Id |
Subject |
Object |
Predicate |
Lexical cue |
T2716 |
11-14 |
DT |
denotes |
The |
T2717 |
15-18 |
NN |
denotes |
use |
T2718 |
39-47 |
VBN |
denotes |
provided |
T2719 |
19-21 |
IN |
denotes |
of |
T2720 |
22-27 |
NN |
denotes |
mouse |
T2721 |
28-34 |
NNS |
denotes |
models |
T2722 |
35-38 |
VBZ |
denotes |
has |
T2723 |
48-56 |
JJ |
denotes |
valuable |
T2724 |
57-64 |
NN |
denotes |
insight |
T2725 |
65-69 |
IN |
denotes |
into |
T2726 |
70-73 |
DT |
denotes |
the |
T2727 |
74-82 |
NN |
denotes |
etiology |
T2728 |
83-85 |
IN |
denotes |
of |
T2729 |
86-95 |
JJ |
denotes |
monogenic |
T2730 |
96-105 |
NNS |
denotes |
syndromes |
T2731 |
106-112 |
VBN |
denotes |
caused |
T2732 |
113-115 |
IN |
denotes |
by |
T2733 |
116-122 |
JJ |
denotes |
single |
T2734 |
128-137 |
NNS |
denotes |
mutations |
T2735 |
123-127 |
NN |
denotes |
gene |
T2736 |
137-138 |
. |
denotes |
. |
T2737 |
138-252 |
sentence |
denotes |
However, such models do not mimic the genetic complexity of disease traits commonly seen in the human population. |
T2738 |
139-146 |
RB |
denotes |
However |
T2739 |
167-172 |
VB |
denotes |
mimic |
T2740 |
146-148 |
, |
denotes |
, |
T2741 |
148-152 |
JJ |
denotes |
such |
T2742 |
153-159 |
NNS |
denotes |
models |
T2743 |
160-162 |
VBP |
denotes |
do |
T2744 |
163-166 |
RB |
denotes |
not |
T2745 |
173-176 |
DT |
denotes |
the |
T2746 |
185-195 |
NN |
denotes |
complexity |
T2747 |
177-184 |
JJ |
denotes |
genetic |
T2748 |
196-198 |
IN |
denotes |
of |
T2749 |
199-206 |
NN |
denotes |
disease |
T2750 |
207-213 |
NNS |
denotes |
traits |
T2751 |
214-222 |
RB |
denotes |
commonly |
T2752 |
223-227 |
VBN |
denotes |
seen |
T2753 |
228-230 |
IN |
denotes |
in |
T2754 |
231-234 |
DT |
denotes |
the |
T2755 |
241-251 |
NN |
denotes |
population |
T2756 |
235-240 |
JJ |
denotes |
human |
T2757 |
251-252 |
. |
denotes |
. |
T2758 |
252-487 |
sentence |
denotes |
Complex traits, such as polygenic growth and obesity are influenced by the small to moderate direct effects of quantitative trait loci (QTL), epistasis between QTL alleles, environmental perturbations and QTL-environment interactions. |
T2759 |
253-260 |
NN |
denotes |
Complex |
T2760 |
261-267 |
NNS |
denotes |
traits |
T2761 |
310-320 |
VBN |
denotes |
influenced |
T2762 |
267-269 |
, |
denotes |
, |
T2763 |
269-273 |
JJ |
denotes |
such |
T2764 |
274-276 |
IN |
denotes |
as |
T2765 |
277-286 |
JJ |
denotes |
polygenic |
T2766 |
287-293 |
NN |
denotes |
growth |
T2767 |
294-297 |
CC |
denotes |
and |
T2768 |
298-305 |
NN |
denotes |
obesity |
T2769 |
306-309 |
VBP |
denotes |
are |
T2770 |
321-323 |
IN |
denotes |
by |
T2771 |
324-327 |
DT |
denotes |
the |
T2772 |
353-360 |
NNS |
denotes |
effects |
T2773 |
328-333 |
JJ |
denotes |
small |
T2774 |
334-336 |
IN |
denotes |
to |
T2775 |
337-345 |
JJ |
denotes |
moderate |
T2776 |
346-352 |
JJ |
denotes |
direct |
T2777 |
361-363 |
IN |
denotes |
of |
T2778 |
364-376 |
JJ |
denotes |
quantitative |
T2779 |
383-387 |
NNS |
denotes |
loci |
T2780 |
377-382 |
NN |
denotes |
trait |
T2781 |
388-389 |
-LRB- |
denotes |
( |
T2782 |
389-392 |
NN |
denotes |
QTL |
T2783 |
392-393 |
-RRB- |
denotes |
) |
T2784 |
393-395 |
, |
denotes |
, |
T2785 |
395-404 |
NN |
denotes |
epistasis |
T2786 |
405-412 |
IN |
denotes |
between |
T2787 |
413-416 |
NN |
denotes |
QTL |
T2788 |
417-424 |
NNS |
denotes |
alleles |
T2789 |
424-426 |
, |
denotes |
, |
T2790 |
426-439 |
JJ |
denotes |
environmental |
T2791 |
440-453 |
NNS |
denotes |
perturbations |
T2792 |
454-457 |
CC |
denotes |
and |
T2793 |
458-461 |
NN |
denotes |
QTL |
T2794 |
462-473 |
NN |
denotes |
environment |
T2795 |
461-462 |
HYPH |
denotes |
- |
T2796 |
474-486 |
NNS |
denotes |
interactions |
T2797 |
486-487 |
. |
denotes |
. |
T2798 |
487-670 |
sentence |
denotes |
To date numerous mouse growth and obesity QTL have been localized [1,2], however, little progress has been made in determining the specific nature of the underlying genetic variants. |
T2799 |
488-490 |
IN |
denotes |
To |
T2800 |
544-553 |
VBN |
denotes |
localized |
T2801 |
491-495 |
NN |
denotes |
date |
T2802 |
496-504 |
JJ |
denotes |
numerous |
T2803 |
530-533 |
NN |
denotes |
QTL |
T2804 |
505-510 |
NN |
denotes |
mouse |
T2805 |
511-517 |
NN |
denotes |
growth |
T2806 |
518-521 |
CC |
denotes |
and |
T2807 |
522-529 |
NN |
denotes |
obesity |
T2808 |
534-538 |
VBP |
denotes |
have |
T2809 |
539-543 |
VBN |
denotes |
been |
T2810 |
595-599 |
VBN |
denotes |
made |
T2811 |
554-555 |
-LRB- |
denotes |
[ |
T2812 |
557-558 |
CD |
denotes |
2 |
T2813 |
555-556 |
CD |
denotes |
1 |
T2814 |
556-557 |
, |
denotes |
, |
T2815 |
558-559 |
-RRB- |
denotes |
] |
T2816 |
559-561 |
, |
denotes |
, |
T2817 |
561-568 |
RB |
denotes |
however |
T2818 |
568-570 |
, |
denotes |
, |
T2819 |
570-576 |
JJ |
denotes |
little |
T2820 |
577-585 |
NN |
denotes |
progress |
T2821 |
586-589 |
VBZ |
denotes |
has |
T2822 |
590-594 |
VBN |
denotes |
been |
T2823 |
600-602 |
IN |
denotes |
in |
T2824 |
603-614 |
VBG |
denotes |
determining |
T2825 |
615-618 |
DT |
denotes |
the |
T2826 |
628-634 |
NN |
denotes |
nature |
T2827 |
619-627 |
JJ |
denotes |
specific |
T2828 |
635-637 |
IN |
denotes |
of |
T2829 |
638-641 |
DT |
denotes |
the |
T2830 |
661-669 |
NNS |
denotes |
variants |
T2831 |
642-652 |
VBG |
denotes |
underlying |
T2832 |
653-660 |
JJ |
denotes |
genetic |
T2833 |
669-670 |
. |
denotes |
. |
T2834 |
670-819 |
sentence |
denotes |
One resource used to fine map QTL are congenic strains which are designed to convert a complex polygenic trait into one that is mono- or oligogenic. |
T2835 |
671-674 |
CD |
denotes |
One |
T2836 |
675-683 |
NN |
denotes |
resource |
T2837 |
705-708 |
VBP |
denotes |
are |
T2838 |
684-688 |
VBN |
denotes |
used |
T2839 |
689-691 |
TO |
denotes |
to |
T2840 |
697-700 |
VB |
denotes |
map |
T2841 |
692-696 |
RB |
denotes |
fine |
T2842 |
701-704 |
NN |
denotes |
QTL |
T2843 |
709-717 |
JJ |
denotes |
congenic |
T2844 |
718-725 |
NNS |
denotes |
strains |
T2845 |
726-731 |
WDT |
denotes |
which |
T2846 |
736-744 |
VBN |
denotes |
designed |
T2847 |
732-735 |
VBP |
denotes |
are |
T2848 |
745-747 |
TO |
denotes |
to |
T2849 |
748-755 |
VB |
denotes |
convert |
T2850 |
756-757 |
DT |
denotes |
a |
T2851 |
776-781 |
NN |
denotes |
trait |
T2852 |
758-765 |
JJ |
denotes |
complex |
T2853 |
766-775 |
JJ |
denotes |
polygenic |
T2854 |
782-786 |
IN |
denotes |
into |
T2855 |
787-790 |
CD |
denotes |
one |
T2856 |
791-795 |
WDT |
denotes |
that |
T2857 |
796-798 |
VBZ |
denotes |
is |
T2858 |
799-803 |
AFX |
denotes |
mono |
T2859 |
808-818 |
JJ |
denotes |
oligogenic |
T2860 |
803-804 |
HYPH |
denotes |
- |
T2861 |
805-807 |
CC |
denotes |
or |
T2862 |
818-819 |
. |
denotes |
. |
T2863 |
819-1039 |
sentence |
denotes |
This is accomplished by eliminating segregating genetic variation outside the locus of interest and reducing the environmental variation influencing a trait by characterizing large numbers of genetically identical mice. |
T2864 |
820-824 |
DT |
denotes |
This |
T2865 |
828-840 |
VBN |
denotes |
accomplished |
T2866 |
825-827 |
VBZ |
denotes |
is |
T2867 |
841-843 |
IN |
denotes |
by |
T2868 |
844-855 |
VBG |
denotes |
eliminating |
T2869 |
856-867 |
VBG |
denotes |
segregating |
T2870 |
868-875 |
JJ |
denotes |
genetic |
T2871 |
876-885 |
NN |
denotes |
variation |
T2872 |
886-893 |
IN |
denotes |
outside |
T2873 |
894-897 |
DT |
denotes |
the |
T2874 |
898-903 |
NN |
denotes |
locus |
T2875 |
904-906 |
IN |
denotes |
of |
T2876 |
907-915 |
NN |
denotes |
interest |
T2877 |
916-919 |
CC |
denotes |
and |
T2878 |
920-928 |
VBG |
denotes |
reducing |
T2879 |
929-932 |
DT |
denotes |
the |
T2880 |
947-956 |
NN |
denotes |
variation |
T2881 |
933-946 |
JJ |
denotes |
environmental |
T2882 |
957-968 |
VBG |
denotes |
influencing |
T2883 |
969-970 |
DT |
denotes |
a |
T2884 |
971-976 |
NN |
denotes |
trait |
T2885 |
977-979 |
IN |
denotes |
by |
T2886 |
980-994 |
VBG |
denotes |
characterizing |
T2887 |
995-1000 |
JJ |
denotes |
large |
T2888 |
1001-1008 |
NNS |
denotes |
numbers |
T2889 |
1009-1011 |
IN |
denotes |
of |
T2890 |
1012-1023 |
RB |
denotes |
genetically |
T2891 |
1024-1033 |
JJ |
denotes |
identical |
T2892 |
1034-1038 |
NNS |
denotes |
mice |
T2893 |
1038-1039 |
. |
denotes |
. |
T2894 |
1039-1168 |
sentence |
denotes |
The Complex Trait Consortium (CTC) considers congenic analysis an excellent method to confirm and subsequently fine map QTL [3]. |
T2895 |
1040-1043 |
DT |
denotes |
The |
T2896 |
1058-1068 |
NNP |
denotes |
Consortium |
T2897 |
1044-1051 |
NNP |
denotes |
Complex |
T2898 |
1052-1057 |
NNP |
denotes |
Trait |
T2899 |
1075-1084 |
VBZ |
denotes |
considers |
T2900 |
1069-1070 |
-LRB- |
denotes |
( |
T2901 |
1070-1073 |
NNP |
denotes |
CTC |
T2902 |
1073-1074 |
-RRB- |
denotes |
) |
T2903 |
1085-1093 |
JJ |
denotes |
congenic |
T2904 |
1094-1102 |
NN |
denotes |
analysis |
T2905 |
1116-1122 |
NN |
denotes |
method |
T2906 |
1103-1105 |
DT |
denotes |
an |
T2907 |
1106-1115 |
JJ |
denotes |
excellent |
T2908 |
1123-1125 |
TO |
denotes |
to |
T2909 |
1126-1133 |
VB |
denotes |
confirm |
T2910 |
1134-1137 |
CC |
denotes |
and |
T2911 |
1138-1150 |
RB |
denotes |
subsequently |
T2912 |
1156-1159 |
VB |
denotes |
map |
T2913 |
1151-1155 |
RB |
denotes |
fine |
T2914 |
1160-1163 |
NN |
denotes |
QTL |
T2915 |
1164-1165 |
-LRB- |
denotes |
[ |
T2916 |
1165-1166 |
CD |
denotes |
3 |
T2917 |
1166-1167 |
-RRB- |
denotes |
] |
T2918 |
1167-1168 |
. |
denotes |
. |
T2919 |
1168-1385 |
sentence |
denotes |
Traditionally, congenic strains are developed by introgressing QTL alleles from a donor strain, whose boundaries are defined by genetic markers, on the genetic background of a recipient strain via 10 backcrosses [4]. |
T2920 |
1169-1182 |
RB |
denotes |
Traditionally |
T2921 |
1205-1214 |
VBN |
denotes |
developed |
T2922 |
1182-1184 |
, |
denotes |
, |
T2923 |
1184-1192 |
JJ |
denotes |
congenic |
T2924 |
1193-1200 |
NNS |
denotes |
strains |
T2925 |
1201-1204 |
VBP |
denotes |
are |
T2926 |
1215-1217 |
IN |
denotes |
by |
T2927 |
1218-1231 |
VBG |
denotes |
introgressing |
T2928 |
1232-1235 |
NN |
denotes |
QTL |
T2929 |
1236-1243 |
NNS |
denotes |
alleles |
T2930 |
1244-1248 |
IN |
denotes |
from |
T2931 |
1249-1250 |
DT |
denotes |
a |
T2932 |
1257-1263 |
NN |
denotes |
strain |
T2933 |
1251-1256 |
NN |
denotes |
donor |
T2934 |
1263-1265 |
, |
denotes |
, |
T2935 |
1265-1270 |
WP$ |
denotes |
whose |
T2936 |
1271-1281 |
NNS |
denotes |
boundaries |
T2937 |
1286-1293 |
VBN |
denotes |
defined |
T2938 |
1282-1285 |
VBP |
denotes |
are |
T2939 |
1294-1296 |
IN |
denotes |
by |
T2940 |
1297-1304 |
JJ |
denotes |
genetic |
T2941 |
1305-1312 |
NNS |
denotes |
markers |
T2942 |
1312-1314 |
, |
denotes |
, |
T2943 |
1314-1316 |
IN |
denotes |
on |
T2944 |
1317-1320 |
DT |
denotes |
the |
T2945 |
1329-1339 |
NN |
denotes |
background |
T2946 |
1321-1328 |
JJ |
denotes |
genetic |
T2947 |
1340-1342 |
IN |
denotes |
of |
T2948 |
1343-1344 |
DT |
denotes |
a |
T2949 |
1355-1361 |
NN |
denotes |
strain |
T2950 |
1345-1354 |
NN |
denotes |
recipient |
T2951 |
1362-1365 |
IN |
denotes |
via |
T2952 |
1366-1368 |
CD |
denotes |
10 |
T2953 |
1369-1380 |
NNS |
denotes |
backcrosses |
T2954 |
1381-1382 |
-LRB- |
denotes |
[ |
T2955 |
1382-1383 |
CD |
denotes |
4 |
T2956 |
1383-1384 |
-RRB- |
denotes |
] |
T2957 |
1384-1385 |
. |
denotes |
. |
T2958 |
1385-1507 |
sentence |
denotes |
While technically straightforward this is a time consuming endeavor taking over three years to construct a single strain. |
T2959 |
1386-1391 |
IN |
denotes |
While |
T2960 |
1404-1419 |
JJ |
denotes |
straightforward |
T2961 |
1392-1403 |
RB |
denotes |
technically |
T2962 |
1425-1427 |
VBZ |
denotes |
is |
T2963 |
1420-1424 |
NN |
denotes |
this |
T2964 |
1428-1429 |
DT |
denotes |
a |
T2965 |
1445-1453 |
NN |
denotes |
endeavor |
T2966 |
1430-1434 |
NN |
denotes |
time |
T2967 |
1435-1444 |
VBG |
denotes |
consuming |
T2968 |
1454-1460 |
VBG |
denotes |
taking |
T2969 |
1461-1465 |
IN |
denotes |
over |
T2970 |
1466-1471 |
CD |
denotes |
three |
T2971 |
1472-1477 |
NNS |
denotes |
years |
T2972 |
1478-1480 |
TO |
denotes |
to |
T2973 |
1481-1490 |
VB |
denotes |
construct |
T2974 |
1491-1492 |
DT |
denotes |
a |
T2975 |
1500-1506 |
NN |
denotes |
strain |
T2976 |
1493-1499 |
JJ |
denotes |
single |
T2977 |
1506-1507 |
. |
denotes |
. |
T2978 |
1507-1650 |
sentence |
denotes |
The speed congenic approach is an alternative to this lengthy process and can reduce the number of required backcrosses from 10 to five [5,6]. |
T2979 |
1508-1511 |
DT |
denotes |
The |
T2980 |
1527-1535 |
NN |
denotes |
approach |
T2981 |
1512-1517 |
NN |
denotes |
speed |
T2982 |
1518-1526 |
JJ |
denotes |
congenic |
T2983 |
1536-1538 |
VBZ |
denotes |
is |
T2984 |
1539-1541 |
DT |
denotes |
an |
T2985 |
1542-1553 |
NN |
denotes |
alternative |
T2986 |
1554-1556 |
IN |
denotes |
to |
T2987 |
1557-1561 |
DT |
denotes |
this |
T2988 |
1570-1577 |
NN |
denotes |
process |
T2989 |
1562-1569 |
JJ |
denotes |
lengthy |
T2990 |
1578-1581 |
CC |
denotes |
and |
T2991 |
1582-1585 |
MD |
denotes |
can |
T2992 |
1586-1592 |
VB |
denotes |
reduce |
T2993 |
1593-1596 |
DT |
denotes |
the |
T2994 |
1597-1603 |
NN |
denotes |
number |
T2995 |
1604-1606 |
IN |
denotes |
of |
T2996 |
1607-1615 |
VBN |
denotes |
required |
T2997 |
1616-1627 |
NNS |
denotes |
backcrosses |
T2998 |
1628-1632 |
IN |
denotes |
from |
T2999 |
1633-1635 |
CD |
denotes |
10 |
T3000 |
1636-1638 |
IN |
denotes |
to |
T3001 |
1639-1643 |
CD |
denotes |
five |
T3002 |
1644-1645 |
-LRB- |
denotes |
[ |
T3003 |
1647-1648 |
CD |
denotes |
6 |
T3004 |
1645-1646 |
CD |
denotes |
5 |
T3005 |
1646-1647 |
, |
denotes |
, |
T3006 |
1648-1649 |
-RRB- |
denotes |
] |
T3007 |
1649-1650 |
. |
denotes |
. |
T3008 |
1650-1798 |
sentence |
denotes |
This strategy uses marker-assisted selection to identify male mice inheriting fewer donor alleles, than expected on average, during each backcross. |
T3009 |
1651-1655 |
DT |
denotes |
This |
T3010 |
1656-1664 |
NN |
denotes |
strategy |
T3011 |
1665-1669 |
VBZ |
denotes |
uses |
T3012 |
1670-1676 |
NN |
denotes |
marker |
T3013 |
1677-1685 |
VBN |
denotes |
assisted |
T3014 |
1676-1677 |
HYPH |
denotes |
- |
T3015 |
1686-1695 |
NN |
denotes |
selection |
T3016 |
1696-1698 |
TO |
denotes |
to |
T3017 |
1699-1707 |
VB |
denotes |
identify |
T3018 |
1708-1712 |
JJ |
denotes |
male |
T3019 |
1713-1717 |
NNS |
denotes |
mice |
T3020 |
1718-1728 |
VBG |
denotes |
inheriting |
T3021 |
1729-1734 |
JJR |
denotes |
fewer |
T3022 |
1741-1748 |
NNS |
denotes |
alleles |
T3023 |
1735-1740 |
NN |
denotes |
donor |
T3024 |
1748-1750 |
, |
denotes |
, |
T3025 |
1750-1754 |
IN |
denotes |
than |
T3026 |
1755-1763 |
VBN |
denotes |
expected |
T3027 |
1764-1766 |
IN |
denotes |
on |
T3028 |
1767-1774 |
JJ |
denotes |
average |
T3029 |
1774-1776 |
, |
denotes |
, |
T3030 |
1776-1782 |
IN |
denotes |
during |
T3031 |
1783-1787 |
DT |
denotes |
each |
T3032 |
1788-1797 |
NN |
denotes |
backcross |
T3033 |
1797-1798 |
. |
denotes |
. |
T3034 |
1798-1965 |
sentence |
denotes |
Numerous traditionally developed and speed congenics have been used to successfully isolate mouse QTL for a wide array of traits, including growth and obesity [7-13]. |
T3035 |
1799-1807 |
JJ |
denotes |
Numerous |
T3036 |
1842-1851 |
NNS |
denotes |
congenics |
T3037 |
1808-1821 |
RB |
denotes |
traditionally |
T3038 |
1822-1831 |
VBN |
denotes |
developed |
T3039 |
1832-1835 |
CC |
denotes |
and |
T3040 |
1836-1841 |
NN |
denotes |
speed |
T3041 |
1862-1866 |
VBN |
denotes |
used |
T3042 |
1852-1856 |
VBP |
denotes |
have |
T3043 |
1857-1861 |
VBN |
denotes |
been |
T3044 |
1867-1869 |
TO |
denotes |
to |
T3045 |
1883-1890 |
VB |
denotes |
isolate |
T3046 |
1870-1882 |
RB |
denotes |
successfully |
T3047 |
1891-1896 |
NN |
denotes |
mouse |
T3048 |
1897-1900 |
NN |
denotes |
QTL |
T3049 |
1901-1904 |
IN |
denotes |
for |
T3050 |
1905-1906 |
DT |
denotes |
a |
T3051 |
1912-1917 |
NN |
denotes |
array |
T3052 |
1907-1911 |
JJ |
denotes |
wide |
T3053 |
1918-1920 |
IN |
denotes |
of |
T3054 |
1921-1927 |
NNS |
denotes |
traits |
T3055 |
1927-1929 |
, |
denotes |
, |
T3056 |
1929-1938 |
VBG |
denotes |
including |
T3057 |
1939-1945 |
NN |
denotes |
growth |
T3058 |
1946-1949 |
CC |
denotes |
and |
T3059 |
1950-1957 |
NN |
denotes |
obesity |
T3060 |
1958-1959 |
-LRB- |
denotes |
[ |
T3061 |
1959-1960 |
CD |
denotes |
7 |
T3062 |
1960-1961 |
SYM |
denotes |
- |
T3063 |
1961-1963 |
CD |
denotes |
13 |
T3064 |
1963-1964 |
-RRB- |
denotes |
] |
T3065 |
1964-1965 |
. |
denotes |
. |
T3066 |
1965-2169 |
sentence |
denotes |
The C57Bl/6J-hg/hg (HG) mouse is a model of systemic overgrowth resulting from a spontaneous deletion on MMU10, which eliminates expression of the Socs2 (suppressor of cytokine signaling 2) gene [14-16]. |
T3067 |
1966-1969 |
DT |
denotes |
The |
T3068 |
1990-1995 |
NN |
denotes |
mouse |
T3069 |
1970-1975 |
NN |
denotes |
C57Bl |
T3070 |
1982-1984 |
NN |
denotes |
hg |
T3071 |
1975-1976 |
HYPH |
denotes |
/ |
T3072 |
1976-1978 |
NN |
denotes |
6J |
T3073 |
1978-1979 |
HYPH |
denotes |
- |
T3074 |
1979-1981 |
NN |
denotes |
hg |
T3075 |
1981-1982 |
HYPH |
denotes |
/ |
T3076 |
1985-1986 |
-LRB- |
denotes |
( |
T3077 |
1986-1988 |
NN |
denotes |
HG |
T3078 |
1988-1989 |
-RRB- |
denotes |
) |
T3079 |
1996-1998 |
VBZ |
denotes |
is |
T3080 |
1999-2000 |
DT |
denotes |
a |
T3081 |
2001-2006 |
NN |
denotes |
model |
T3082 |
2007-2009 |
IN |
denotes |
of |
T3083 |
2010-2018 |
JJ |
denotes |
systemic |
T3084 |
2019-2029 |
NN |
denotes |
overgrowth |
T3085 |
2030-2039 |
VBG |
denotes |
resulting |
T3086 |
2040-2044 |
IN |
denotes |
from |
T3087 |
2045-2046 |
DT |
denotes |
a |
T3088 |
2059-2067 |
NN |
denotes |
deletion |
T3089 |
2047-2058 |
JJ |
denotes |
spontaneous |
T3090 |
2068-2070 |
IN |
denotes |
on |
T3091 |
2071-2076 |
NN |
denotes |
MMU10 |
T3092 |
2076-2078 |
, |
denotes |
, |
T3093 |
2078-2083 |
WDT |
denotes |
which |
T3094 |
2084-2094 |
VBZ |
denotes |
eliminates |
T3095 |
2095-2105 |
NN |
denotes |
expression |
T3096 |
2106-2108 |
IN |
denotes |
of |
T3097 |
2109-2112 |
DT |
denotes |
the |
T3098 |
2156-2160 |
NN |
denotes |
gene |
T3099 |
2113-2118 |
NN |
denotes |
Socs2 |
T3100 |
2119-2120 |
-LRB- |
denotes |
( |
T3101 |
2120-2130 |
NN |
denotes |
suppressor |
T3102 |
2131-2133 |
IN |
denotes |
of |
T3103 |
2134-2142 |
NN |
denotes |
cytokine |
T3104 |
2143-2152 |
NN |
denotes |
signaling |
T3105 |
2153-2154 |
CD |
denotes |
2 |
T3106 |
2154-2155 |
-RRB- |
denotes |
) |
T3107 |
2161-2162 |
-LRB- |
denotes |
[ |
T3108 |
2162-2164 |
CD |
denotes |
14 |
T3109 |
2164-2165 |
SYM |
denotes |
- |
T3110 |
2165-2167 |
CD |
denotes |
16 |
T3111 |
2167-2168 |
-RRB- |
denotes |
] |
T3112 |
2168-2169 |
. |
denotes |
. |
T3113 |
2169-2357 |
sentence |
denotes |
The role of Socs2 in the HG phenotype was confirmed by an independently engineered Socs2-/- knockout mouse which shared a number of phenotypes in common with HG, including gigantism [17]. |
T3114 |
2170-2173 |
DT |
denotes |
The |
T3115 |
2174-2178 |
NN |
denotes |
role |
T3116 |
2212-2221 |
VBN |
denotes |
confirmed |
T3117 |
2179-2181 |
IN |
denotes |
of |
T3118 |
2182-2187 |
NN |
denotes |
Socs2 |
T3119 |
2188-2190 |
IN |
denotes |
in |
T3120 |
2191-2194 |
DT |
denotes |
the |
T3121 |
2198-2207 |
NN |
denotes |
phenotype |
T3122 |
2195-2197 |
NN |
denotes |
HG |
T3123 |
2208-2211 |
VBD |
denotes |
was |
T3124 |
2222-2224 |
IN |
denotes |
by |
T3125 |
2225-2227 |
DT |
denotes |
an |
T3126 |
2271-2276 |
NN |
denotes |
mouse |
T3127 |
2228-2241 |
RB |
denotes |
independently |
T3128 |
2242-2252 |
VBN |
denotes |
engineered |
T3129 |
2253-2258 |
NN |
denotes |
Socs2 |
T3130 |
2262-2270 |
NN |
denotes |
knockout |
T3131 |
2258-2259 |
SYM |
denotes |
- |
T3132 |
2259-2260 |
HYPH |
denotes |
/ |
T3133 |
2260-2261 |
SYM |
denotes |
- |
T3134 |
2277-2282 |
WDT |
denotes |
which |
T3135 |
2283-2289 |
VBD |
denotes |
shared |
T3136 |
2290-2291 |
DT |
denotes |
a |
T3137 |
2292-2298 |
NN |
denotes |
number |
T3138 |
2299-2301 |
IN |
denotes |
of |
T3139 |
2302-2312 |
NNS |
denotes |
phenotypes |
T3140 |
2313-2315 |
IN |
denotes |
in |
T3141 |
2316-2322 |
NN |
denotes |
common |
T3142 |
2323-2327 |
IN |
denotes |
with |
T3143 |
2328-2330 |
NN |
denotes |
HG |
T3144 |
2330-2332 |
, |
denotes |
, |
T3145 |
2332-2341 |
VBG |
denotes |
including |
T3146 |
2342-2351 |
NN |
denotes |
gigantism |
T3147 |
2352-2353 |
-LRB- |
denotes |
[ |
T3148 |
2353-2355 |
CD |
denotes |
17 |
T3149 |
2355-2356 |
-RRB- |
denotes |
] |
T3150 |
2356-2357 |
. |
denotes |
. |
T3151 |
2357-2451 |
sentence |
denotes |
QTL which alter the phenotypic effects of another locus are referred to as modifier QTL [18]. |
T3152 |
2358-2361 |
NN |
denotes |
QTL |
T3153 |
2418-2426 |
VBN |
denotes |
referred |
T3154 |
2362-2367 |
WDT |
denotes |
which |
T3155 |
2368-2373 |
VBP |
denotes |
alter |
T3156 |
2374-2377 |
DT |
denotes |
the |
T3157 |
2389-2396 |
NNS |
denotes |
effects |
T3158 |
2378-2388 |
JJ |
denotes |
phenotypic |
T3159 |
2397-2399 |
IN |
denotes |
of |
T3160 |
2400-2407 |
DT |
denotes |
another |
T3161 |
2408-2413 |
NN |
denotes |
locus |
T3162 |
2414-2417 |
VBP |
denotes |
are |
T3163 |
2427-2429 |
IN |
denotes |
to |
T3164 |
2430-2432 |
IN |
denotes |
as |
T3165 |
2433-2441 |
NN |
denotes |
modifier |
T3166 |
2442-2445 |
NN |
denotes |
QTL |
T3167 |
2446-2447 |
-LRB- |
denotes |
[ |
T3168 |
2447-2449 |
CD |
denotes |
18 |
T3169 |
2449-2450 |
-RRB- |
denotes |
] |
T3170 |
2450-2451 |
. |
denotes |
. |
T3171 |
2451-2678 |
sentence |
denotes |
Modifier QTL have been mapped for numerous traits and in these studies the modified locus is typically a known gene containing a spontaneously arisen or engineered mutation with major phenotypic effects, such as hg [11,19-21]. |
T3172 |
2452-2460 |
NN |
denotes |
Modifier |
T3173 |
2461-2464 |
NN |
denotes |
QTL |
T3174 |
2475-2481 |
VBN |
denotes |
mapped |
T3175 |
2465-2469 |
VBP |
denotes |
have |
T3176 |
2470-2474 |
VBN |
denotes |
been |
T3177 |
2482-2485 |
IN |
denotes |
for |
T3178 |
2486-2494 |
JJ |
denotes |
numerous |
T3179 |
2495-2501 |
NNS |
denotes |
traits |
T3180 |
2502-2505 |
CC |
denotes |
and |
T3181 |
2506-2508 |
IN |
denotes |
in |
T3182 |
2542-2544 |
VBZ |
denotes |
is |
T3183 |
2509-2514 |
DT |
denotes |
these |
T3184 |
2515-2522 |
NNS |
denotes |
studies |
T3185 |
2523-2526 |
DT |
denotes |
the |
T3186 |
2536-2541 |
NN |
denotes |
locus |
T3187 |
2527-2535 |
VBN |
denotes |
modified |
T3188 |
2545-2554 |
RB |
denotes |
typically |
T3189 |
2555-2556 |
DT |
denotes |
a |
T3190 |
2563-2567 |
NN |
denotes |
gene |
T3191 |
2557-2562 |
VBN |
denotes |
known |
T3192 |
2568-2578 |
VBG |
denotes |
containing |
T3193 |
2579-2580 |
DT |
denotes |
a |
T3194 |
2616-2624 |
NN |
denotes |
mutation |
T3195 |
2581-2594 |
RB |
denotes |
spontaneously |
T3196 |
2595-2601 |
VBN |
denotes |
arisen |
T3197 |
2602-2604 |
CC |
denotes |
or |
T3198 |
2605-2615 |
VBN |
denotes |
engineered |
T3199 |
2625-2629 |
IN |
denotes |
with |
T3200 |
2630-2635 |
JJ |
denotes |
major |
T3201 |
2647-2654 |
NNS |
denotes |
effects |
T3202 |
2636-2646 |
JJ |
denotes |
phenotypic |
T3203 |
2654-2656 |
, |
denotes |
, |
T3204 |
2656-2660 |
JJ |
denotes |
such |
T3205 |
2661-2663 |
IN |
denotes |
as |
T3206 |
2664-2666 |
NN |
denotes |
hg |
T3207 |
2667-2668 |
-LRB- |
denotes |
[ |
T3208 |
2668-2670 |
CD |
denotes |
11 |
T3209 |
2670-2671 |
, |
denotes |
, |
T3210 |
2671-2673 |
CD |
denotes |
19 |
T3211 |
2673-2674 |
SYM |
denotes |
- |
T3212 |
2674-2676 |
CD |
denotes |
21 |
T3213 |
2676-2677 |
-RRB- |
denotes |
] |
T3214 |
2677-2678 |
. |
denotes |
. |
T3215 |
2678-2832 |
sentence |
denotes |
Epistasis forms the basis of these interactions, implying that the known gene and its modifiers are members of the same biochemical or metabolic pathway. |
T3216 |
2679-2688 |
NN |
denotes |
Epistasis |
T3217 |
2689-2694 |
VBZ |
denotes |
forms |
T3218 |
2695-2698 |
DT |
denotes |
the |
T3219 |
2699-2704 |
NN |
denotes |
basis |
T3220 |
2705-2707 |
IN |
denotes |
of |
T3221 |
2708-2713 |
DT |
denotes |
these |
T3222 |
2714-2726 |
NNS |
denotes |
interactions |
T3223 |
2726-2728 |
, |
denotes |
, |
T3224 |
2728-2736 |
VBG |
denotes |
implying |
T3225 |
2737-2741 |
IN |
denotes |
that |
T3226 |
2775-2778 |
VBP |
denotes |
are |
T3227 |
2742-2745 |
DT |
denotes |
the |
T3228 |
2752-2756 |
NN |
denotes |
gene |
T3229 |
2746-2751 |
VBN |
denotes |
known |
T3230 |
2757-2760 |
CC |
denotes |
and |
T3231 |
2761-2764 |
PRP$ |
denotes |
its |
T3232 |
2765-2774 |
NNS |
denotes |
modifiers |
T3233 |
2779-2786 |
NNS |
denotes |
members |
T3234 |
2787-2789 |
IN |
denotes |
of |
T3235 |
2790-2793 |
DT |
denotes |
the |
T3236 |
2824-2831 |
NN |
denotes |
pathway |
T3237 |
2794-2798 |
JJ |
denotes |
same |
T3238 |
2799-2810 |
JJ |
denotes |
biochemical |
T3239 |
2811-2813 |
CC |
denotes |
or |
T3240 |
2814-2823 |
JJ |
denotes |
metabolic |
T3241 |
2831-2832 |
. |
denotes |
. |
T3242 |
2832-2932 |
sentence |
denotes |
Different genetic backgrounds have been shown to modify the growth-enhancing effects of hg [15,22]. |
T3243 |
2833-2842 |
JJ |
denotes |
Different |
T3244 |
2851-2862 |
NNS |
denotes |
backgrounds |
T3245 |
2843-2850 |
JJ |
denotes |
genetic |
T3246 |
2873-2878 |
VBN |
denotes |
shown |
T3247 |
2863-2867 |
VBP |
denotes |
have |
T3248 |
2868-2872 |
VBN |
denotes |
been |
T3249 |
2879-2881 |
TO |
denotes |
to |
T3250 |
2882-2888 |
VB |
denotes |
modify |
T3251 |
2889-2892 |
DT |
denotes |
the |
T3252 |
2910-2917 |
NNS |
denotes |
effects |
T3253 |
2893-2899 |
NN |
denotes |
growth |
T3254 |
2900-2909 |
VBG |
denotes |
enhancing |
T3255 |
2899-2900 |
HYPH |
denotes |
- |
T3256 |
2918-2920 |
IN |
denotes |
of |
T3257 |
2921-2923 |
NN |
denotes |
hg |
T3258 |
2924-2925 |
-LRB- |
denotes |
[ |
T3259 |
2928-2930 |
CD |
denotes |
22 |
T3260 |
2925-2927 |
CD |
denotes |
15 |
T3261 |
2927-2928 |
, |
denotes |
, |
T3262 |
2930-2931 |
-RRB- |
denotes |
] |
T3263 |
2931-2932 |
. |
denotes |
. |
T3264 |
2932-3159 |
sentence |
denotes |
Since the primary function of Socs2 is to negatively regulate growth hormone (Gh) [23], it is likely these background effects are the result of polymorphism influencing interactions between members of the Gh signaling pathway. |
T3265 |
2933-2938 |
IN |
denotes |
Since |
T3266 |
2969-2971 |
VBZ |
denotes |
is |
T3267 |
2939-2942 |
DT |
denotes |
the |
T3268 |
2951-2959 |
NN |
denotes |
function |
T3269 |
2943-2950 |
JJ |
denotes |
primary |
T3270 |
2960-2962 |
IN |
denotes |
of |
T3271 |
2963-2968 |
NN |
denotes |
Socs2 |
T3272 |
3024-3026 |
VBZ |
denotes |
is |
T3273 |
2972-2974 |
TO |
denotes |
to |
T3274 |
2986-2994 |
VB |
denotes |
regulate |
T3275 |
2975-2985 |
RB |
denotes |
negatively |
T3276 |
2995-3001 |
NN |
denotes |
growth |
T3277 |
3002-3009 |
NN |
denotes |
hormone |
T3278 |
3010-3011 |
-LRB- |
denotes |
( |
T3279 |
3011-3013 |
NN |
denotes |
Gh |
T3280 |
3013-3014 |
-RRB- |
denotes |
) |
T3281 |
3015-3016 |
-LRB- |
denotes |
[ |
T3282 |
3016-3018 |
CD |
denotes |
23 |
T3283 |
3018-3019 |
-RRB- |
denotes |
] |
T3284 |
3019-3021 |
, |
denotes |
, |
T3285 |
3021-3023 |
PRP |
denotes |
it |
T3286 |
3027-3033 |
JJ |
denotes |
likely |
T3287 |
3034-3039 |
DT |
denotes |
these |
T3288 |
3051-3058 |
NNS |
denotes |
effects |
T3289 |
3040-3050 |
NN |
denotes |
background |
T3290 |
3059-3062 |
VBP |
denotes |
are |
T3291 |
3063-3066 |
DT |
denotes |
the |
T3292 |
3067-3073 |
NN |
denotes |
result |
T3293 |
3074-3076 |
IN |
denotes |
of |
T3294 |
3077-3089 |
NN |
denotes |
polymorphism |
T3295 |
3090-3101 |
VBG |
denotes |
influencing |
T3296 |
3102-3114 |
NNS |
denotes |
interactions |
T3297 |
3115-3122 |
IN |
denotes |
between |
T3298 |
3123-3130 |
NNS |
denotes |
members |
T3299 |
3131-3133 |
IN |
denotes |
of |
T3300 |
3134-3137 |
DT |
denotes |
the |
T3301 |
3151-3158 |
NN |
denotes |
pathway |
T3302 |
3138-3140 |
NN |
denotes |
Gh |
T3303 |
3141-3150 |
NN |
denotes |
signaling |
T3304 |
3158-3159 |
. |
denotes |
. |
T3305 |
3159-3328 |
sentence |
denotes |
Thus, identification of hg modifiers has the potential to uncover novel members of metabolically important pathways or previously unknown convergences between pathways. |
T3306 |
3160-3164 |
RB |
denotes |
Thus |
T3307 |
3197-3200 |
VBZ |
denotes |
has |
T3308 |
3164-3166 |
, |
denotes |
, |
T3309 |
3166-3180 |
NN |
denotes |
identification |
T3310 |
3181-3183 |
IN |
denotes |
of |
T3311 |
3184-3186 |
NN |
denotes |
hg |
T3312 |
3187-3196 |
NNS |
denotes |
modifiers |
T3313 |
3201-3204 |
DT |
denotes |
the |
T3314 |
3205-3214 |
NN |
denotes |
potential |
T3315 |
3215-3217 |
TO |
denotes |
to |
T3316 |
3218-3225 |
VB |
denotes |
uncover |
T3317 |
3226-3231 |
JJ |
denotes |
novel |
T3318 |
3232-3239 |
NNS |
denotes |
members |
T3319 |
3240-3242 |
IN |
denotes |
of |
T3320 |
3243-3256 |
RB |
denotes |
metabolically |
T3321 |
3257-3266 |
JJ |
denotes |
important |
T3322 |
3267-3275 |
NNS |
denotes |
pathways |
T3323 |
3276-3278 |
CC |
denotes |
or |
T3324 |
3279-3289 |
RB |
denotes |
previously |
T3325 |
3290-3297 |
JJ |
denotes |
unknown |
T3326 |
3298-3310 |
NNS |
denotes |
convergences |
T3327 |
3311-3318 |
IN |
denotes |
between |
T3328 |
3319-3327 |
NNS |
denotes |
pathways |
T3329 |
3327-3328 |
. |
denotes |
. |
T3330 |
3328-3447 |
sentence |
denotes |
As an initial step, growth and carcass composition QTL were identified in a cross between CAST/EiJ (CAST) and HG [24]. |
T3331 |
3329-3331 |
IN |
denotes |
As |
T3332 |
3389-3399 |
VBN |
denotes |
identified |
T3333 |
3332-3334 |
DT |
denotes |
an |
T3334 |
3343-3347 |
NN |
denotes |
step |
T3335 |
3335-3342 |
JJ |
denotes |
initial |
T3336 |
3347-3349 |
, |
denotes |
, |
T3337 |
3349-3355 |
NN |
denotes |
growth |
T3338 |
3380-3383 |
NN |
denotes |
QTL |
T3339 |
3356-3359 |
CC |
denotes |
and |
T3340 |
3360-3367 |
NN |
denotes |
carcass |
T3341 |
3368-3379 |
NN |
denotes |
composition |
T3342 |
3384-3388 |
VBD |
denotes |
were |
T3343 |
3400-3402 |
IN |
denotes |
in |
T3344 |
3403-3404 |
DT |
denotes |
a |
T3345 |
3405-3410 |
NN |
denotes |
cross |
T3346 |
3411-3418 |
IN |
denotes |
between |
T3347 |
3419-3423 |
NN |
denotes |
CAST |
T3348 |
3424-3427 |
NN |
denotes |
EiJ |
T3349 |
3423-3424 |
HYPH |
denotes |
/ |
T3350 |
3428-3429 |
-LRB- |
denotes |
( |
T3351 |
3429-3433 |
NN |
denotes |
CAST |
T3352 |
3433-3434 |
-RRB- |
denotes |
) |
T3353 |
3435-3438 |
CC |
denotes |
and |
T3354 |
3439-3441 |
NN |
denotes |
HG |
T3355 |
3442-3443 |
-LRB- |
denotes |
[ |
T3356 |
3443-3445 |
CD |
denotes |
24 |
T3357 |
3445-3446 |
-RRB- |
denotes |
] |
T3358 |
3446-3447 |
. |
denotes |
. |
T3359 |
3447-3686 |
sentence |
denotes |
In the F2, mice homozygous for the hg deletion (hg/hg) and wild type (+/+) were assayed and hg modifier QTL were defined as those either absent in +/+ mice and segregating in hg/hg or loci with altered gene action dependent on background. |
T3360 |
3448-3450 |
IN |
denotes |
In |
T3361 |
3528-3535 |
VBN |
denotes |
assayed |
T3362 |
3451-3454 |
DT |
denotes |
the |
T3363 |
3455-3457 |
NN |
denotes |
F2 |
T3364 |
3457-3459 |
, |
denotes |
, |
T3365 |
3459-3463 |
NNS |
denotes |
mice |
T3366 |
3464-3474 |
JJ |
denotes |
homozygous |
T3367 |
3475-3478 |
IN |
denotes |
for |
T3368 |
3479-3482 |
DT |
denotes |
the |
T3369 |
3486-3494 |
NN |
denotes |
deletion |
T3370 |
3483-3485 |
NN |
denotes |
hg |
T3371 |
3495-3496 |
-LRB- |
denotes |
( |
T3372 |
3499-3501 |
NN |
denotes |
hg |
T3373 |
3496-3498 |
NN |
denotes |
hg |
T3374 |
3498-3499 |
HYPH |
denotes |
/ |
T3375 |
3501-3502 |
-RRB- |
denotes |
) |
T3376 |
3503-3506 |
CC |
denotes |
and |
T3377 |
3507-3511 |
JJ |
denotes |
wild |
T3378 |
3512-3516 |
NN |
denotes |
type |
T3379 |
3517-3518 |
-LRB- |
denotes |
( |
T3380 |
3518-3519 |
SYM |
denotes |
+ |
T3381 |
3520-3521 |
SYM |
denotes |
+ |
T3382 |
3519-3520 |
HYPH |
denotes |
/ |
T3383 |
3521-3522 |
-RRB- |
denotes |
) |
T3384 |
3523-3527 |
VBD |
denotes |
were |
T3385 |
3536-3539 |
CC |
denotes |
and |
T3386 |
3540-3542 |
NN |
denotes |
hg |
T3387 |
3543-3551 |
NN |
denotes |
modifier |
T3388 |
3552-3555 |
NN |
denotes |
QTL |
T3389 |
3561-3568 |
VBN |
denotes |
defined |
T3390 |
3556-3560 |
VBD |
denotes |
were |
T3391 |
3569-3571 |
IN |
denotes |
as |
T3392 |
3572-3577 |
DT |
denotes |
those |
T3393 |
3578-3584 |
CC |
denotes |
either |
T3394 |
3585-3591 |
JJ |
denotes |
absent |
T3395 |
3592-3594 |
IN |
denotes |
in |
T3396 |
3595-3596 |
SYM |
denotes |
+ |
T3397 |
3597-3598 |
SYM |
denotes |
+ |
T3398 |
3596-3597 |
HYPH |
denotes |
/ |
T3399 |
3599-3603 |
NNS |
denotes |
mice |
T3400 |
3604-3607 |
CC |
denotes |
and |
T3401 |
3608-3619 |
VBG |
denotes |
segregating |
T3402 |
3620-3622 |
IN |
denotes |
in |
T3403 |
3623-3625 |
NN |
denotes |
hg |
T3404 |
3626-3628 |
NN |
denotes |
hg |
T3405 |
3625-3626 |
HYPH |
denotes |
/ |
T3406 |
3629-3631 |
CC |
denotes |
or |
T3407 |
3632-3636 |
NNS |
denotes |
loci |
T3408 |
3637-3641 |
IN |
denotes |
with |
T3409 |
3642-3649 |
VBN |
denotes |
altered |
T3410 |
3655-3661 |
NN |
denotes |
action |
T3411 |
3650-3654 |
NN |
denotes |
gene |
T3412 |
3662-3671 |
JJ |
denotes |
dependent |
T3413 |
3672-3674 |
IN |
denotes |
on |
T3414 |
3675-3685 |
NN |
denotes |
background |
T3415 |
3685-3686 |
. |
denotes |
. |
T3416 |
3686-4089 |
sentence |
denotes |
Four hg modifier QTL (Wg2 on MMU2, Carfhg2 on MMU9, Carp2 on MMU11 and Feml3 on MMU17) were identified, along with 12 additional QTL (Q1Ucd1 on MMU1 (which only reached a suggestive level of statistical significance), Wg1, Carp1, Cara1 and Feml1 on MMU2, Carfhg1 on MMU5, Wg3 on MMU8, Feml2 on MMU9, Wg4 and Cara2 on MMU11 and Carp3 and Cara3 on MMU17) whose expression was independent of the hg locus. |
T3417 |
3687-3691 |
CD |
denotes |
Four |
T3418 |
3692-3694 |
NN |
denotes |
hg |
T3419 |
3704-3707 |
NN |
denotes |
QTL |
T3420 |
3695-3703 |
NN |
denotes |
modifier |
T3421 |
3779-3789 |
VBN |
denotes |
identified |
T3422 |
3708-3709 |
-LRB- |
denotes |
( |
T3423 |
3709-3712 |
NN |
denotes |
Wg2 |
T3424 |
3713-3715 |
IN |
denotes |
on |
T3425 |
3716-3720 |
NN |
denotes |
MMU2 |
T3426 |
3720-3722 |
, |
denotes |
, |
T3427 |
3722-3729 |
NN |
denotes |
Carfhg2 |
T3428 |
3730-3732 |
IN |
denotes |
on |
T3429 |
3733-3737 |
NN |
denotes |
MMU9 |
T3430 |
3737-3739 |
, |
denotes |
, |
T3431 |
3739-3744 |
NN |
denotes |
Carp2 |
T3432 |
3745-3747 |
IN |
denotes |
on |
T3433 |
3748-3753 |
NN |
denotes |
MMU11 |
T3434 |
3754-3757 |
CC |
denotes |
and |
T3435 |
3758-3763 |
NN |
denotes |
Feml3 |
T3436 |
3764-3766 |
IN |
denotes |
on |
T3437 |
3767-3772 |
NN |
denotes |
MMU17 |
T3438 |
3772-3773 |
-RRB- |
denotes |
) |
T3439 |
3774-3778 |
VBD |
denotes |
were |
T3440 |
3789-3791 |
, |
denotes |
, |
T3441 |
3791-3796 |
IN |
denotes |
along |
T3442 |
3797-3801 |
IN |
denotes |
with |
T3443 |
3802-3804 |
CD |
denotes |
12 |
T3444 |
3816-3819 |
NN |
denotes |
QTL |
T3445 |
3805-3815 |
JJ |
denotes |
additional |
T3446 |
3820-3821 |
-LRB- |
denotes |
( |
T3447 |
3821-3827 |
NN |
denotes |
Q1Ucd1 |
T3448 |
3828-3830 |
IN |
denotes |
on |
T3449 |
3831-3835 |
NN |
denotes |
MMU1 |
T3450 |
3836-3837 |
-LRB- |
denotes |
( |
T3451 |
3837-3842 |
WDT |
denotes |
which |
T3452 |
3848-3855 |
VBD |
denotes |
reached |
T3453 |
3843-3847 |
RB |
denotes |
only |
T3454 |
3856-3857 |
DT |
denotes |
a |
T3455 |
3869-3874 |
NN |
denotes |
level |
T3456 |
3858-3868 |
JJ |
denotes |
suggestive |
T3457 |
3875-3877 |
IN |
denotes |
of |
T3458 |
3878-3889 |
JJ |
denotes |
statistical |
T3459 |
3890-3902 |
NN |
denotes |
significance |
T3460 |
3902-3903 |
-RRB- |
denotes |
) |
T3461 |
3903-3905 |
, |
denotes |
, |
T3462 |
3905-3908 |
NN |
denotes |
Wg1 |
T3463 |
3908-3910 |
, |
denotes |
, |
T3464 |
3910-3915 |
NN |
denotes |
Carp1 |
T3465 |
3915-3917 |
, |
denotes |
, |
T3466 |
3917-3922 |
NN |
denotes |
Cara1 |
T3467 |
3923-3926 |
CC |
denotes |
and |
T3468 |
3927-3932 |
NN |
denotes |
Feml1 |
T3469 |
3933-3935 |
IN |
denotes |
on |
T3470 |
3936-3940 |
NN |
denotes |
MMU2 |
T3471 |
3940-3942 |
, |
denotes |
, |
T3472 |
3942-3949 |
NN |
denotes |
Carfhg1 |
T3473 |
3950-3952 |
IN |
denotes |
on |
T3474 |
3953-3957 |
NN |
denotes |
MMU5 |
T3475 |
3957-3959 |
, |
denotes |
, |
T3476 |
3959-3962 |
NN |
denotes |
Wg3 |
T3477 |
3963-3965 |
IN |
denotes |
on |
T3478 |
3966-3970 |
NN |
denotes |
MMU8 |
T3479 |
3970-3972 |
, |
denotes |
, |
T3480 |
3972-3977 |
NN |
denotes |
Feml2 |
T3481 |
3978-3980 |
IN |
denotes |
on |
T3482 |
3981-3985 |
NN |
denotes |
MMU9 |
T3483 |
3985-3987 |
, |
denotes |
, |
T3484 |
3987-3990 |
NN |
denotes |
Wg4 |
T3485 |
3991-3994 |
CC |
denotes |
and |
T3486 |
3995-4000 |
NN |
denotes |
Cara2 |
T3487 |
4001-4003 |
IN |
denotes |
on |
T3488 |
4004-4009 |
NN |
denotes |
MMU11 |
T3489 |
4010-4013 |
CC |
denotes |
and |
T3490 |
4014-4019 |
NN |
denotes |
Carp3 |
T3491 |
4020-4023 |
CC |
denotes |
and |
T3492 |
4024-4029 |
NN |
denotes |
Cara3 |
T3493 |
4030-4032 |
IN |
denotes |
on |
T3494 |
4033-4038 |
NN |
denotes |
MMU17 |
T3495 |
4038-4039 |
-RRB- |
denotes |
) |
T3496 |
4040-4045 |
WP$ |
denotes |
whose |
T3497 |
4046-4056 |
NN |
denotes |
expression |
T3498 |
4057-4060 |
VBD |
denotes |
was |
T3499 |
4061-4072 |
JJ |
denotes |
independent |
T3500 |
4073-4075 |
IN |
denotes |
of |
T3501 |
4076-4079 |
DT |
denotes |
the |
T3502 |
4083-4088 |
NN |
denotes |
locus |
T3503 |
4080-4082 |
NN |
denotes |
hg |
T3504 |
4088-4089 |
. |
denotes |
. |
T3505 |
4089-4190 |
sentence |
denotes |
In the current study we have developed speed congenic strains to isolate all the aforementioned QTL. |
T3506 |
4090-4092 |
IN |
denotes |
In |
T3507 |
4119-4128 |
VBN |
denotes |
developed |
T3508 |
4093-4096 |
DT |
denotes |
the |
T3509 |
4105-4110 |
NN |
denotes |
study |
T3510 |
4097-4104 |
JJ |
denotes |
current |
T3511 |
4111-4113 |
PRP |
denotes |
we |
T3512 |
4114-4118 |
VBP |
denotes |
have |
T3513 |
4129-4134 |
NN |
denotes |
speed |
T3514 |
4135-4143 |
JJ |
denotes |
congenic |
T3515 |
4144-4151 |
NNS |
denotes |
strains |
T3516 |
4152-4154 |
TO |
denotes |
to |
T3517 |
4155-4162 |
VB |
denotes |
isolate |
T3518 |
4163-4166 |
PDT |
denotes |
all |
T3519 |
4186-4189 |
NN |
denotes |
QTL |
T3520 |
4167-4170 |
DT |
denotes |
the |
T3521 |
4171-4185 |
JJ |
denotes |
aforementioned |
T3522 |
4189-4190 |
. |
denotes |
. |
T3523 |
4190-4418 |
sentence |
denotes |
Single strains on an HG background were created for each chromosomal region outside of MMU2, while a comprehensive panel of overlapping strains with identical donor regions on both B6 and HG backgrounds were developed for MMU2. |
T3524 |
4191-4197 |
JJ |
denotes |
Single |
T3525 |
4198-4205 |
NNS |
denotes |
strains |
T3526 |
4231-4238 |
VBN |
denotes |
created |
T3527 |
4206-4208 |
IN |
denotes |
on |
T3528 |
4209-4211 |
DT |
denotes |
an |
T3529 |
4215-4225 |
NN |
denotes |
background |
T3530 |
4212-4214 |
NN |
denotes |
HG |
T3531 |
4226-4230 |
VBD |
denotes |
were |
T3532 |
4239-4242 |
IN |
denotes |
for |
T3533 |
4243-4247 |
DT |
denotes |
each |
T3534 |
4260-4266 |
NN |
denotes |
region |
T3535 |
4248-4259 |
JJ |
denotes |
chromosomal |
T3536 |
4267-4274 |
IN |
denotes |
outside |
T3537 |
4275-4277 |
IN |
denotes |
of |
T3538 |
4278-4282 |
NN |
denotes |
MMU2 |
T3539 |
4282-4284 |
, |
denotes |
, |
T3540 |
4284-4289 |
IN |
denotes |
while |
T3541 |
4399-4408 |
VBN |
denotes |
developed |
T3542 |
4290-4291 |
DT |
denotes |
a |
T3543 |
4306-4311 |
NN |
denotes |
panel |
T3544 |
4292-4305 |
JJ |
denotes |
comprehensive |
T3545 |
4312-4314 |
IN |
denotes |
of |
T3546 |
4315-4326 |
VBG |
denotes |
overlapping |
T3547 |
4327-4334 |
NNS |
denotes |
strains |
T3548 |
4335-4339 |
IN |
denotes |
with |
T3549 |
4340-4349 |
JJ |
denotes |
identical |
T3550 |
4356-4363 |
NNS |
denotes |
regions |
T3551 |
4350-4355 |
NN |
denotes |
donor |
T3552 |
4364-4366 |
IN |
denotes |
on |
T3553 |
4367-4371 |
CC |
denotes |
both |
T3554 |
4372-4374 |
NN |
denotes |
B6 |
T3555 |
4382-4393 |
NNS |
denotes |
backgrounds |
T3556 |
4375-4378 |
CC |
denotes |
and |
T3557 |
4379-4381 |
NN |
denotes |
HG |
T3558 |
4394-4398 |
VBD |
denotes |
were |
T3559 |
4409-4412 |
IN |
denotes |
for |
T3560 |
4413-4417 |
NN |
denotes |
MMU2 |
T3561 |
4417-4418 |
. |
denotes |
. |
T3562 |
4418-4521 |
sentence |
denotes |
The MMU2 panel was developed to confirm the presence of multiple QTL and test for QTL-hg interactions. |
T3563 |
4419-4422 |
DT |
denotes |
The |
T3564 |
4428-4433 |
NN |
denotes |
panel |
T3565 |
4423-4427 |
NN |
denotes |
MMU2 |
T3566 |
4438-4447 |
VBN |
denotes |
developed |
T3567 |
4434-4437 |
VBD |
denotes |
was |
T3568 |
4448-4450 |
TO |
denotes |
to |
T3569 |
4451-4458 |
VB |
denotes |
confirm |
T3570 |
4459-4462 |
DT |
denotes |
the |
T3571 |
4463-4471 |
NN |
denotes |
presence |
T3572 |
4472-4474 |
IN |
denotes |
of |
T3573 |
4475-4483 |
JJ |
denotes |
multiple |
T3574 |
4484-4487 |
NN |
denotes |
QTL |
T3575 |
4488-4491 |
CC |
denotes |
and |
T3576 |
4492-4496 |
VB |
denotes |
test |
T3577 |
4497-4500 |
IN |
denotes |
for |
T3578 |
4501-4504 |
NN |
denotes |
QTL |
T3579 |
4505-4507 |
NN |
denotes |
hg |
T3580 |
4504-4505 |
HYPH |
denotes |
- |
T3581 |
4508-4520 |
NNS |
denotes |
interactions |
T3582 |
4520-4521 |
. |
denotes |
. |
T3583 |
4521-4713 |
sentence |
denotes |
Additionally, based on the knowledge of potential QTL-hg interactions, genes involved in Gh signaling, whose genomic location overlapped hg modifier QTL on MMU2, 9, 11 and 17, were sequenced. |
T3584 |
4522-4534 |
RB |
denotes |
Additionally |
T3585 |
4703-4712 |
VBN |
denotes |
sequenced |
T3586 |
4534-4536 |
, |
denotes |
, |
T3587 |
4536-4541 |
VBN |
denotes |
based |
T3588 |
4542-4544 |
IN |
denotes |
on |
T3589 |
4545-4548 |
DT |
denotes |
the |
T3590 |
4549-4558 |
NN |
denotes |
knowledge |
T3591 |
4559-4561 |
IN |
denotes |
of |
T3592 |
4562-4571 |
JJ |
denotes |
potential |
T3593 |
4579-4591 |
NNS |
denotes |
interactions |
T3594 |
4572-4575 |
NN |
denotes |
QTL |
T3595 |
4576-4578 |
NN |
denotes |
hg |
T3596 |
4575-4576 |
HYPH |
denotes |
- |
T3597 |
4591-4593 |
, |
denotes |
, |
T3598 |
4593-4598 |
NNS |
denotes |
genes |
T3599 |
4599-4607 |
VBN |
denotes |
involved |
T3600 |
4608-4610 |
IN |
denotes |
in |
T3601 |
4611-4613 |
NN |
denotes |
Gh |
T3602 |
4614-4623 |
NN |
denotes |
signaling |
T3603 |
4623-4625 |
, |
denotes |
, |
T3604 |
4625-4630 |
WP$ |
denotes |
whose |
T3605 |
4639-4647 |
NN |
denotes |
location |
T3606 |
4631-4638 |
JJ |
denotes |
genomic |
T3607 |
4648-4658 |
VBD |
denotes |
overlapped |
T3608 |
4659-4661 |
NN |
denotes |
hg |
T3609 |
4671-4674 |
NN |
denotes |
QTL |
T3610 |
4662-4670 |
NN |
denotes |
modifier |
T3611 |
4675-4677 |
IN |
denotes |
on |
T3612 |
4678-4681 |
NN |
denotes |
MMU |
T3613 |
4681-4682 |
CD |
denotes |
2 |
T3614 |
4682-4684 |
, |
denotes |
, |
T3615 |
4684-4685 |
CD |
denotes |
9 |
T3616 |
4685-4687 |
, |
denotes |
, |
T3617 |
4687-4689 |
CD |
denotes |
11 |
T3618 |
4690-4693 |
CC |
denotes |
and |
T3619 |
4694-4696 |
CD |
denotes |
17 |
T3620 |
4696-4698 |
, |
denotes |
, |
T3621 |
4698-4702 |
VBD |
denotes |
were |
T3622 |
4712-4713 |
. |
denotes |
. |
R295 |
T2734 |
T2732 |
pobj |
mutations,by |
R296 |
T2735 |
T2734 |
compound |
gene,mutations |
R305 |
T2746 |
T2739 |
dobj |
complexity,mimic |
R306 |
T2747 |
T2746 |
amod |
genetic,complexity |
R314 |
T2755 |
T2753 |
pobj |
population,in |
R315 |
T2756 |
T2755 |
amod |
human,population |
R329 |
T2772 |
T2770 |
pobj |
effects,by |
R330 |
T2773 |
T2772 |
amod |
small,effects |
R331 |
T2774 |
T2773 |
prep |
to,small |
R332 |
T2775 |
T2774 |
amod |
moderate,to |
R333 |
T2776 |
T2772 |
amod |
direct,effects |
R336 |
T2779 |
T2777 |
pobj |
loci,of |
R337 |
T2780 |
T2779 |
compound |
trait,loci |
R351 |
T2794 |
T2796 |
compound |
environment,interactions |
R352 |
T2795 |
T2794 |
punct |
-,environment |
R356 |
T2800 |
T2810 |
ccomp |
localized,made |
R357 |
T2801 |
T2799 |
pobj |
date,To |
R358 |
T2802 |
T2803 |
amod |
numerous,QTL |
R359 |
T2803 |
T2800 |
nsubjpass |
QTL,localized |
R360 |
T2804 |
T2805 |
nmod |
mouse,growth |
R361 |
T2805 |
T2803 |
nmod |
growth,QTL |
R362 |
T2806 |
T2805 |
cc |
and,growth |
R363 |
T2807 |
T2805 |
conj |
obesity,growth |
R364 |
T2808 |
T2800 |
aux |
have,localized |
R365 |
T2809 |
T2800 |
auxpass |
been,localized |
R367 |
T2812 |
T2800 |
parataxis |
2,localized |
R368 |
T2813 |
T2812 |
nummod |
1,2 |
R369 |
T2814 |
T2812 |
punct |
",",2 |
R381 |
T2826 |
T2824 |
dobj |
nature,determining |
R382 |
T2827 |
T2826 |
amod |
specific,nature |
R385 |
T2830 |
T2828 |
pobj |
variants,of |
R386 |
T2831 |
T2830 |
amod |
underlying,variants |
R387 |
T2832 |
T2830 |
amod |
genetic,variants |
R393 |
T2840 |
T2838 |
advcl |
map,used |
R394 |
T2841 |
T2840 |
advmod |
fine,map |
R398 |
T2903 |
T2904 |
amod |
congenic,analysis |
R399 |
T2845 |
T2846 |
dep |
which,designed |
R400 |
T2904 |
T2905 |
nsubj |
analysis,method |
R401 |
T2846 |
T2844 |
relcl |
designed,strains |
R402 |
T2905 |
T2899 |
ccomp |
method,considers |
R403 |
T2906 |
T2905 |
det |
an,method |
R404 |
T2847 |
T2846 |
auxpass |
are,designed |
R405 |
T2907 |
T2905 |
amod |
excellent,method |
R406 |
T2908 |
T2909 |
aux |
to,confirm |
R407 |
T2909 |
T2905 |
advcl |
confirm,method |
R408 |
T2848 |
T2849 |
aux |
to,convert |
R409 |
T2910 |
T2909 |
cc |
and,confirm |
R410 |
T2911 |
T2912 |
advmod |
subsequently,map |
R411 |
T2849 |
T2846 |
advcl |
convert,designed |
R412 |
T2912 |
T2909 |
conj |
map,confirm |
R413 |
T2913 |
T2912 |
advmod |
fine,map |
R414 |
T2914 |
T2912 |
dobj |
QTL,map |
R415 |
T2915 |
T2916 |
punct |
[,3 |
R416 |
T2850 |
T2851 |
det |
a,trait |
R417 |
T2916 |
T2899 |
parataxis |
3,considers |
R418 |
T2917 |
T2916 |
punct |
],3 |
R419 |
T2851 |
T2849 |
dobj |
trait,convert |
R420 |
T2918 |
T2899 |
punct |
.,considers |
R421 |
T2920 |
T2921 |
advmod |
Traditionally,developed |
R422 |
T2852 |
T2851 |
amod |
complex,trait |
R423 |
T2922 |
T2921 |
punct |
", ",developed |
R424 |
T2923 |
T2924 |
amod |
congenic,strains |
R425 |
T2853 |
T2851 |
amod |
polygenic,trait |
R426 |
T2924 |
T2921 |
nsubjpass |
strains,developed |
R427 |
T2925 |
T2921 |
auxpass |
are,developed |
R428 |
T2854 |
T2849 |
prep |
into,convert |
R429 |
T2926 |
T2921 |
prep |
by,developed |
R430 |
T2927 |
T2926 |
pcomp |
introgressing,by |
R431 |
T2928 |
T2929 |
compound |
QTL,alleles |
R432 |
T2929 |
T2927 |
dobj |
alleles,introgressing |
R433 |
T2855 |
T2854 |
pobj |
one,into |
R434 |
T2930 |
T2929 |
prep |
from,alleles |
R435 |
T2931 |
T2932 |
det |
a,strain |
R436 |
T2932 |
T2930 |
pobj |
strain,from |
R437 |
T2933 |
T2932 |
compound |
donor,strain |
R438 |
T2934 |
T2932 |
punct |
", ",strain |
R439 |
T2856 |
T2857 |
dep |
that,is |
R440 |
T2935 |
T2936 |
poss |
whose,boundaries |
R441 |
T2936 |
T2937 |
dep |
boundaries,defined |
R442 |
T2857 |
T2855 |
relcl |
is,one |
R443 |
T2937 |
T2932 |
relcl |
defined,strain |
R444 |
T2858 |
T2859 |
advmod |
mono,oligogenic |
R445 |
T2938 |
T2937 |
auxpass |
are,defined |
R446 |
T2939 |
T2937 |
agent |
by,defined |
R447 |
T2940 |
T2941 |
amod |
genetic,markers |
R448 |
T2859 |
T2857 |
acomp |
oligogenic,is |
R449 |
T2941 |
T2939 |
pobj |
markers,by |
R450 |
T2942 |
T2927 |
punct |
", ",introgressing |
R451 |
T2943 |
T2927 |
prep |
on,introgressing |
R452 |
T2860 |
T2859 |
punct |
-,oligogenic |
R453 |
T2944 |
T2945 |
det |
the,background |
R454 |
T2945 |
T2943 |
pobj |
background,on |
R455 |
T2946 |
T2945 |
amod |
genetic,background |
R456 |
T2861 |
T2859 |
cc |
or,oligogenic |
R457 |
T2947 |
T2945 |
prep |
of,background |
R458 |
T2948 |
T2949 |
det |
a,strain |
R459 |
T2949 |
T2947 |
pobj |
strain,of |
R460 |
T2862 |
T2837 |
punct |
.,are |
R461 |
T2950 |
T2949 |
compound |
recipient,strain |
R462 |
T2951 |
T2927 |
prep |
via,introgressing |
R463 |
T2864 |
T2865 |
nsubjpass |
This,accomplished |
R464 |
T2952 |
T2953 |
nummod |
10,backcrosses |
R465 |
T2953 |
T2951 |
pobj |
backcrosses,via |
R466 |
T2954 |
T2955 |
punct |
[,4 |
R467 |
T2955 |
T2927 |
parataxis |
4,introgressing |
R468 |
T2956 |
T2955 |
punct |
],4 |
R469 |
T2866 |
T2865 |
auxpass |
is,accomplished |
R470 |
T2957 |
T2921 |
punct |
.,developed |
R471 |
T2959 |
T2960 |
mark |
While,straightforward |
R472 |
T2960 |
T2962 |
advcl |
straightforward,is |
R473 |
T2867 |
T2865 |
prep |
by,accomplished |
R474 |
T2961 |
T2960 |
advmod |
technically,straightforward |
R475 |
T2963 |
T2962 |
nsubj |
this,is |
R476 |
T2868 |
T2867 |
pcomp |
eliminating,by |
R477 |
T2964 |
T2965 |
det |
a,endeavor |
R478 |
T2965 |
T2962 |
attr |
endeavor,is |
R479 |
T2966 |
T2967 |
npadvmod |
time,consuming |
R480 |
T2869 |
T2868 |
xcomp |
segregating,eliminating |
R481 |
T2967 |
T2965 |
amod |
consuming,endeavor |
R482 |
T2968 |
T2965 |
acl |
taking,endeavor |
R483 |
T2969 |
T2970 |
quantmod |
over,three |
R484 |
T2870 |
T2871 |
amod |
genetic,variation |
R485 |
T2970 |
T2971 |
nummod |
three,years |
R486 |
T2971 |
T2968 |
dobj |
years,taking |
R487 |
T2972 |
T2973 |
aux |
to,construct |
R488 |
T2973 |
T2968 |
advcl |
construct,taking |
R489 |
T2974 |
T2975 |
det |
a,strain |
R490 |
T2975 |
T2973 |
dobj |
strain,construct |
R491 |
T2871 |
T2869 |
dobj |
variation,segregating |
R492 |
T2976 |
T2975 |
amod |
single,strain |
R493 |
T2977 |
T2962 |
punct |
.,is |
R494 |
T2872 |
T2869 |
prep |
outside,segregating |
R495 |
T2979 |
T2980 |
det |
The,approach |
R496 |
T2980 |
T2983 |
nsubj |
approach,is |
R497 |
T2873 |
T2874 |
det |
the,locus |
R498 |
T2981 |
T2982 |
npadvmod |
speed,congenic |
R499 |
T2982 |
T2980 |
amod |
congenic,approach |
R500 |
T2984 |
T2985 |
det |
an,alternative |
R501 |
T2874 |
T2872 |
pobj |
locus,outside |
R502 |
T2985 |
T2983 |
attr |
alternative,is |
R503 |
T2986 |
T2985 |
prep |
to,alternative |
R504 |
T2875 |
T2874 |
prep |
of,locus |
R505 |
T2987 |
T2988 |
det |
this,process |
R506 |
T2988 |
T2986 |
pobj |
process,to |
R507 |
T2989 |
T2988 |
amod |
lengthy,process |
R508 |
T2876 |
T2875 |
pobj |
interest,of |
R509 |
T2990 |
T2983 |
cc |
and,is |
R510 |
T2991 |
T2992 |
aux |
can,reduce |
R511 |
T2992 |
T2983 |
conj |
reduce,is |
R512 |
T2993 |
T2994 |
det |
the,number |
R513 |
T2877 |
T2868 |
cc |
and,eliminating |
R514 |
T2994 |
T2992 |
dobj |
number,reduce |
R515 |
T2995 |
T2994 |
prep |
of,number |
R516 |
T2878 |
T2868 |
conj |
reducing,eliminating |
R517 |
T2996 |
T2997 |
amod |
required,backcrosses |
R518 |
T2997 |
T2995 |
pobj |
backcrosses,of |
R519 |
T2998 |
T2992 |
prep |
from,reduce |
R520 |
T2879 |
T2880 |
det |
the,variation |
R521 |
T2999 |
T2998 |
pobj |
10,from |
R522 |
T3000 |
T2992 |
prep |
to,reduce |
R523 |
T3001 |
T3000 |
pobj |
five,to |
R524 |
T2880 |
T2878 |
dobj |
variation,reducing |
R525 |
T3002 |
T3003 |
punct |
[,6 |
R526 |
T3003 |
T2992 |
parataxis |
6,reduce |
R527 |
T3004 |
T3003 |
nummod |
5,6 |
R528 |
T2881 |
T2880 |
amod |
environmental,variation |
R529 |
T3005 |
T3003 |
punct |
",",6 |
R530 |
T3006 |
T3003 |
punct |
],6 |
R531 |
T3007 |
T2983 |
punct |
.,is |
R532 |
T2882 |
T2880 |
acl |
influencing,variation |
R533 |
T2883 |
T2884 |
det |
a,trait |
R534 |
T2884 |
T2882 |
dobj |
trait,influencing |
R535 |
T3009 |
T3010 |
det |
This,strategy |
R536 |
T2885 |
T2878 |
prep |
by,reducing |
R537 |
T3010 |
T3011 |
nsubj |
strategy,uses |
R538 |
T3012 |
T3013 |
npadvmod |
marker,assisted |
R539 |
T3013 |
T3015 |
amod |
assisted,selection |
R540 |
T3014 |
T3013 |
punct |
-,assisted |
R541 |
T3015 |
T3011 |
dobj |
selection,uses |
R542 |
T2886 |
T2885 |
pcomp |
characterizing,by |
R543 |
T3016 |
T3017 |
aux |
to,identify |
R544 |
T3017 |
T3011 |
advcl |
identify,uses |
R545 |
T3018 |
T3019 |
amod |
male,mice |
R546 |
T2887 |
T2888 |
amod |
large,numbers |
R547 |
T3019 |
T3017 |
dobj |
mice,identify |
R548 |
T2888 |
T2886 |
dobj |
numbers,characterizing |
R549 |
T3020 |
T3019 |
acl |
inheriting,mice |
R550 |
T3021 |
T3022 |
amod |
fewer,alleles |
R551 |
T2889 |
T2888 |
prep |
of,numbers |
R552 |
T2890 |
T2891 |
advmod |
genetically,identical |
R553 |
T3022 |
T3020 |
dobj |
alleles,inheriting |
R554 |
T3023 |
T3022 |
compound |
donor,alleles |
R555 |
T2891 |
T2892 |
amod |
identical,mice |
R556 |
T3024 |
T3022 |
punct |
", ",alleles |
R557 |
T3025 |
T3026 |
mark |
than,expected |
R558 |
T3026 |
T3022 |
advcl |
expected,alleles |
R559 |
T2892 |
T2889 |
pobj |
mice,of |
R560 |
T3027 |
T3020 |
prep |
on,inheriting |
R561 |
T3028 |
T3027 |
amod |
average,on |
R562 |
T3029 |
T3011 |
punct |
", ",uses |
R563 |
T2893 |
T2865 |
punct |
.,accomplished |
R564 |
T3030 |
T3011 |
prep |
during,uses |
R565 |
T3031 |
T3032 |
det |
each,backcross |
R566 |
T2895 |
T2896 |
det |
The,Consortium |
R567 |
T3032 |
T3030 |
pobj |
backcross,during |
R568 |
T3033 |
T3011 |
punct |
.,uses |
R569 |
T2896 |
T2899 |
nsubj |
Consortium,considers |
R570 |
T3035 |
T3036 |
amod |
Numerous,congenics |
R571 |
T3036 |
T3041 |
nsubjpass |
congenics,used |
R572 |
T3037 |
T3038 |
advmod |
traditionally,developed |
R573 |
T2897 |
T2898 |
compound |
Complex,Trait |
R574 |
T3038 |
T3036 |
amod |
developed,congenics |
R575 |
T3039 |
T3038 |
cc |
and,developed |
R576 |
T3040 |
T3038 |
conj |
speed,developed |
R577 |
T2898 |
T2896 |
compound |
Trait,Consortium |
R578 |
T3042 |
T3041 |
aux |
have,used |
R579 |
T3043 |
T3041 |
auxpass |
been,used |
R580 |
T2900 |
T2896 |
punct |
(,Consortium |
R581 |
T3044 |
T3045 |
aux |
to,isolate |
R582 |
T3045 |
T3041 |
advcl |
isolate,used |
R583 |
T3046 |
T3045 |
advmod |
successfully,isolate |
R584 |
T3047 |
T3048 |
compound |
mouse,QTL |
R585 |
T2901 |
T2896 |
appos |
CTC,Consortium |
R586 |
T3048 |
T3045 |
dobj |
QTL,isolate |
R587 |
T3049 |
T3045 |
prep |
for,isolate |
R588 |
T2902 |
T2899 |
punct |
),considers |
R590 |
T3051 |
T3049 |
pobj |
array,for |
R591 |
T3052 |
T3051 |
amod |
wide,array |
R595 |
T3114 |
T3115 |
det |
The,role |
R596 |
T3056 |
T3051 |
prep |
including,array |
R597 |
T3057 |
T3056 |
pobj |
growth,including |
R598 |
T3058 |
T3057 |
cc |
and,growth |
R599 |
T3059 |
T3057 |
conj |
obesity,growth |
R600 |
T3115 |
T3116 |
nsubjpass |
role,confirmed |
R601 |
T3060 |
T3061 |
punct |
[,7 |
R602 |
T3061 |
T3041 |
parataxis |
7,used |
R603 |
T3062 |
T3063 |
punct |
-,13 |
R604 |
T3117 |
T3115 |
prep |
of,role |
R605 |
T3063 |
T3061 |
prep |
13,7 |
R606 |
T3064 |
T3061 |
punct |
],7 |
R607 |
T3065 |
T3041 |
punct |
.,used |
R608 |
T3067 |
T3068 |
det |
The,mouse |
R609 |
T3118 |
T3117 |
pobj |
Socs2,of |
R610 |
T3068 |
T3079 |
nsubj |
mouse,is |
R611 |
T3069 |
T3070 |
nmod |
C57Bl,hg |
R613 |
T3119 |
T3115 |
prep |
in,role |
R614 |
T3071 |
T3070 |
punct |
/,hg |
R615 |
T3072 |
T3070 |
nmod |
6J,hg |
R616 |
T3073 |
T3070 |
punct |
-,hg |
R617 |
T3074 |
T3070 |
nmod |
hg,hg |
R618 |
T3120 |
T3121 |
det |
the,phenotype |
R619 |
T3075 |
T3070 |
punct |
/,hg |
R620 |
T3076 |
T3077 |
punct |
(,HG |
R621 |
T3077 |
T3070 |
parataxis |
HG,hg |
R622 |
T3121 |
T3119 |
pobj |
phenotype,in |
R623 |
T3078 |
T3077 |
punct |
),HG |
R624 |
T3080 |
T3081 |
det |
a,model |
R625 |
T3122 |
T3121 |
compound |
HG,phenotype |
R626 |
T3081 |
T3079 |
attr |
model,is |
R627 |
T3082 |
T3081 |
prep |
of,model |
R628 |
T3083 |
T3084 |
amod |
systemic,overgrowth |
R629 |
T3123 |
T3116 |
auxpass |
was,confirmed |
R630 |
T3084 |
T3082 |
pobj |
overgrowth,of |
R631 |
T3085 |
T3081 |
acl |
resulting,model |
R632 |
T3124 |
T3116 |
agent |
by,confirmed |
R633 |
T3086 |
T3085 |
prep |
from,resulting |
R634 |
T3087 |
T3088 |
det |
a,deletion |
R635 |
T3088 |
T3086 |
pobj |
deletion,from |
R636 |
T3125 |
T3126 |
det |
an,mouse |
R637 |
T3089 |
T3088 |
amod |
spontaneous,deletion |
R638 |
T3090 |
T3088 |
prep |
on,deletion |
R639 |
T3091 |
T3090 |
pobj |
MMU10,on |
R640 |
T3126 |
T3124 |
pobj |
mouse,by |
R641 |
T3092 |
T3091 |
punct |
", ",MMU10 |
R642 |
T3093 |
T3094 |
dep |
which,eliminates |
R643 |
T3094 |
T3091 |
relcl |
eliminates,MMU10 |
R644 |
T3095 |
T3094 |
dobj |
expression,eliminates |
R645 |
T3096 |
T3095 |
prep |
of,expression |
R646 |
T3097 |
T3098 |
det |
the,gene |
R647 |
T3127 |
T3128 |
advmod |
independently,engineered |
R648 |
T3098 |
T3096 |
pobj |
gene,of |
R649 |
T3099 |
T3098 |
nmod |
Socs2,gene |
R650 |
T3100 |
T3099 |
punct |
(,Socs2 |
R651 |
T3101 |
T3099 |
appos |
suppressor,Socs2 |
R652 |
T3128 |
T3126 |
amod |
engineered,mouse |
R653 |
T3102 |
T3101 |
prep |
of,suppressor |
R654 |
T3103 |
T3104 |
compound |
cytokine,signaling |
R655 |
T3104 |
T3102 |
pobj |
signaling,of |
R656 |
T3129 |
T3130 |
nmod |
Socs2,knockout |
R657 |
T3105 |
T3104 |
nummod |
2,signaling |
R658 |
T3106 |
T3098 |
punct |
),gene |
R659 |
T3107 |
T3108 |
punct |
[,14 |
R660 |
T3130 |
T3126 |
compound |
knockout,mouse |
R661 |
T3131 |
T3129 |
punct |
-,Socs2 |
R662 |
T3108 |
T3079 |
parataxis |
14,is |
R663 |
T3109 |
T3110 |
punct |
-,16 |
R664 |
T3132 |
T3129 |
punct |
/,Socs2 |
R665 |
T3133 |
T3129 |
punct |
-,Socs2 |
R666 |
T3110 |
T3108 |
prep |
16,14 |
R667 |
T3134 |
T3135 |
dep |
which,shared |
R668 |
T3111 |
T3108 |
punct |
],14 |
R669 |
T3112 |
T3079 |
punct |
.,is |
R674 |
T3220 |
T3219 |
prep |
of,basis |
R675 |
T3221 |
T3222 |
det |
these,interactions |
R676 |
T3139 |
T3138 |
pobj |
phenotypes,of |
R677 |
T3222 |
T3220 |
pobj |
interactions,of |
R678 |
T3223 |
T3217 |
punct |
", ",forms |
R679 |
T3224 |
T3217 |
advcl |
implying,forms |
R680 |
T3140 |
T3137 |
prep |
in,number |
R681 |
T3225 |
T3226 |
mark |
that,are |
R682 |
T3226 |
T3224 |
ccomp |
are,implying |
R683 |
T3141 |
T3140 |
pobj |
common,in |
R684 |
T3227 |
T3228 |
det |
the,gene |
R685 |
T3228 |
T3226 |
nsubj |
gene,are |
R686 |
T3229 |
T3228 |
amod |
known,gene |
R687 |
T3230 |
T3228 |
cc |
and,gene |
R688 |
T3142 |
T3141 |
prep |
with,common |
R689 |
T3231 |
T3232 |
poss |
its,modifiers |
R690 |
T3232 |
T3228 |
conj |
modifiers,gene |
R691 |
T3143 |
T3142 |
pobj |
HG,with |
R692 |
T3233 |
T3226 |
attr |
members,are |
R693 |
T3234 |
T3233 |
prep |
of,members |
R694 |
T3144 |
T3137 |
punct |
", ",number |
R695 |
T3235 |
T3236 |
det |
the,pathway |
R696 |
T3236 |
T3234 |
pobj |
pathway,of |
R697 |
T3237 |
T3236 |
amod |
same,pathway |
R698 |
T3238 |
T3236 |
amod |
biochemical,pathway |
R699 |
T3239 |
T3238 |
cc |
or,biochemical |
R700 |
T3240 |
T3238 |
conj |
metabolic,biochemical |
R701 |
T3241 |
T3217 |
punct |
.,forms |
R702 |
T3145 |
T3137 |
prep |
including,number |
R703 |
T3243 |
T3244 |
amod |
Different,backgrounds |
R704 |
T3244 |
T3246 |
nsubjpass |
backgrounds,shown |
R705 |
T3245 |
T3244 |
amod |
genetic,backgrounds |
R706 |
T3146 |
T3145 |
pobj |
gigantism,including |
R707 |
T3247 |
T3246 |
aux |
have,shown |
R708 |
T3248 |
T3246 |
auxpass |
been,shown |
R709 |
T3147 |
T3148 |
punct |
[,17 |
R710 |
T3249 |
T3250 |
aux |
to,modify |
R711 |
T3148 |
T3116 |
parataxis |
17,confirmed |
R712 |
T3250 |
T3246 |
xcomp |
modify,shown |
R713 |
T3251 |
T3252 |
det |
the,effects |
R714 |
T3252 |
T3250 |
dobj |
effects,modify |
R715 |
T3149 |
T3148 |
punct |
],17 |
R716 |
T3253 |
T3254 |
npadvmod |
growth,enhancing |
R717 |
T3254 |
T3252 |
amod |
enhancing,effects |
R718 |
T3255 |
T3254 |
punct |
-,enhancing |
R719 |
T3150 |
T3116 |
punct |
.,confirmed |
R720 |
T3256 |
T3252 |
prep |
of,effects |
R721 |
T3257 |
T3256 |
pobj |
hg,of |
R722 |
T3258 |
T3259 |
punct |
[,22 |
R723 |
T3259 |
T3246 |
parataxis |
22,shown |
R724 |
T3152 |
T3153 |
nsubjpass |
QTL,referred |
R725 |
T3260 |
T3259 |
nummod |
15,22 |
R726 |
T3261 |
T3259 |
punct |
",",22 |
R727 |
T3262 |
T3259 |
punct |
],22 |
R728 |
T3154 |
T3155 |
dep |
which,alter |
R729 |
T3263 |
T3246 |
punct |
.,shown |
R730 |
T3265 |
T3266 |
mark |
Since,is |
R731 |
T3266 |
T3272 |
advcl |
is,is |
R732 |
T3267 |
T3268 |
det |
the,function |
R733 |
T3155 |
T3152 |
relcl |
alter,QTL |
R734 |
T3268 |
T3266 |
nsubj |
function,is |
R735 |
T3269 |
T3268 |
amod |
primary,function |
R736 |
T3270 |
T3268 |
prep |
of,function |
R737 |
T3156 |
T3157 |
det |
the,effects |
R738 |
T3271 |
T3270 |
pobj |
Socs2,of |
R739 |
T3157 |
T3155 |
dobj |
effects,alter |
R740 |
T3273 |
T3274 |
aux |
to,regulate |
R741 |
T3274 |
T3266 |
xcomp |
regulate,is |
R742 |
T3275 |
T3274 |
advmod |
negatively,regulate |
R743 |
T3276 |
T3277 |
compound |
growth,hormone |
R744 |
T3158 |
T3157 |
amod |
phenotypic,effects |
R745 |
T3277 |
T3274 |
dobj |
hormone,regulate |
R746 |
T3278 |
T3277 |
punct |
(,hormone |
R747 |
T3159 |
T3157 |
prep |
of,effects |
R748 |
T3279 |
T3277 |
appos |
Gh,hormone |
R749 |
T3280 |
T3266 |
punct |
),is |
R750 |
T3281 |
T3282 |
punct |
[,23 |
R751 |
T3282 |
T3266 |
parataxis |
23,is |
R752 |
T3160 |
T3161 |
det |
another,locus |
R753 |
T3283 |
T3282 |
punct |
],23 |
R754 |
T3284 |
T3272 |
punct |
", ",is |
R755 |
T3285 |
T3272 |
nsubj |
it,is |
R756 |
T3161 |
T3159 |
pobj |
locus,of |
R757 |
T3286 |
T3272 |
acomp |
likely,is |
R758 |
T3287 |
T3288 |
det |
these,effects |
R759 |
T3162 |
T3153 |
auxpass |
are,referred |
R760 |
T3288 |
T3290 |
nsubj |
effects,are |
R761 |
T3289 |
T3288 |
compound |
background,effects |
R762 |
T3163 |
T3153 |
prep |
to,referred |
R763 |
T3290 |
T3286 |
advcl |
are,likely |
R764 |
T3291 |
T3292 |
det |
the,result |
R765 |
T3164 |
T3153 |
prep |
as,referred |
R766 |
T3292 |
T3290 |
attr |
result,are |
R767 |
T3293 |
T3292 |
prep |
of,result |
R768 |
T3294 |
T3295 |
nsubj |
polymorphism,influencing |
R769 |
T3295 |
T3293 |
pcomp |
influencing,of |
R770 |
T3165 |
T3166 |
compound |
modifier,QTL |
R771 |
T3296 |
T3295 |
dobj |
interactions,influencing |
R772 |
T3297 |
T3296 |
prep |
between,interactions |
R773 |
T3298 |
T3297 |
pobj |
members,between |
R774 |
T3166 |
T3164 |
pobj |
QTL,as |
R775 |
T3299 |
T3298 |
prep |
of,members |
R776 |
T3300 |
T3301 |
det |
the,pathway |
R777 |
T3301 |
T3299 |
pobj |
pathway,of |
R778 |
T3167 |
T3168 |
punct |
[,18 |
R779 |
T3302 |
T3303 |
compound |
Gh,signaling |
R780 |
T3303 |
T3301 |
compound |
signaling,pathway |
R781 |
T3304 |
T3272 |
punct |
.,is |
R782 |
T3168 |
T3153 |
parataxis |
18,referred |
R783 |
T3306 |
T3307 |
advmod |
Thus,has |
R784 |
T3169 |
T3168 |
punct |
],18 |
R785 |
T3308 |
T3307 |
punct |
", ",has |
R786 |
T3309 |
T3307 |
nsubj |
identification,has |
R787 |
T3170 |
T3153 |
punct |
.,referred |
R788 |
T3310 |
T3309 |
prep |
of,identification |
R789 |
T3311 |
T3312 |
compound |
hg,modifiers |
R790 |
T3312 |
T3310 |
pobj |
modifiers,of |
R791 |
T3172 |
T3173 |
compound |
Modifier,QTL |
R792 |
T3313 |
T3314 |
det |
the,potential |
R793 |
T3314 |
T3307 |
dobj |
potential,has |
R794 |
T3315 |
T3316 |
aux |
to,uncover |
R795 |
T3316 |
T3314 |
acl |
uncover,potential |
R796 |
T3173 |
T3174 |
nsubjpass |
QTL,mapped |
R797 |
T3317 |
T3318 |
amod |
novel,members |
R798 |
T3318 |
T3316 |
dobj |
members,uncover |
R799 |
T3319 |
T3318 |
prep |
of,members |
R800 |
T3320 |
T3321 |
advmod |
metabolically,important |
R801 |
T3321 |
T3322 |
amod |
important,pathways |
R802 |
T3322 |
T3319 |
pobj |
pathways,of |
R803 |
T3175 |
T3174 |
aux |
have,mapped |
R804 |
T3323 |
T3318 |
cc |
or,members |
R805 |
T3324 |
T3325 |
advmod |
previously,unknown |
R806 |
T3325 |
T3326 |
amod |
unknown,convergences |
R807 |
T3176 |
T3174 |
auxpass |
been,mapped |
R808 |
T3177 |
T3174 |
prep |
for,mapped |
R809 |
T3326 |
T3318 |
conj |
convergences,members |
R810 |
T3178 |
T3179 |
amod |
numerous,traits |
R811 |
T3327 |
T3326 |
prep |
between,convergences |
R812 |
T3328 |
T3327 |
pobj |
pathways,between |
R813 |
T3329 |
T3307 |
punct |
.,has |
R814 |
T3179 |
T3177 |
pobj |
traits,for |
R815 |
T3331 |
T3332 |
prep |
As,identified |
R816 |
T3180 |
T3174 |
cc |
and,mapped |
R817 |
T3333 |
T3334 |
det |
an,step |
R818 |
T3334 |
T3331 |
pobj |
step,As |
R819 |
T3181 |
T3182 |
prep |
in,is |
R820 |
T3335 |
T3334 |
amod |
initial,step |
R821 |
T3336 |
T3332 |
punct |
", ",identified |
R822 |
T3337 |
T3338 |
nmod |
growth,QTL |
R823 |
T3182 |
T3174 |
conj |
is,mapped |
R824 |
T3338 |
T3332 |
nsubjpass |
QTL,identified |
R825 |
T3339 |
T3337 |
cc |
and,growth |
R826 |
T3340 |
T3341 |
compound |
carcass,composition |
R827 |
T3183 |
T3184 |
det |
these,studies |
R828 |
T3341 |
T3337 |
conj |
composition,growth |
R829 |
T3342 |
T3332 |
auxpass |
were,identified |
R830 |
T3343 |
T3332 |
prep |
in,identified |
R831 |
T3184 |
T3181 |
pobj |
studies,in |
R832 |
T3344 |
T3345 |
det |
a,cross |
R833 |
T3345 |
T3343 |
pobj |
cross,in |
R834 |
T3346 |
T3345 |
prep |
between,cross |
R835 |
T3185 |
T3186 |
det |
the,locus |
R836 |
T3347 |
T3348 |
compound |
CAST,EiJ |
R837 |
T3348 |
T3346 |
pobj |
EiJ,between |
R838 |
T3349 |
T3348 |
punct |
/,EiJ |
R839 |
T3186 |
T3182 |
nsubj |
locus,is |
R840 |
T3350 |
T3348 |
punct |
(,EiJ |
R841 |
T3351 |
T3348 |
appos |
CAST,EiJ |
R842 |
T3352 |
T3348 |
punct |
),EiJ |
R843 |
T3187 |
T3186 |
amod |
modified,locus |
R844 |
T3353 |
T3348 |
cc |
and,EiJ |
R845 |
T3354 |
T3348 |
conj |
HG,EiJ |
R846 |
T3355 |
T3356 |
punct |
[,24 |
R847 |
T3188 |
T3182 |
advmod |
typically,is |
R848 |
T3356 |
T3332 |
parataxis |
24,identified |
R849 |
T3357 |
T3356 |
punct |
],24 |
R850 |
T3358 |
T3332 |
punct |
.,identified |
R851 |
T3189 |
T3190 |
det |
a,gene |
R852 |
T3360 |
T3361 |
prep |
In,assayed |
R853 |
T3362 |
T3363 |
det |
the,F2 |
R854 |
T3363 |
T3360 |
pobj |
F2,In |
R855 |
T3364 |
T3361 |
punct |
", ",assayed |
R856 |
T3190 |
T3182 |
attr |
gene,is |
R857 |
T3365 |
T3361 |
nsubjpass |
mice,assayed |
R858 |
T3366 |
T3365 |
amod |
homozygous,mice |
R859 |
T3191 |
T3190 |
amod |
known,gene |
R860 |
T3367 |
T3366 |
prep |
for,homozygous |
R861 |
T3192 |
T3190 |
acl |
containing,gene |
R862 |
T3368 |
T3369 |
det |
the,deletion |
R863 |
T3369 |
T3367 |
pobj |
deletion,for |
R864 |
T3193 |
T3194 |
det |
a,mutation |
R865 |
T3370 |
T3369 |
compound |
hg,deletion |
R866 |
T3371 |
T3372 |
punct |
(,hg |
R867 |
T3372 |
T3369 |
parataxis |
hg,deletion |
R868 |
T3194 |
T3192 |
dobj |
mutation,containing |
R869 |
T3373 |
T3372 |
compound |
hg,hg |
R870 |
T3374 |
T3372 |
punct |
/,hg |
R871 |
T3375 |
T3372 |
punct |
),hg |
R872 |
T3195 |
T3196 |
advmod |
spontaneously,arisen |
R873 |
T3376 |
T3366 |
cc |
and,homozygous |
R874 |
T3196 |
T3194 |
amod |
arisen,mutation |
R875 |
T3377 |
T3378 |
amod |
wild,type |
R876 |
T3378 |
T3366 |
conj |
type,homozygous |
R877 |
T3379 |
T3378 |
punct |
(,type |
R878 |
T3380 |
T3381 |
punct |
+,+ |
R879 |
T3197 |
T3196 |
cc |
or,arisen |
R880 |
T3381 |
T3378 |
appos |
+,type |
R881 |
T3382 |
T3381 |
punct |
/,+ |
R882 |
T3383 |
T3361 |
punct |
),assayed |
R883 |
T3198 |
T3196 |
conj |
engineered,arisen |
R884 |
T3384 |
T3361 |
auxpass |
were,assayed |
R885 |
T3385 |
T3361 |
cc |
and,assayed |
R886 |
T3199 |
T3194 |
prep |
with,mutation |
R887 |
T3386 |
T3387 |
compound |
hg,modifier |
R888 |
T3387 |
T3388 |
compound |
modifier,QTL |
R889 |
T3388 |
T3389 |
nsubjpass |
QTL,defined |
R890 |
T3389 |
T3361 |
conj |
defined,assayed |
R891 |
T3200 |
T3201 |
amod |
major,effects |
R892 |
T3390 |
T3389 |
auxpass |
were,defined |
R893 |
T3391 |
T3389 |
prep |
as,defined |
R894 |
T3392 |
T3391 |
pobj |
those,as |
R895 |
T3201 |
T3199 |
pobj |
effects,with |
R896 |
T3393 |
T3392 |
preconj |
either,those |
R897 |
T3394 |
T3392 |
amod |
absent,those |
R898 |
T3202 |
T3201 |
amod |
phenotypic,effects |
R899 |
T3395 |
T3394 |
prep |
in,absent |
R900 |
T3396 |
T3397 |
punct |
+,+ |
R901 |
T3397 |
T3399 |
punct |
+,mice |
R902 |
T3398 |
T3397 |
punct |
/,+ |
R903 |
T3203 |
T3194 |
punct |
", ",mutation |
R904 |
T3399 |
T3395 |
pobj |
mice,in |
R905 |
T3400 |
T3394 |
cc |
and,absent |
R906 |
T3401 |
T3394 |
conj |
segregating,absent |
R907 |
T3402 |
T3401 |
prep |
in,segregating |
R908 |
T3403 |
T3404 |
compound |
hg,hg |
R909 |
T3404 |
T3402 |
pobj |
hg,in |
R910 |
T3204 |
T3205 |
amod |
such,as |
R911 |
T3405 |
T3404 |
punct |
/,hg |
R912 |
T3406 |
T3392 |
cc |
or,those |
R913 |
T3407 |
T3392 |
conj |
loci,those |
R914 |
T3408 |
T3407 |
prep |
with,loci |
R915 |
T3205 |
T3194 |
prep |
as,mutation |
R916 |
T3409 |
T3410 |
amod |
altered,action |
R917 |
T3410 |
T3408 |
pobj |
action,with |
R918 |
T3411 |
T3410 |
compound |
gene,action |
R919 |
T3206 |
T3205 |
pobj |
hg,as |
R920 |
T3412 |
T3410 |
amod |
dependent,action |
R921 |
T3413 |
T3412 |
prep |
on,dependent |
R922 |
T3414 |
T3413 |
pobj |
background,on |
R923 |
T3207 |
T3208 |
punct |
[,11 |
R924 |
T3415 |
T3389 |
punct |
.,defined |
R925 |
T3208 |
T3182 |
parataxis |
11,is |
R926 |
T3417 |
T3418 |
nummod |
Four,hg |
R927 |
T3418 |
T3419 |
compound |
hg,QTL |
R928 |
T3419 |
T3421 |
nsubjpass |
QTL,identified |
R929 |
T3420 |
T3419 |
compound |
modifier,QTL |
R930 |
T3209 |
T3208 |
punct |
",",11 |
R931 |
T3422 |
T3419 |
punct |
(,QTL |
R932 |
T3423 |
T3419 |
appos |
Wg2,QTL |
R933 |
T3210 |
T3208 |
appos |
19,11 |
R934 |
T3424 |
T3423 |
prep |
on,Wg2 |
R935 |
T3425 |
T3424 |
pobj |
MMU2,on |
R936 |
T3426 |
T3423 |
punct |
", ",Wg2 |
R937 |
T3211 |
T3212 |
punct |
-,21 |
R938 |
T3427 |
T3423 |
conj |
Carfhg2,Wg2 |
R939 |
T3428 |
T3427 |
prep |
on,Carfhg2 |
R940 |
T3429 |
T3428 |
pobj |
MMU9,on |
R941 |
T3212 |
T3210 |
prep |
21,19 |
R942 |
T3430 |
T3427 |
punct |
", ",Carfhg2 |
R943 |
T3213 |
T3208 |
punct |
],11 |
R944 |
T3214 |
T3182 |
punct |
.,is |
R945 |
T3431 |
T3427 |
conj |
Carp2,Carfhg2 |
R946 |
T3216 |
T3217 |
nsubj |
Epistasis,forms |
R947 |
T3432 |
T3431 |
prep |
on,Carp2 |
R948 |
T3433 |
T3432 |
pobj |
MMU11,on |
R949 |
T3434 |
T3431 |
cc |
and,Carp2 |
R950 |
T3435 |
T3431 |
conj |
Feml3,Carp2 |
R951 |
T3218 |
T3219 |
det |
the,basis |
R952 |
T3436 |
T3435 |
prep |
on,Feml3 |
R953 |
T3437 |
T3436 |
pobj |
MMU17,on |
R954 |
T3438 |
T3419 |
punct |
),QTL |
R955 |
T3219 |
T3217 |
dobj |
basis,forms |
R961 |
T3444 |
T3442 |
pobj |
QTL,with |
R962 |
T3445 |
T3444 |
amod |
additional,QTL |
R963 |
T3537 |
T3536 |
prep |
of,outside |
R964 |
T3446 |
T3444 |
punct |
(,QTL |
R965 |
T3447 |
T3444 |
appos |
Q1Ucd1,QTL |
R966 |
T3448 |
T3447 |
prep |
on,Q1Ucd1 |
R967 |
T3538 |
T3537 |
pobj |
MMU2,of |
R968 |
T3449 |
T3448 |
pobj |
MMU1,on |
R969 |
T3450 |
T3444 |
punct |
(,QTL |
R970 |
T3539 |
T3526 |
punct |
", ",created |
R971 |
T3451 |
T3452 |
dep |
which,reached |
R972 |
T3452 |
T3444 |
relcl |
reached,QTL |
R973 |
T3453 |
T3452 |
advmod |
only,reached |
R974 |
T3540 |
T3541 |
mark |
while,developed |
R975 |
T3454 |
T3455 |
det |
a,level |
R976 |
T3455 |
T3452 |
dobj |
level,reached |
R977 |
T3456 |
T3455 |
amod |
suggestive,level |
R978 |
T3541 |
T3526 |
advcl |
developed,created |
R979 |
T3457 |
T3455 |
prep |
of,level |
R980 |
T3458 |
T3459 |
amod |
statistical,significance |
R981 |
T3459 |
T3457 |
pobj |
significance,of |
R982 |
T3460 |
T3444 |
punct |
),QTL |
R983 |
T3542 |
T3543 |
det |
a,panel |
R984 |
T3461 |
T3444 |
punct |
", ",QTL |
R985 |
T3462 |
T3444 |
appos |
Wg1,QTL |
R986 |
T3543 |
T3541 |
nsubjpass |
panel,developed |
R987 |
T3463 |
T3462 |
punct |
", ",Wg1 |
R988 |
T3464 |
T3462 |
conj |
Carp1,Wg1 |
R989 |
T3465 |
T3464 |
punct |
", ",Carp1 |
R990 |
T3544 |
T3543 |
amod |
comprehensive,panel |
R991 |
T3466 |
T3464 |
conj |
Cara1,Carp1 |
R992 |
T3467 |
T3466 |
cc |
and,Cara1 |
R993 |
T3468 |
T3466 |
conj |
Feml1,Cara1 |
R994 |
T3545 |
T3543 |
prep |
of,panel |
R995 |
T3469 |
T3466 |
prep |
on,Cara1 |
R996 |
T3470 |
T3469 |
pobj |
MMU2,on |
R997 |
T3471 |
T3466 |
punct |
", ",Cara1 |
R998 |
T3546 |
T3547 |
amod |
overlapping,strains |
R999 |
T3472 |
T3466 |
conj |
Carfhg1,Cara1 |
R1000 |
T3473 |
T3472 |
prep |
on,Carfhg1 |
R1001 |
T3474 |
T3473 |
pobj |
MMU5,on |
R1002 |
T3547 |
T3545 |
pobj |
strains,of |
R1003 |
T3475 |
T3472 |
punct |
", ",Carfhg1 |
R1004 |
T3476 |
T3472 |
conj |
Wg3,Carfhg1 |
R1005 |
T3477 |
T3476 |
prep |
on,Wg3 |
R1006 |
T3548 |
T3547 |
prep |
with,strains |
R1007 |
T3478 |
T3477 |
pobj |
MMU8,on |
R1008 |
T3479 |
T3476 |
punct |
", ",Wg3 |
R1009 |
T3480 |
T3476 |
conj |
Feml2,Wg3 |
R1010 |
T3481 |
T3480 |
prep |
on,Feml2 |
R1011 |
T3482 |
T3481 |
pobj |
MMU9,on |
R1012 |
T3483 |
T3480 |
punct |
", ",Feml2 |
R1013 |
T3549 |
T3550 |
amod |
identical,regions |
R1014 |
T3484 |
T3480 |
conj |
Wg4,Feml2 |
R1015 |
T3485 |
T3484 |
cc |
and,Wg4 |
R1016 |
T3486 |
T3484 |
conj |
Cara2,Wg4 |
R1017 |
T3550 |
T3548 |
pobj |
regions,with |
R1018 |
T3487 |
T3484 |
prep |
on,Wg4 |
R1019 |
T3488 |
T3487 |
pobj |
MMU11,on |
R1020 |
T3489 |
T3484 |
cc |
and,Wg4 |
R1021 |
T3490 |
T3484 |
conj |
Carp3,Wg4 |
R1022 |
T3551 |
T3550 |
compound |
donor,regions |
R1023 |
T3491 |
T3490 |
cc |
and,Carp3 |
R1024 |
T3492 |
T3490 |
conj |
Cara3,Carp3 |
R1025 |
T3552 |
T3543 |
prep |
on,panel |
R1026 |
T3493 |
T3490 |
prep |
on,Carp3 |
R1027 |
T3494 |
T3493 |
pobj |
MMU17,on |
R1028 |
T3495 |
T3490 |
punct |
),Carp3 |
R1029 |
T3553 |
T3554 |
preconj |
both,B6 |
R1030 |
T3554 |
T3555 |
nmod |
B6,backgrounds |
R1031 |
T3496 |
T3497 |
poss |
whose,expression |
R1032 |
T3555 |
T3552 |
pobj |
backgrounds,on |
R1033 |
T3497 |
T3498 |
dep |
expression,was |
R1034 |
T3498 |
T3490 |
relcl |
was,Carp3 |
R1035 |
T3499 |
T3498 |
acomp |
independent,was |
R1036 |
T3556 |
T3554 |
cc |
and,B6 |
R1037 |
T3500 |
T3499 |
prep |
of,independent |
R1038 |
T3501 |
T3502 |
det |
the,locus |
R1039 |
T3502 |
T3500 |
pobj |
locus,of |
R1040 |
T3557 |
T3554 |
conj |
HG,B6 |
R1041 |
T3503 |
T3502 |
compound |
hg,locus |
R1042 |
T3504 |
T3421 |
punct |
.,identified |
R1043 |
T3558 |
T3541 |
auxpass |
were,developed |
R1044 |
T3506 |
T3507 |
prep |
In,developed |
R1045 |
T3559 |
T3541 |
prep |
for,developed |
R1046 |
T3508 |
T3509 |
det |
the,study |
R1047 |
T3509 |
T3506 |
pobj |
study,In |
R1048 |
T3510 |
T3509 |
amod |
current,study |
R1049 |
T3511 |
T3507 |
nsubj |
we,developed |
R1050 |
T3560 |
T3559 |
pobj |
MMU2,for |
R1051 |
T3512 |
T3507 |
aux |
have,developed |
R1052 |
T3513 |
T3514 |
npadvmod |
speed,congenic |
R1053 |
T3514 |
T3515 |
amod |
congenic,strains |
R1054 |
T3561 |
T3526 |
punct |
.,created |
R1055 |
T3515 |
T3507 |
dobj |
strains,developed |
R1056 |
T3516 |
T3517 |
aux |
to,isolate |
R1057 |
T3517 |
T3507 |
advcl |
isolate,developed |
R1058 |
T3563 |
T3564 |
det |
The,panel |
R1059 |
T3518 |
T3519 |
predet |
all,QTL |
R1060 |
T3519 |
T3517 |
dobj |
QTL,isolate |
R1061 |
T3520 |
T3519 |
det |
the,QTL |
R1062 |
T3521 |
T3519 |
amod |
aforementioned,QTL |
R1063 |
T3522 |
T3507 |
punct |
.,developed |
R1064 |
T3524 |
T3525 |
amod |
Single,strains |
R1065 |
T3564 |
T3566 |
nsubjpass |
panel,developed |
R1066 |
T3525 |
T3526 |
nsubjpass |
strains,created |
R1067 |
T3527 |
T3525 |
prep |
on,strains |
R1068 |
T3565 |
T3564 |
compound |
MMU2,panel |
R1069 |
T3528 |
T3529 |
det |
an,background |
R1070 |
T3529 |
T3527 |
pobj |
background,on |
R1071 |
T3530 |
T3529 |
compound |
HG,background |
R1072 |
T3567 |
T3566 |
auxpass |
was,developed |
R1073 |
T3531 |
T3526 |
auxpass |
were,created |
R1074 |
T3532 |
T3526 |
prep |
for,created |
R1075 |
T3568 |
T3569 |
aux |
to,confirm |
R1076 |
T3533 |
T3534 |
det |
each,region |
R1077 |
T3569 |
T3566 |
advcl |
confirm,developed |
R1078 |
T3534 |
T3532 |
pobj |
region,for |
R1079 |
T3535 |
T3534 |
amod |
chromosomal,region |
R1080 |
T3536 |
T3534 |
prep |
outside,region |
R1090 |
T3579 |
T3581 |
compound |
hg,interactions |
R1091 |
T3580 |
T3579 |
punct |
-,hg |
R1102 |
T3593 |
T3591 |
pobj |
interactions,of |
R1103 |
T3594 |
T3595 |
compound |
QTL,hg |
R1105 |
T3596 |
T3595 |
punct |
-,hg |
R1114 |
T3605 |
T3607 |
dep |
location,overlapped |
R1115 |
T3606 |
T3605 |
amod |
genomic,location |
R1118 |
T3609 |
T3607 |
dobj |
QTL,overlapped |
R1119 |
T3610 |
T3609 |
compound |
modifier,QTL |
R278 |
T2716 |
T2717 |
det |
The,use |
R279 |
T2717 |
T2718 |
nsubj |
use,provided |
R280 |
T2719 |
T2717 |
prep |
of,use |
R281 |
T2720 |
T2721 |
compound |
mouse,models |
R282 |
T2721 |
T2719 |
pobj |
models,of |
R283 |
T2722 |
T2718 |
aux |
has,provided |
R284 |
T2723 |
T2724 |
amod |
valuable,insight |
R285 |
T2724 |
T2718 |
dobj |
insight,provided |
R286 |
T2725 |
T2724 |
prep |
into,insight |
R287 |
T2726 |
T2727 |
det |
the,etiology |
R288 |
T2727 |
T2725 |
pobj |
etiology,into |
R289 |
T2728 |
T2727 |
prep |
of,etiology |
R290 |
T2729 |
T2730 |
amod |
monogenic,syndromes |
R291 |
T2730 |
T2728 |
pobj |
syndromes,of |
R292 |
T2731 |
T2730 |
acl |
caused,syndromes |
R293 |
T2732 |
T2731 |
agent |
by,caused |
R294 |
T2733 |
T2734 |
amod |
single,mutations |
R297 |
T2736 |
T2718 |
punct |
.,provided |
R298 |
T2738 |
T2739 |
advmod |
However,mimic |
R299 |
T2740 |
T2739 |
punct |
", ",mimic |
R300 |
T2741 |
T2742 |
amod |
such,models |
R301 |
T2742 |
T2739 |
nsubj |
models,mimic |
R302 |
T2743 |
T2739 |
aux |
do,mimic |
R303 |
T2744 |
T2739 |
neg |
not,mimic |
R304 |
T2745 |
T2746 |
det |
the,complexity |
R307 |
T2748 |
T2746 |
prep |
of,complexity |
R308 |
T2749 |
T2750 |
compound |
disease,traits |
R309 |
T2750 |
T2748 |
pobj |
traits,of |
R310 |
T2751 |
T2752 |
advmod |
commonly,seen |
R311 |
T2752 |
T2750 |
acl |
seen,traits |
R312 |
T2753 |
T2752 |
prep |
in,seen |
R313 |
T2754 |
T2755 |
det |
the,population |
R316 |
T2757 |
T2739 |
punct |
.,mimic |
R317 |
T2759 |
T2760 |
compound |
Complex,traits |
R318 |
T2760 |
T2761 |
nsubjpass |
traits,influenced |
R319 |
T2762 |
T2760 |
punct |
", ",traits |
R320 |
T2763 |
T2764 |
amod |
such,as |
R321 |
T2764 |
T2760 |
prep |
as,traits |
R322 |
T2765 |
T2766 |
amod |
polygenic,growth |
R323 |
T2766 |
T2764 |
pobj |
growth,as |
R324 |
T2767 |
T2766 |
cc |
and,growth |
R325 |
T2768 |
T2766 |
conj |
obesity,growth |
R326 |
T2769 |
T2761 |
auxpass |
are,influenced |
R327 |
T2770 |
T2761 |
agent |
by,influenced |
R328 |
T2771 |
T2772 |
det |
the,effects |
R334 |
T2777 |
T2772 |
prep |
of,effects |
R335 |
T2778 |
T2779 |
amod |
quantitative,loci |
R338 |
T2781 |
T2779 |
punct |
(,loci |
R339 |
T2782 |
T2779 |
appos |
QTL,loci |
R340 |
T2783 |
T2779 |
punct |
),loci |
R341 |
T2784 |
T2779 |
punct |
", ",loci |
R342 |
T2785 |
T2779 |
conj |
epistasis,loci |
R343 |
T2786 |
T2785 |
prep |
between,epistasis |
R344 |
T2787 |
T2788 |
compound |
QTL,alleles |
R345 |
T2788 |
T2786 |
pobj |
alleles,between |
R346 |
T2789 |
T2785 |
punct |
", ",epistasis |
R347 |
T2790 |
T2791 |
amod |
environmental,perturbations |
R348 |
T2791 |
T2785 |
conj |
perturbations,epistasis |
R349 |
T2792 |
T2791 |
cc |
and,perturbations |
R350 |
T2793 |
T2794 |
compound |
QTL,environment |
R353 |
T2796 |
T2791 |
conj |
interactions,perturbations |
R354 |
T2797 |
T2761 |
punct |
.,influenced |
R355 |
T2799 |
T2800 |
prep |
To,localized |
R366 |
T2811 |
T2812 |
punct |
[,2 |
R370 |
T2815 |
T2812 |
punct |
],2 |
R371 |
T2816 |
T2810 |
punct |
", ",made |
R372 |
T2817 |
T2810 |
advmod |
however,made |
R373 |
T2818 |
T2810 |
punct |
", ",made |
R374 |
T2819 |
T2820 |
amod |
little,progress |
R375 |
T2820 |
T2810 |
nsubjpass |
progress,made |
R376 |
T2821 |
T2810 |
aux |
has,made |
R377 |
T2822 |
T2810 |
auxpass |
been,made |
R378 |
T2823 |
T2810 |
prep |
in,made |
R379 |
T2824 |
T2823 |
pcomp |
determining,in |
R380 |
T2825 |
T2826 |
det |
the,nature |
R383 |
T2828 |
T2826 |
prep |
of,nature |
R384 |
T2829 |
T2830 |
det |
the,variants |
R388 |
T2833 |
T2810 |
punct |
.,made |
R389 |
T2835 |
T2836 |
nummod |
One,resource |
R390 |
T2836 |
T2837 |
nsubj |
resource,are |
R391 |
T2838 |
T2836 |
acl |
used,resource |
R392 |
T2839 |
T2840 |
aux |
to,map |
R395 |
T2842 |
T2840 |
dobj |
QTL,map |
R396 |
T2843 |
T2844 |
amod |
congenic,strains |
R397 |
T2844 |
T2837 |
attr |
strains,are |
R589 |
T3050 |
T3051 |
det |
a,array |
R592 |
T3053 |
T3051 |
prep |
of,array |
R593 |
T3054 |
T3053 |
pobj |
traits,of |
R594 |
T3055 |
T3051 |
punct |
", ",array |
R612 |
T3070 |
T3068 |
nmod |
hg,mouse |
R670 |
T3135 |
T3126 |
relcl |
shared,mouse |
R671 |
T3136 |
T3137 |
det |
a,number |
R672 |
T3137 |
T3135 |
dobj |
number,shared |
R673 |
T3138 |
T3137 |
prep |
of,number |
R956 |
T3439 |
T3421 |
auxpass |
were,identified |
R957 |
T3440 |
T3421 |
punct |
", ",identified |
R958 |
T3441 |
T3421 |
prep |
along,identified |
R959 |
T3442 |
T3441 |
prep |
with,along |
R960 |
T3443 |
T3444 |
nummod |
12,QTL |
R1081 |
T3570 |
T3571 |
det |
the,presence |
R1082 |
T3571 |
T3569 |
dobj |
presence,confirm |
R1083 |
T3572 |
T3571 |
prep |
of,presence |
R1084 |
T3573 |
T3574 |
amod |
multiple,QTL |
R1085 |
T3574 |
T3572 |
pobj |
QTL,of |
R1086 |
T3575 |
T3569 |
cc |
and,confirm |
R1087 |
T3576 |
T3569 |
conj |
test,confirm |
R1088 |
T3577 |
T3576 |
prep |
for,test |
R1089 |
T3578 |
T3579 |
compound |
QTL,hg |
R1092 |
T3581 |
T3577 |
pobj |
interactions,for |
R1093 |
T3582 |
T3566 |
punct |
.,developed |
R1094 |
T3584 |
T3585 |
advmod |
Additionally,sequenced |
R1095 |
T3586 |
T3585 |
punct |
", ",sequenced |
R1096 |
T3587 |
T3585 |
prep |
based,sequenced |
R1097 |
T3588 |
T3587 |
prep |
on,based |
R1098 |
T3589 |
T3590 |
det |
the,knowledge |
R1099 |
T3590 |
T3588 |
pobj |
knowledge,on |
R1100 |
T3591 |
T3590 |
prep |
of,knowledge |
R1101 |
T3592 |
T3593 |
amod |
potential,interactions |
R1104 |
T3595 |
T3593 |
compound |
hg,interactions |
R1106 |
T3597 |
T3585 |
punct |
", ",sequenced |
R1107 |
T3598 |
T3585 |
nsubjpass |
genes,sequenced |
R1108 |
T3599 |
T3598 |
acl |
involved,genes |
R1109 |
T3600 |
T3599 |
prep |
in,involved |
R1110 |
T3601 |
T3602 |
compound |
Gh,signaling |
R1111 |
T3602 |
T3600 |
pobj |
signaling,in |
R1112 |
T3603 |
T3598 |
punct |
", ",genes |
R1113 |
T3604 |
T3605 |
poss |
whose,location |
R1116 |
T3607 |
T3598 |
relcl |
overlapped,genes |
R1117 |
T3608 |
T3609 |
compound |
hg,QTL |
R1120 |
T3611 |
T3609 |
prep |
on,QTL |
R1121 |
T3612 |
T3613 |
nmod |
MMU,2 |
R1122 |
T3613 |
T3611 |
pobj |
2,on |
R1123 |
T3614 |
T3613 |
punct |
", ",2 |
R1124 |
T3615 |
T3613 |
conj |
9,2 |
R1125 |
T3616 |
T3615 |
punct |
", ",9 |
R1126 |
T3617 |
T3615 |
conj |
11,9 |
R1127 |
T3618 |
T3617 |
cc |
and,11 |
R1128 |
T3619 |
T3617 |
conj |
17,11 |
R1129 |
T3620 |
T3585 |
punct |
", ",sequenced |
R1130 |
T3621 |
T3585 |
auxpass |
were,sequenced |
R1131 |
T3622 |
T3585 |
punct |
.,sequenced |