Id |
Subject |
Object |
Predicate |
Lexical cue |
T3017 |
0-3 |
DT |
denotes |
The |
T3018 |
14-20 |
NN |
denotes |
genome |
T3019 |
4-13 |
JJ |
denotes |
mammalian |
T3020 |
21-28 |
VBZ |
denotes |
carries |
T3021 |
29-32 |
CD |
denotes |
two |
T3022 |
48-57 |
NNS |
denotes |
orthologs |
T3023 |
32-34 |
, |
denotes |
, |
T3024 |
34-37 |
CC |
denotes |
and |
T3025 |
38-42 |
RB |
denotes |
only |
T3026 |
43-46 |
CD |
denotes |
two |
T3027 |
46-48 |
, |
denotes |
, |
T3028 |
58-60 |
IN |
denotes |
of |
T3029 |
61-71 |
NNP |
denotes |
Drosophila |
T3030 |
72-76 |
NN |
denotes |
Pygo |
T3031 |
76-78 |
, |
denotes |
, |
T3032 |
78-83 |
NN |
denotes |
Pygo1 |
T3033 |
84-87 |
CC |
denotes |
and |
T3034 |
88-93 |
NN |
denotes |
Pygo2 |
T3035 |
94-95 |
-LRB- |
denotes |
[ |
T3036 |
99-101 |
CD |
denotes |
14 |
T3037 |
95-96 |
CD |
denotes |
8 |
T3038 |
96-97 |
, |
denotes |
, |
T3039 |
97-98 |
CD |
denotes |
9 |
T3040 |
98-99 |
, |
denotes |
, |
T3041 |
101-102 |
-RRB- |
denotes |
] |
T3042 |
102-103 |
. |
denotes |
. |
T3043 |
103-317 |
sentence |
denotes |
In this report, we generated targeted mutations of Pygo1 and Pygo2 to determine their functions, with a particular interest in the contributions of these genes to canonical Wnt signaling during kidney development. |
T3044 |
104-106 |
IN |
denotes |
In |
T3045 |
123-132 |
VBD |
denotes |
generated |
T3046 |
107-111 |
DT |
denotes |
this |
T3047 |
112-118 |
NN |
denotes |
report |
T3048 |
118-120 |
, |
denotes |
, |
T3049 |
120-122 |
PRP |
denotes |
we |
T3050 |
133-141 |
VBN |
denotes |
targeted |
T3051 |
142-151 |
NNS |
denotes |
mutations |
T3052 |
152-154 |
IN |
denotes |
of |
T3053 |
155-160 |
NN |
denotes |
Pygo1 |
T3054 |
161-164 |
CC |
denotes |
and |
T3055 |
165-170 |
NN |
denotes |
Pygo2 |
T3056 |
171-173 |
TO |
denotes |
to |
T3057 |
174-183 |
VB |
denotes |
determine |
T3058 |
184-189 |
PRP$ |
denotes |
their |
T3059 |
190-199 |
NNS |
denotes |
functions |
T3060 |
199-201 |
, |
denotes |
, |
T3061 |
201-205 |
IN |
denotes |
with |
T3062 |
206-207 |
DT |
denotes |
a |
T3063 |
219-227 |
NN |
denotes |
interest |
T3064 |
208-218 |
JJ |
denotes |
particular |
T3065 |
228-230 |
IN |
denotes |
in |
T3066 |
231-234 |
DT |
denotes |
the |
T3067 |
235-248 |
NNS |
denotes |
contributions |
T3068 |
249-251 |
IN |
denotes |
of |
T3069 |
252-257 |
DT |
denotes |
these |
T3070 |
258-263 |
NNS |
denotes |
genes |
T3071 |
264-266 |
IN |
denotes |
to |
T3072 |
267-276 |
JJ |
denotes |
canonical |
T3073 |
281-290 |
NN |
denotes |
signaling |
T3074 |
277-280 |
NN |
denotes |
Wnt |
T3075 |
291-297 |
IN |
denotes |
during |
T3076 |
298-304 |
NN |
denotes |
kidney |
T3077 |
305-316 |
NN |
denotes |
development |
T3078 |
316-317 |
. |
denotes |
. |
T3079 |
317-478 |
sentence |
denotes |
The resulting double-homozygous mutant embryos showed a context-dependent reduction in canonical Wnt signaling as measured by Wnt reporter transgene expression. |
T3080 |
318-321 |
DT |
denotes |
The |
T3081 |
357-364 |
NNS |
denotes |
embryos |
T3082 |
322-331 |
VBG |
denotes |
resulting |
T3083 |
332-338 |
JJ |
denotes |
double |
T3084 |
339-349 |
JJ |
denotes |
homozygous |
T3085 |
338-339 |
HYPH |
denotes |
- |
T3086 |
350-356 |
NN |
denotes |
mutant |
T3087 |
365-371 |
VBD |
denotes |
showed |
T3088 |
372-373 |
DT |
denotes |
a |
T3089 |
392-401 |
NN |
denotes |
reduction |
T3090 |
374-381 |
NN |
denotes |
context |
T3091 |
382-391 |
JJ |
denotes |
dependent |
T3092 |
381-382 |
HYPH |
denotes |
- |
T3093 |
402-404 |
IN |
denotes |
in |
T3094 |
405-414 |
JJ |
denotes |
canonical |
T3095 |
419-428 |
NN |
denotes |
signaling |
T3096 |
415-418 |
NN |
denotes |
Wnt |
T3097 |
429-431 |
IN |
denotes |
as |
T3098 |
432-440 |
VBN |
denotes |
measured |
T3099 |
441-443 |
IN |
denotes |
by |
T3100 |
444-447 |
NN |
denotes |
Wnt |
T3101 |
448-456 |
NN |
denotes |
reporter |
T3102 |
457-466 |
NN |
denotes |
transgene |
T3103 |
467-477 |
NN |
denotes |
expression |
T3104 |
477-478 |
. |
denotes |
. |
T3105 |
478-601 |
sentence |
denotes |
Development remained, however, surprisingly normal, with survival to birth and few apparent defects in most organ systems. |
T3106 |
479-490 |
NN |
denotes |
Development |
T3107 |
491-499 |
VBD |
denotes |
remained |
T3108 |
499-501 |
, |
denotes |
, |
T3109 |
501-508 |
RB |
denotes |
however |
T3110 |
508-510 |
, |
denotes |
, |
T3111 |
510-522 |
RB |
denotes |
surprisingly |
T3112 |
523-529 |
JJ |
denotes |
normal |
T3113 |
529-531 |
, |
denotes |
, |
T3114 |
531-535 |
IN |
denotes |
with |
T3115 |
536-544 |
NN |
denotes |
survival |
T3116 |
545-547 |
IN |
denotes |
to |
T3117 |
548-553 |
NN |
denotes |
birth |
T3118 |
554-557 |
CC |
denotes |
and |
T3119 |
558-561 |
JJ |
denotes |
few |
T3120 |
571-578 |
NNS |
denotes |
defects |
T3121 |
562-570 |
JJ |
denotes |
apparent |
T3122 |
579-581 |
IN |
denotes |
in |
T3123 |
582-586 |
JJS |
denotes |
most |
T3124 |
593-600 |
NNS |
denotes |
systems |
T3125 |
587-592 |
NN |
denotes |
organ |
T3126 |
600-601 |
. |
denotes |
. |
T3127 |
601-927 |
sentence |
denotes |
Our phenotypic analysis focused on the kidney, which showed altered branching morphogenesis of the ureteric bud, and expansion of the zone of condensed mesenchyme surrounding the ureteric bud, yet relatively normal nephron formation, as measured by histology, confocal analysis, in situ hybridization and microarray analysis. |
T3128 |
602-605 |
PRP$ |
denotes |
Our |
T3129 |
617-625 |
NN |
denotes |
analysis |
T3130 |
606-616 |
JJ |
denotes |
phenotypic |
T3131 |
626-633 |
VBD |
denotes |
focused |
T3132 |
634-636 |
IN |
denotes |
on |
T3133 |
637-640 |
DT |
denotes |
the |
T3134 |
641-647 |
NN |
denotes |
kidney |
T3135 |
647-649 |
, |
denotes |
, |
T3136 |
649-654 |
WDT |
denotes |
which |
T3137 |
655-661 |
VBD |
denotes |
showed |
T3138 |
662-669 |
VBN |
denotes |
altered |
T3139 |
680-693 |
NN |
denotes |
morphogenesis |
T3140 |
670-679 |
NN |
denotes |
branching |
T3141 |
694-696 |
IN |
denotes |
of |
T3142 |
697-700 |
DT |
denotes |
the |
T3143 |
710-713 |
NN |
denotes |
bud |
T3144 |
701-709 |
JJ |
denotes |
ureteric |
T3145 |
713-715 |
, |
denotes |
, |
T3146 |
715-718 |
CC |
denotes |
and |
T3147 |
719-728 |
NN |
denotes |
expansion |
T3148 |
729-731 |
IN |
denotes |
of |
T3149 |
732-735 |
DT |
denotes |
the |
T3150 |
736-740 |
NN |
denotes |
zone |
T3151 |
741-743 |
IN |
denotes |
of |
T3152 |
744-753 |
VBN |
denotes |
condensed |
T3153 |
754-764 |
NN |
denotes |
mesenchyme |
T3154 |
765-776 |
VBG |
denotes |
surrounding |
T3155 |
777-780 |
DT |
denotes |
the |
T3156 |
790-793 |
NN |
denotes |
bud |
T3157 |
781-789 |
JJ |
denotes |
ureteric |
T3158 |
793-795 |
, |
denotes |
, |
T3159 |
795-798 |
CC |
denotes |
yet |
T3160 |
799-809 |
RB |
denotes |
relatively |
T3161 |
810-816 |
JJ |
denotes |
normal |
T3162 |
825-834 |
NN |
denotes |
formation |
T3163 |
817-824 |
NN |
denotes |
nephron |
T3164 |
834-836 |
, |
denotes |
, |
T3165 |
836-838 |
IN |
denotes |
as |
T3166 |
839-847 |
VBN |
denotes |
measured |
T3167 |
848-850 |
IN |
denotes |
by |
T3168 |
851-860 |
NN |
denotes |
histology |
T3169 |
860-862 |
, |
denotes |
, |
T3170 |
862-870 |
JJ |
denotes |
confocal |
T3171 |
871-879 |
NN |
denotes |
analysis |
T3172 |
879-881 |
, |
denotes |
, |
T3173 |
881-883 |
FW |
denotes |
in |
T3174 |
884-888 |
FW |
denotes |
situ |
T3175 |
889-902 |
NN |
denotes |
hybridization |
T3176 |
903-906 |
CC |
denotes |
and |
T3177 |
907-917 |
NN |
denotes |
microarray |
T3178 |
918-926 |
NN |
denotes |
analysis |
T3179 |
926-927 |
. |
denotes |
. |
T3180 |
927-1092 |
sentence |
denotes |
The obvious conclusion is that in mammals, unlike Drosophila, Pygopus-mediated canonical Wnt signaling is not absolutely necessary in most developing organ systems. |
T3181 |
928-931 |
DT |
denotes |
The |
T3182 |
940-950 |
NN |
denotes |
conclusion |
T3183 |
932-939 |
JJ |
denotes |
obvious |
T3184 |
951-953 |
VBZ |
denotes |
is |
T3185 |
954-958 |
IN |
denotes |
that |
T3186 |
1031-1033 |
VBZ |
denotes |
is |
T3187 |
959-961 |
IN |
denotes |
in |
T3188 |
962-969 |
NNS |
denotes |
mammals |
T3189 |
969-971 |
, |
denotes |
, |
T3190 |
971-977 |
IN |
denotes |
unlike |
T3191 |
978-988 |
NNP |
denotes |
Drosophila |
T3192 |
988-990 |
, |
denotes |
, |
T3193 |
990-997 |
NN |
denotes |
Pygopus |
T3194 |
998-1006 |
VBN |
denotes |
mediated |
T3195 |
997-998 |
HYPH |
denotes |
- |
T3196 |
1021-1030 |
NN |
denotes |
signaling |
T3197 |
1007-1016 |
JJ |
denotes |
canonical |
T3198 |
1017-1020 |
NN |
denotes |
Wnt |
T3199 |
1034-1037 |
RB |
denotes |
not |
T3200 |
1038-1048 |
RB |
denotes |
absolutely |
T3201 |
1049-1058 |
JJ |
denotes |
necessary |
T3202 |
1059-1061 |
IN |
denotes |
in |
T3203 |
1062-1066 |
JJS |
denotes |
most |
T3204 |
1084-1091 |
NNS |
denotes |
systems |
T3205 |
1067-1077 |
VBG |
denotes |
developing |
T3206 |
1078-1083 |
NN |
denotes |
organ |
T3207 |
1091-1092 |
. |
denotes |
. |
R751 |
T3017 |
T3018 |
det |
The,genome |
R752 |
T3018 |
T3020 |
nsubj |
genome,carries |
R753 |
T3019 |
T3018 |
amod |
mammalian,genome |
R754 |
T3021 |
T3022 |
nummod |
two,orthologs |
R755 |
T3022 |
T3020 |
dobj |
orthologs,carries |
R756 |
T3023 |
T3021 |
punct |
", ",two |
R757 |
T3024 |
T3021 |
cc |
and,two |
R758 |
T3025 |
T3026 |
advmod |
only,two |
R759 |
T3026 |
T3021 |
conj |
two,two |
R760 |
T3027 |
T3022 |
punct |
", ",orthologs |
R761 |
T3028 |
T3022 |
prep |
of,orthologs |
R762 |
T3029 |
T3030 |
compound |
Drosophila,Pygo |
R763 |
T3030 |
T3028 |
pobj |
Pygo,of |
R764 |
T3031 |
T3022 |
punct |
", ",orthologs |
R765 |
T3032 |
T3022 |
appos |
Pygo1,orthologs |
R766 |
T3033 |
T3032 |
cc |
and,Pygo1 |
R767 |
T3140 |
T3139 |
compound |
branching,morphogenesis |
R768 |
T3141 |
T3139 |
prep |
of,morphogenesis |
R769 |
T3142 |
T3143 |
det |
the,bud |
R770 |
T3143 |
T3141 |
pobj |
bud,of |
R771 |
T3144 |
T3143 |
amod |
ureteric,bud |
R772 |
T3145 |
T3139 |
punct |
", ",morphogenesis |
R773 |
T3146 |
T3139 |
cc |
and,morphogenesis |
R774 |
T3147 |
T3139 |
conj |
expansion,morphogenesis |
R775 |
T3148 |
T3147 |
prep |
of,expansion |
R776 |
T3149 |
T3150 |
det |
the,zone |
R777 |
T3150 |
T3148 |
pobj |
zone,of |
R778 |
T3151 |
T3150 |
prep |
of,zone |
R779 |
T3152 |
T3153 |
amod |
condensed,mesenchyme |
R780 |
T3153 |
T3151 |
pobj |
mesenchyme,of |
R781 |
T3034 |
T3032 |
conj |
Pygo2,Pygo1 |
R782 |
T3154 |
T3150 |
acl |
surrounding,zone |
R783 |
T3035 |
T3036 |
punct |
[,14 |
R784 |
T3155 |
T3156 |
det |
the,bud |
R785 |
T3036 |
T3020 |
parataxis |
14,carries |
R786 |
T3037 |
T3036 |
nummod |
8,14 |
R787 |
T3156 |
T3154 |
dobj |
bud,surrounding |
R788 |
T3038 |
T3036 |
punct |
",",14 |
R789 |
T3039 |
T3036 |
nummod |
9,14 |
R790 |
T3040 |
T3036 |
punct |
",",14 |
R791 |
T3157 |
T3156 |
amod |
ureteric,bud |
R792 |
T3041 |
T3036 |
punct |
],14 |
R793 |
T3042 |
T3020 |
punct |
.,carries |
R794 |
T3158 |
T3139 |
punct |
", ",morphogenesis |
R795 |
T3044 |
T3045 |
prep |
In,generated |
R796 |
T3159 |
T3139 |
cc |
yet,morphogenesis |
R797 |
T3046 |
T3047 |
det |
this,report |
R798 |
T3047 |
T3044 |
pobj |
report,In |
R799 |
T3160 |
T3161 |
advmod |
relatively,normal |
R800 |
T3048 |
T3045 |
punct |
", ",generated |
R801 |
T3049 |
T3045 |
nsubj |
we,generated |
R802 |
T3050 |
T3051 |
amod |
targeted,mutations |
R803 |
T3051 |
T3045 |
dobj |
mutations,generated |
R804 |
T3161 |
T3162 |
amod |
normal,formation |
R805 |
T3052 |
T3051 |
prep |
of,mutations |
R806 |
T3053 |
T3052 |
pobj |
Pygo1,of |
R807 |
T3054 |
T3053 |
cc |
and,Pygo1 |
R808 |
T3162 |
T3139 |
conj |
formation,morphogenesis |
R809 |
T3055 |
T3053 |
conj |
Pygo2,Pygo1 |
R810 |
T3056 |
T3057 |
aux |
to,determine |
R811 |
T3163 |
T3162 |
compound |
nephron,formation |
R812 |
T3057 |
T3045 |
advcl |
determine,generated |
R813 |
T3058 |
T3059 |
poss |
their,functions |
R814 |
T3059 |
T3057 |
dobj |
functions,determine |
R815 |
T3060 |
T3057 |
punct |
", ",determine |
R816 |
T3061 |
T3057 |
prep |
with,determine |
R817 |
T3062 |
T3063 |
det |
a,interest |
R818 |
T3164 |
T3137 |
punct |
", ",showed |
R819 |
T3063 |
T3061 |
pobj |
interest,with |
R820 |
T3064 |
T3063 |
amod |
particular,interest |
R821 |
T3065 |
T3063 |
prep |
in,interest |
R822 |
T3165 |
T3166 |
mark |
as,measured |
R823 |
T3066 |
T3067 |
det |
the,contributions |
R824 |
T3067 |
T3065 |
pobj |
contributions,in |
R825 |
T3166 |
T3137 |
advcl |
measured,showed |
R826 |
T3068 |
T3067 |
prep |
of,contributions |
R827 |
T3069 |
T3070 |
det |
these,genes |
R828 |
T3167 |
T3166 |
prep |
by,measured |
R829 |
T3070 |
T3068 |
pobj |
genes,of |
R830 |
T3071 |
T3067 |
prep |
to,contributions |
R831 |
T3072 |
T3073 |
amod |
canonical,signaling |
R832 |
T3168 |
T3167 |
pobj |
histology,by |
R833 |
T3073 |
T3071 |
pobj |
signaling,to |
R834 |
T3074 |
T3073 |
compound |
Wnt,signaling |
R835 |
T3075 |
T3073 |
prep |
during,signaling |
R836 |
T3169 |
T3168 |
punct |
", ",histology |
R837 |
T3076 |
T3077 |
compound |
kidney,development |
R838 |
T3077 |
T3075 |
pobj |
development,during |
R839 |
T3078 |
T3045 |
punct |
.,generated |
R840 |
T3170 |
T3171 |
amod |
confocal,analysis |
R841 |
T3080 |
T3081 |
det |
The,embryos |
R842 |
T3081 |
T3087 |
nsubj |
embryos,showed |
R843 |
T3171 |
T3168 |
conj |
analysis,histology |
R844 |
T3082 |
T3081 |
amod |
resulting,embryos |
R845 |
T3083 |
T3084 |
amod |
double,homozygous |
R846 |
T3084 |
T3081 |
amod |
homozygous,embryos |
R847 |
T3172 |
T3171 |
punct |
", ",analysis |
R848 |
T3085 |
T3084 |
punct |
-,homozygous |
R849 |
T3086 |
T3081 |
compound |
mutant,embryos |
R850 |
T3173 |
T3174 |
advmod |
in,situ |
R851 |
T3088 |
T3089 |
det |
a,reduction |
R852 |
T3089 |
T3087 |
dobj |
reduction,showed |
R853 |
T3174 |
T3175 |
amod |
situ,hybridization |
R854 |
T3090 |
T3091 |
npadvmod |
context,dependent |
R855 |
T3091 |
T3089 |
amod |
dependent,reduction |
R856 |
T3092 |
T3091 |
punct |
-,dependent |
R857 |
T3093 |
T3089 |
prep |
in,reduction |
R858 |
T3175 |
T3171 |
conj |
hybridization,analysis |
R859 |
T3094 |
T3095 |
amod |
canonical,signaling |
R860 |
T3095 |
T3093 |
pobj |
signaling,in |
R861 |
T3176 |
T3175 |
cc |
and,hybridization |
R862 |
T3096 |
T3095 |
compound |
Wnt,signaling |
R863 |
T3097 |
T3098 |
mark |
as,measured |
R864 |
T3098 |
T3087 |
advcl |
measured,showed |
R865 |
T3177 |
T3178 |
compound |
microarray,analysis |
R866 |
T3099 |
T3098 |
agent |
by,measured |
R867 |
T3100 |
T3101 |
compound |
Wnt,reporter |
R868 |
T3178 |
T3175 |
conj |
analysis,hybridization |
R869 |
T3101 |
T3102 |
compound |
reporter,transgene |
R870 |
T3102 |
T3103 |
compound |
transgene,expression |
R871 |
T3103 |
T3099 |
pobj |
expression,by |
R872 |
T3179 |
T3131 |
punct |
.,focused |
R873 |
T3104 |
T3087 |
punct |
.,showed |
R874 |
T3181 |
T3182 |
det |
The,conclusion |
R875 |
T3106 |
T3107 |
nsubj |
Development,remained |
R876 |
T3108 |
T3107 |
punct |
", ",remained |
R877 |
T3182 |
T3184 |
nsubj |
conclusion,is |
R878 |
T3109 |
T3107 |
advmod |
however,remained |
R879 |
T3110 |
T3107 |
punct |
", ",remained |
R880 |
T3111 |
T3112 |
advmod |
surprisingly,normal |
R881 |
T3183 |
T3182 |
amod |
obvious,conclusion |
R882 |
T3112 |
T3107 |
acomp |
normal,remained |
R883 |
T3113 |
T3107 |
punct |
", ",remained |
R884 |
T3114 |
T3107 |
prep |
with,remained |
R885 |
T3185 |
T3186 |
mark |
that,is |
R886 |
T3115 |
T3114 |
pobj |
survival,with |
R887 |
T3116 |
T3115 |
prep |
to,survival |
R888 |
T3117 |
T3116 |
pobj |
birth,to |
R889 |
T3118 |
T3115 |
cc |
and,survival |
R890 |
T3186 |
T3184 |
ccomp |
is,is |
R891 |
T3119 |
T3120 |
amod |
few,defects |
R892 |
T3120 |
T3115 |
conj |
defects,survival |
R893 |
T3121 |
T3120 |
amod |
apparent,defects |
R894 |
T3122 |
T3120 |
prep |
in,defects |
R895 |
T3123 |
T3124 |
amod |
most,systems |
R896 |
T3187 |
T3186 |
prep |
in,is |
R897 |
T3124 |
T3122 |
pobj |
systems,in |
R898 |
T3125 |
T3124 |
compound |
organ,systems |
R899 |
T3126 |
T3107 |
punct |
.,remained |
R900 |
T3128 |
T3129 |
poss |
Our,analysis |
R901 |
T3188 |
T3187 |
pobj |
mammals,in |
R902 |
T3129 |
T3131 |
nsubj |
analysis,focused |
R903 |
T3130 |
T3129 |
amod |
phenotypic,analysis |
R905 |
T3132 |
T3131 |
prep |
on,focused |
R906 |
T3133 |
T3134 |
det |
the,kidney |
R907 |
T3190 |
T3188 |
prep |
unlike,mammals |
R908 |
T3134 |
T3132 |
pobj |
kidney,on |
R909 |
T3135 |
T3134 |
punct |
", ",kidney |
R910 |
T3136 |
T3137 |
dep |
which,showed |
R911 |
T3137 |
T3134 |
relcl |
showed,kidney |
R912 |
T3191 |
T3190 |
pobj |
Drosophila,unlike |
R913 |
T3138 |
T3139 |
amod |
altered,morphogenesis |
R914 |
T3139 |
T3137 |
dobj |
morphogenesis,showed |
R915 |
T3192 |
T3186 |
punct |
", ",is |
R916 |
T3193 |
T3194 |
npadvmod |
Pygopus,mediated |
R917 |
T3194 |
T3196 |
amod |
mediated,signaling |
R918 |
T3195 |
T3194 |
punct |
-,mediated |
R919 |
T3196 |
T3186 |
nsubj |
signaling,is |
R920 |
T3197 |
T3196 |
amod |
canonical,signaling |
R921 |
T3198 |
T3196 |
compound |
Wnt,signaling |
R922 |
T3199 |
T3186 |
neg |
not,is |
R923 |
T3200 |
T3201 |
advmod |
absolutely,necessary |
R924 |
T3201 |
T3186 |
acomp |
necessary,is |
R925 |
T3202 |
T3186 |
prep |
in,is |
R926 |
T3203 |
T3204 |
amod |
most,systems |
R927 |
T3204 |
T3202 |
pobj |
systems,in |
R928 |
T3205 |
T3204 |
amod |
developing,systems |
R929 |
T3206 |
T3204 |
compound |
organ,systems |
R930 |
T3207 |
T3184 |
punct |
.,is |
R904 |
T3189 |
T3188 |
punct |
", ",mammals |