 |
prep |
with,homology |
| R188 |
T362 |
T363 |
compound |
Pseudomonas,sp |
| R189 |
T363 |
T361 |
pobj |
sp,with |
| R190 |
T364 |
T351 |
punct |
.,built |
| R191 |
T366 |
T367 |
nummod |
101,sedolisin |
| R192 |
T368 |
T367 |
punct |
.,sedolisin |
| R193 |
T372 |
T373 |
nsubj |
CLN2,conserved |
| R194 |
T374 |
T373 |
aux |
is,conserved |
| R195 |
T375 |
T376 |
advmod |
very,highly |
| R196 |
T376 |
T373 |
advmod |
highly,conserved |
| R197 |
T377 |
T373 |
cc |
and,conserved |
| R198 |
T378 |
T379 |
advmod |
widely,distributed |
| R199 |
T379 |
T373 |
conj |
distributed,conserved |
| R200 |
T380 |
T373 |
prep |
among,conserved |
| R201 |
T381 |
T382 |
amod |
higher,organisms |
| R202 |
T382 |
T380 |
pobj |
organisms,among |
| R203 |
T383 |
T373 |
cc |
and,conserved |
| R204 |
T384 |
T385 |
aux |
may,play |
| R205 |
T385 |
T373 |
conj |
play,conserved |
| R206 |
T386 |
T387 |
det |
an,role |
| R207 |
T387 |
T385 |
dobj |
role,play |
| R209 |
T389 |
T385 |
prep |
in,play |
| R210 |
T390 |
T391 |
poss |
their,cycles |
| R211 |
T391 |
T389 |
pobj |
cycles,in |
| R213 |
T393 |
T373 |
punct |
.,conserved |
| R214 |
T395 |
T396 |
det |
The,model |
| R215 |
T396 |
T397 |
nsubj |
model,indicates |
| R216 |
T398 |
T396 |
acl |
presented,model |
| R217 |
T399 |
T398 |
advmod |
here,presented |
| R218 |
T400 |
T401 |
det |
a,site |
| R219 |
T401 |
T397 |
dobj |
site,indicates |
| R225 |
T407 |
T408 |
dep |
that,conserved |
| R226 |
T408 |
T401 |
relcl |
conserved,site |
| R230 |
T412 |
T408 |
prep |
among,conserved |
| R231 |
T413 |
T414 |
det |
all,enzymes |
| R232 |
T414 |
T412 |
pobj |
enzymes,among |
| R235 |
T417 |
T397 |
punct |
.,indicates |
| R236 |
T419 |
T420 |
det |
This,result |
| R237 |
T420 |
T421 |
nsubj |
result,is |
| R238 |
T422 |
T421 |
advmod |
somehow,is |
| R239 |
T423 |
T421 |
acomp |
surprising,is |
| R240 |
T424 |
T421 |
prep |
for,is |
| R241 |
T425 |
T426 |
det |
a,tripeptidase |
| R242 |
T426 |
T424 |
pobj |
tripeptidase,for |
| R243 |
T427 |
T428 |
advmod |
where,anticipated |
| R244 |
T428 |
T426 |
relcl |
anticipated,tripeptidase |
| R249 |
T433 |
T431 |
pobj |
pocket,of |
| R250 |
T434 |
T435 |
advmod |
more,constrained |
| R251 |
T435 |
T433 |
amod |
constrained,pocket |
| R252 |
T436 |
T433 |
compound |
binding,pocket |
| R254 |
T438 |
T421 |
punct |
.,is |
| R255 |
T440 |
T441 |
det |
This,model |
| R256 |
T441 |
T443 |
nsubj |
model,be |
| R258 |
T444 |
T443 |
aux |
should,be |
| R259 |
T445 |
T443 |
acomp |
useful,be |
| R260 |
T446 |
T443 |
prep |
in,be |
| R261 |
T447 |
T448 |
det |
the,search |
| R262 |
T448 |
T446 |
pobj |
search,in |
| R263 |
T449 |
T448 |
prep |
for,search |
| R264 |
T450 |
T451 |
det |
the,substrates |
| R265 |
T451 |
T449 |
pobj |
substrates,for |
| R267 |
T453 |
T451 |
prep |
of,substrates |
| R268 |
T454 |
T455 |
det |
these,enzymes |
| R269 |
T455 |
T453 |
pobj |
enzymes,of |
| R270 |
T456 |
T446 |
cc |
and,in |
| R271 |
T457 |
T446 |
conj |
in,in |
| R272 |
T458 |
T459 |
det |
the,design |
| R273 |
T459 |
T457 |
pobj |
design,in |
| R274 |
T460 |
T459 |
prep |
of,design |
| R275 |
T461 |
T462 |
amod |
more,inhibitors |
| R276 |
T462 |
T460 |
pobj |
inhibitors,of |
| R278 |
T464 |
T462 |
prep |
of,inhibitors |
| R279 |
T465 |
T464 |
pobj |
CLN2,of |
| R280 |
T466 |
T443 |
punct |
.,be |
| R70 |
T232 |
T227 |
amod |
ceroid,lipofuscinosis |
| R79 |
T243 |
T242 |
amod |
genomic,sequences |
| R87 |
T251 |
T250 |
amod |
systematic,analyses |
| R89 |
T253 |
T250 |
prep |
of,analyses |
| R90 |
T254 |
T255 |
poss |
their,distribution |
| R91 |
T255 |
T253 |
pobj |
distribution,of |
| R92 |
T256 |
T250 |
cc |
nor,analyses |
| R93 |
T257 |
T250 |
conj |
modeling,analyses |
| R94 |
T258 |
T257 |
prep |
of,modeling |
| R95 |
T259 |
T260 |
poss |
their,structures |
| R96 |
T260 |
T258 |
pobj |
structures,of |
| R97 |
T261 |
T252 |
aux |
have,attempted |
| R98 |
T262 |
T252 |
auxpass |
been,attempted |
| R99 |
T263 |
T252 |
advmod |
previously,attempted |
| R113 |
T281 |
T280 |
amod |
genomic,sequences |
| R149 |
T320 |
T321 |
punct |
", ",distinct |
| R153 |
T324 |
T321 |
punct |
", ",distinct |
| R165 |
T337 |
T336 |
advmod |
well,as |
| R175 |
T348 |
T349 |
advmod |
three,dimensional |
| R208 |
T388 |
T387 |
amod |
important,role |
| R212 |
T392 |
T391 |
compound |
life,cycles |
| R220 |
T402 |
T403 |
advmod |
very,open |
| R221 |
T403 |
T401 |
amod |
open,site |
| R222 |
T404 |
T403 |
cc |
and,open |
| R223 |
T405 |
T403 |
conj |
accessible,open |
| R224 |
T406 |
T401 |
amod |
active,site |
| R227 |
T409 |
T408 |
auxpass |
is,conserved |
| R228 |
T410 |
T411 |
advmod |
almost,completely |
| R229 |
T411 |
T408 |
advmod |
completely,conserved |
| R233 |
T415 |
T414 |
amod |
known,enzymes |
| R234 |
T416 |
T414 |
compound |
CLN2,enzymes |
| R245 |
T429 |
T430 |
det |
the,presence |
| R27 |
T186 |
T185 |
punct |
-,peptidase |
| R45 |
T205 |
T204 |
punct |
-,carboxyl |
| R61 |
T223 |
T222 |
amod |
fatal,disease |
| R62 |
T224 |
T222 |
amod |
neurodegenerative,disease |
| R66 |
T228 |
T229 |
amod |
late,infantile |
| R69 |
T231 |
T227 |
amod |
neuronal,lipofuscinosis |
| R246 |
T430 |
T428 |
nsubjpass |
presence,anticipated |
| R247 |
T431 |
T430 |
prep |
of,presence |
| R248 |
T432 |
T433 |
det |
a,pocket |
| R253 |
T437 |
T428 |
auxpass |
was,anticipated |
| R257 |
T442 |
T441 |
amod |
structural,model |
| R266 |
T452 |
T451 |
amod |
physiological,substrates |
| R277 |
T463 |
T462 |
amod |
specific,inhibitors |