PMC:149350 / 10642-22751 JSONTXT

{"target":"http://pubannotation.org/docs/sourcedb/PMC/sourceid/149350","sourcedb":"PMC","sourceid":"149350","source_url":"http://www.ncbi.nlm.nih.gov/pmc/149350","text":"Results\nEquatorial and axial (sagittal) dimensions of lenses, and gross lenticular appearance, were recorded (Table 1). AlphaA/BKO lenses at 5 wks of age were significantly smaller (p \u003c 0.05), both equatorially and axially, than 5 wk old wild type lenses and remain smaller throughout life. Fifty-four wk old alphaA/BKO lenses were much smaller than similarly aged normal lenses, and were similar in size to 5 wk old wild type lenses. Both the 5 wk old wild type lenses and the 54 wk old alphaA/BKO lenses were significantly smaller (p \u003c 0.01) than the 46 or 72 wk old wild type lenses. AlphaA/BKO lenses at 5 wks of age exhibited vacuoles at the equatorial region (Figure 1A see arrows) and an area of slight light scatter at the posterior subcapsular regions (Figure 1B see arrowheads). By 54 wks of age, alphaA/BKO lenses exhibited dense cortical and nuclear cataracts (Figure 1C). No posterior sutures were observed in any of the alphaA/BKO lenses examined. All wild type lenses were transparent upon removal from the eye (Figure 1D,1E and 1F) and had well defined posterior sutures.\nFigure 1 Light micrographs of lenses dissected from fixed eyes of alphaA/BKO (A-C) and wild type (D-F) mice. All micrographs were generated using a 4 × /0.10 objective on a Zeiss LSM 410 using transmitted light mode. Images in (A\u0026B) were enlarged compared to images (C-F) (see scale bars). For all micrographs except (B) the objective focal point was set at the equatorial plane of the lens, while (B) was set at the posterior pole of the lens. (A\u0026B) Representative micrographs of lenses from 5 wk alphaA/BKO mice, showing changes in the equatorial (A) and posterior sub capsular (B) regions. (C) Representative micrograph of 54 wk alphaA/BKO mouse lens showing dense whole lens cataract. (D-F) Representative clear lenses from 5 wk (D), 46 wk (E), and wk 72 (F) wild type mice. Arrows (A) indicate vacuoles in the equatorial region of a representative 5 wk alphaA/BKO lens. The arrowheads (B) indicate an area of minor light scattering deep to the posterior capsule in a representative 5 wk alphaA/BKO lens.\nTable 1 Dimensions and gross lenticular appearance of lenses.\nLenses Sample size (n) Mean +/- standard deviation Gross lenticular appearance\nAxial diameter mm Equatorial diameter mm\n5 wk alphaA/BKO 6 1.3567 +/-0.0321 1.55+/-0.0458 Equatorial and posterior subcapsular light scattering\n54 wk alphaA/BKO 6 1.78+/-0.0516 1.87+/-0.0837 Dense whole lens cataract\n5 wk wild type 6 1.8625+/-0.0585 1.905+/-0.087 clear\n46 wk wild type 6 2.2075+/-0.0929 2.3775+/-0.015 clear\n72 wk wild type 6 2.5633+/-0.0252 2.4567+/-0.0058 clear Examination of mid-axial sections stained with DiI and SYTOX green revealed unique differences in gross morphology and nucleic acid staining between alphaA/BKO lenses and wild type lenses (Figure 2). In five week old alphaA/BKO lenses, Sytox green stained nuclei (green) in the anterior epithelium, equatorial/bow, and posterior subcapsular regions (Figure 2A). Cell nuclei localized to the equatorial/bow region were disorganized as compared to wild type. There were enlarged extracellular spaces between cells in the equatorial/bow region in 5 wk old alphaA/BKO lenses. Fifty-four wk old alphaA/BKO lenses (Figure 2B) did not stain for nucleic acid in the posterior subcapsular region, however, the entire anterior subcapsular lens region stained for nucleic acid (refer to Figure 3I for high magnification/resolution image of this area). A defined equatorial/bow region was not observed in these older lenses, and fiber cells extending to the posterior capsule of the lens were not found. Disorganized cellular material capped the anterior region of the embryonic/fetal nucleus, and a distinctive indentation of the lens mass at the equatorial region was apparent in mid-axial sections as well as whole lenses. In alphaA/BKO lenses the embryonic/fetal nucleus (primary lens fibers) had migrated posteriorly and appeared to be in contact with the posterior capsule. In 5 wk, 46 wk or 72 wk old wild type lenses (Figure 2C,2D and 2E), however, there was no posterior or anterior subcapsular nucleic acid staining and the embryonic-fetal nucleus was centrally located, not in contact with the posterior capsule.\nFigure 2 Representative confocal optical sections taken from mid-axial vibratome sections of alphaA/BKO (A\u0026B) and wild type (C-E) lenses. All optical sections were generated using a 4 × /0.10 objective on a Zeiss LSM 410. The red color represents DiI staining (membrane), while the green color represents SYTOX green staining (DNA). (A\u0026B) were taken from 5 wk and 54 wk alphaA/BKO lenses, respectively. (C-E) were taken from 5 wk, 46 wk, and 72 wk wild type lenses, respectively. The arrowheads indicate representative anterior epithelial nuclei stained with SYTOX green.\nFigure 3 Representative confocal optical sections from alphaA/BKO (A-J) and wild type (K-T) lenses. All optical sections were generated using a 63 × /1.4 objective on a Zeiss LSM 410. The red color represents DiI staining (membrane), while the green color represents SYTOX green staining (DNA). (A-E) are from 5 wk alphaA/BKO lenses. (F-J) are from 54 wk alphaA/BKO lenses. (K-O) are from 5 wk wild type lenses. (P-T) are from 46 wk wild type lenses. (A, F, K, and P) are anterior central cortex and epithelium. (B, G, L, and Q) are equatorial bow region. (C, H, M, and R) are embryonic/fetal nucleus. (D, I, N, and S) are deep anterior cortex. (E, J, O, and T) are posterior subcapsular region. (*) indicates areas devoid of cellular material (B and G). Arrow indicates posterior suture (O). Arrowheads indicate representative nuclei stained with SYTOX green (A, B, E, F, G, I, K, L, P, and Q). At higher magnification, examination of sections stained with DiI and SYTOX green revealed ultrastructural differences between alphaA/BKO lenses and wild type lenses (Figure 3). Anterior epithelial staining in 5 wk old alphaA/BKO (Figure 3A) and 5 wk old wild type lenses were similar (Figure 3K) with respect to nuclear staining with SYTOX green. However, some differences in 54 wk old alphaA/BKO lenses (Figure 3F) were observed in the central anterior epithelium compared to 5 wk (Figure 3K), 46 wk (Figure 3P), and 72 wk (data not shown) wild type lenses. These differences included changes in central epithelial nuclear staining, nuclear shape and epithelial thickness and continuity. Superficial and deep anterior cortical staining was grossly different between alphaA/BKO (Figure 3A,3D,3F and 3I) and wild type lenses (Figure 3K,3N,3P and 3S). In 5 and 54 wk old alphaA/BKO lenses, superficial and deep anterior cortical regions, stained for lipid membranes, did not reveal any patterns typical of fiber cells cut in cross-section or longitudinal section. In 5 wk alphaA/BKO lenses there was a high density of stained membranes per unit area throughout the cortex, with no nucleic acid staining in these regions. In 54 wk alphaA/BKO lenses, large, irregularly shaped cells were observed, interspersed among regions of high membrane staining density per unit area. These large objects were not vacuoles, because examination of the interior of these structures by transmitted and reflected light microscopy showed that the membranes encompassed cellular material (data not shown). In addition, nucleic acid staining was observed (Figure 3F and 3I) within many of these cells exhibiting large cross-sectional profiles. In the wild type lenses, typical cross-sectional patterns of fiber cells organized in radial columns were observed (Figure 3K,3N,3P and 3S).\nUltrastructural differences at the equatorial/bow region were also observed between alphaA/BKO lenses (Figure 3B and 3G) and wild type lenses (Figure 3L and 3Q). In contrast to wild type lenses (Figure 3L and 3Q), the alphaA/BKO lenses (Figure 3B and 3G) contained large areas devoid of cellular material, their nuclei were not limited to a well defined equatorial/bow region, and ordered radial columns of elongating fiber cells extending from the posterior capsule to the anterior epithelium were not observed. In addition to the equatorial region, ultrastructural differences between alphaA/BKO lenses (Figure 3C and 3H) and wild type lenses (Figure 3M and 3R) were observed in fiber cells of the embryonic and fetal nucleus. In 5 wk alphaA/BKO lenses (Figure 3C), a greater variation in cross sectional diameter was observed, with cell diameters ranging from less than 2 microns up to approximately 25 microns, compared to the 5 wk and 46 wk wild type lenses, whose diameters ranged from less than 2 microns up to approximately 10 microns (Figure 3M and 3R). At 54 wk, these cells in the alphaA/BKO lenses were larger in cross sectional diameter than wild type, and exhibited a much greater variation in the cross sectional size and cell shape.\nCells adjacent to the posterior capsule of 5 wk alphaA/BKO lenses (Figure 3E) exhibited nucleic acid staining, were small and non-elongated, with numerous membrane projections., These were, however, not observed in 54 wk alphaA/BKO lenses. In 5 wk alphaA/BKO lenses, deep to the posterior capsule, larger diameter cells, similar to those observed in the embryonic/fetal nucleus were observed (data not shown). Fiber cells of similar dimension and appearance to embryonic/fetal nuclear fibers were seen at the posterior capsule in 54 wk alphaA/BKO lenses (Figure 3J). In both 5 and 54 wk alphaA/BKO lenses, no posterior sutures could be found in any axial or equatorial vibratome sections examined. Posterior sutures were readily found in vibratome sections of wild type lenses (Figure 3O, arrow). Fiber cells attached to the posterior capsule and extending anteriorly were observed in 5 wk (Figure 3O), 46 wk (Figure 3T) and 72 wk (data not shown) wild type lenses.\nSEM confirmed many of the observations made by confocal microscopy. In the equatorial/bow region, a disorganized array of relatively small irregular shaped cells were observed in 5 wk (Figure 4A) and 54 wk (Figure 4D) alphaA/BKO lenses, while elongated fiber cells organized into radial columns, with ball and socket interdigitations, were observed in 5 wk (Figure 4G), 46 wk (Figure 4J) and 72 wk (data not shown) wild type lenses. In the anterior cortical region of 5 wk alphaA/BKO lenses, radial columns of cells with numerous cell surface projections were observed (Figure 4B), but these radial columns were no longer present at 54 weeks of age (Figure 4E). In these older lenses, irregularly shaped, convoluted cells, with no recognizable pattern of organization, were present. In contrast, radial columns of elongated fiber cells of uniform size and shape, with ball and socket interdigitations, were present in the anterior cortical region of 5 wk (Figure 4H), 46 wk (Figure 4K) and 72 week (data not shown) wild type lenses. The posterior subcapsular regions of 5 wk (Figure 4C) and 54 wk (Figure 4F) alphaA/BKO lenses were filled with irregularly shaped cells with numerous cell surface projections. In the fetal/embryonic nucleus, elongated fiber cells with numerous long, finger-like projections and furrowed membranes were evident (not shown). No elongated cells extending from the bow region to the posterior capsule of alphaA/BKO lenses were observed. In contrast, radial columns of elongated fiber cells of uniform size and shape containing ball and socket interdigitations were observed in the posterior subcapsular region in 5 wk (Figure 4I), 46 wk (Figure 4L) and 72 wk (data not shown) wild type lenses, and fiber cells in contact with the posterior capsule could be traced back to the equatorial bow region and anterior epithelium.\nFigure 4 Representative scanning electron micrographs of alphaA/BKO (A-F) and wild type (G-L) lenses. (A-C) Representative images from 5 wk alphaA/BKO lenses. (D-F) Representative images from 54 wk alphaA/BKO lenses. (G-I) Representative images from 5 wk wild type lenses. (J-L) Representative images from 46 wk wild type lenses. (A, D, G, and J) Equatorial/bow region. (B, E, H, and K) Anterior cortex. 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