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genes were identified as potential candidate genes for six of the eight small-effect QTL according to their gene expression patterns during CIA and their genomic locations. 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No candidate genes were located in QTL 4 and 7, suggesting that QTG polymorphisms of the susceptibility genes inside these two QTL might not affect the phenotype by regulating gene expression. Two of the 21 candidate genes were reported to be involved in arthritis. Mapk14, a candidate gene for locus 8 and also called p38 mitogen-activated protein kinase (MAPK) alpha, regulates the production of arthritis-essential cytokines, such as tumour necrosis factor and interleukin-1 [32]. Moreover, inhibitors of p38 MAPK could attenuate CIA in rats [33], and p38 MAPK is becoming a potential therapeutic target in RA [32]. Stat5a, a candidate gene for loci 6, is an essential molecule for lymphoid development and differentiation [34]. Stat5a-deficient mice were reported to lose tolerance, resulting in the development of autoimmune diseases. Stat5a is suggested to contribute to tolerance through maintenance of the CD4+CD25+ regulatory T cell population [35]."}

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