Id |
Subject |
Object |
Predicate |
Lexical cue |
T10927 |
0-3 |
NN |
denotes |
ATR |
T10928 |
4-5 |
NN |
denotes |
X |
T10929 |
3-4 |
HYPH |
denotes |
- |
T10930 |
6-14 |
NN |
denotes |
syndrome |
T10931 |
15-17 |
VBZ |
denotes |
is |
T10932 |
18-21 |
DT |
denotes |
the |
T10933 |
42-49 |
NN |
denotes |
disease |
T10934 |
22-27 |
JJ |
denotes |
first |
T10935 |
28-33 |
JJ |
denotes |
human |
T10936 |
34-41 |
JJ |
denotes |
genetic |
T10937 |
50-55 |
VBN |
denotes |
known |
T10938 |
56-58 |
TO |
denotes |
to |
T10939 |
62-68 |
VBN |
denotes |
caused |
T10940 |
59-61 |
VB |
denotes |
be |
T10941 |
69-71 |
IN |
denotes |
by |
T10942 |
72-81 |
NNS |
denotes |
mutations |
T10943 |
82-84 |
IN |
denotes |
in |
T10944 |
85-86 |
DT |
denotes |
a |
T10945 |
109-115 |
NN |
denotes |
factor |
T10946 |
87-96 |
NN |
denotes |
chromatin |
T10947 |
97-108 |
NN |
denotes |
remodelling |
T10948 |
115-116 |
. |
denotes |
. |
T10949 |
116-219 |
sentence |
denotes |
At present we do not know how ATRX influences gene expression or what effect it has on cell behaviour. |
T10950 |
117-119 |
IN |
denotes |
At |
T10951 |
138-142 |
VB |
denotes |
know |
T10952 |
120-127 |
JJ |
denotes |
present |
T10953 |
128-130 |
PRP |
denotes |
we |
T10954 |
131-133 |
VBP |
denotes |
do |
T10955 |
134-137 |
RB |
denotes |
not |
T10956 |
143-146 |
WRB |
denotes |
how |
T10957 |
152-162 |
VBZ |
denotes |
influences |
T10958 |
147-151 |
NN |
denotes |
ATRX |
T10959 |
163-167 |
NN |
denotes |
gene |
T10960 |
168-178 |
NN |
denotes |
expression |
T10961 |
179-181 |
CC |
denotes |
or |
T10962 |
182-186 |
WDT |
denotes |
what |
T10963 |
187-193 |
NN |
denotes |
effect |
T10964 |
197-200 |
VBZ |
denotes |
has |
T10965 |
194-196 |
PRP |
denotes |
it |
T10966 |
201-203 |
IN |
denotes |
on |
T10967 |
204-208 |
NN |
denotes |
cell |
T10968 |
209-218 |
NN |
denotes |
behaviour |
T10969 |
218-219 |
. |
denotes |
. |
T10970 |
219-395 |
sentence |
denotes |
Nevertheless, we have previously noted that none of the natural mutations causing ATR-X syndrome are nulls, which suggests that it plays a critical role in normal development. |
T10971 |
220-232 |
RB |
denotes |
Nevertheless |
T10972 |
253-258 |
VBN |
denotes |
noted |
T10973 |
232-234 |
, |
denotes |
, |
T10974 |
234-236 |
PRP |
denotes |
we |
T10975 |
237-241 |
VBP |
denotes |
have |
T10976 |
242-252 |
RB |
denotes |
previously |
T10977 |
259-263 |
IN |
denotes |
that |
T10978 |
317-320 |
VBP |
denotes |
are |
T10979 |
264-268 |
NN |
denotes |
none |
T10980 |
269-271 |
IN |
denotes |
of |
T10981 |
272-275 |
DT |
denotes |
the |
T10982 |
284-293 |
NNS |
denotes |
mutations |
T10983 |
276-283 |
JJ |
denotes |
natural |
T10984 |
294-301 |
VBG |
denotes |
causing |
T10985 |
302-305 |
NN |
denotes |
ATR |
T10986 |
306-307 |
NN |
denotes |
X |
T10987 |
305-306 |
HYPH |
denotes |
- |
T10988 |
308-316 |
NN |
denotes |
syndrome |
T10989 |
321-326 |
NNS |
denotes |
nulls |
T10990 |
326-328 |
, |
denotes |
, |
T10991 |
328-333 |
WDT |
denotes |
which |
T10992 |
334-342 |
VBZ |
denotes |
suggests |
T10993 |
343-347 |
IN |
denotes |
that |
T10994 |
351-356 |
VBZ |
denotes |
plays |
T10995 |
348-350 |
PRP |
denotes |
it |
T10996 |
357-358 |
DT |
denotes |
a |
T10997 |
368-372 |
NN |
denotes |
role |
T10998 |
359-367 |
JJ |
denotes |
critical |
T10999 |
373-375 |
IN |
denotes |
in |
T11000 |
376-382 |
JJ |
denotes |
normal |
T11001 |
383-394 |
NN |
denotes |
development |
T11002 |
394-395 |
. |
denotes |
. |
T11003 |
395-597 |
sentence |
denotes |
Results of conditional inactivation of Atrx in the developing mouse forebrain, based on the Atrx flox allele described here, shows that Atrx exerts a major effect on terminally differentiating neurons. |
T11004 |
396-403 |
NNS |
denotes |
Results |
T11005 |
521-526 |
VBZ |
denotes |
shows |
T11006 |
404-406 |
IN |
denotes |
of |
T11007 |
407-418 |
JJ |
denotes |
conditional |
T11008 |
419-431 |
NN |
denotes |
inactivation |
T11009 |
432-434 |
IN |
denotes |
of |
T11010 |
435-439 |
NN |
denotes |
Atrx |
T11011 |
440-442 |
IN |
denotes |
in |
T11012 |
443-446 |
DT |
denotes |
the |
T11013 |
464-473 |
NN |
denotes |
forebrain |
T11014 |
447-457 |
VBG |
denotes |
developing |
T11015 |
458-463 |
NN |
denotes |
mouse |
T11016 |
473-475 |
, |
denotes |
, |
T11017 |
475-480 |
VBN |
denotes |
based |
T11018 |
481-483 |
IN |
denotes |
on |
T11019 |
484-487 |
DT |
denotes |
the |
T11020 |
498-504 |
NN |
denotes |
allele |
T11021 |
488-497 |
NN |
denotes |
Atrx flox |
T11022 |
505-514 |
VBN |
denotes |
described |
T11023 |
515-519 |
RB |
denotes |
here |
T11024 |
519-521 |
, |
denotes |
, |
T11025 |
527-531 |
IN |
denotes |
that |
T11026 |
537-543 |
VBZ |
denotes |
exerts |
T11027 |
532-536 |
NN |
denotes |
Atrx |
T11028 |
544-545 |
DT |
denotes |
a |
T11029 |
552-558 |
NN |
denotes |
effect |
T11030 |
546-551 |
JJ |
denotes |
major |
T11031 |
559-561 |
IN |
denotes |
on |
T11032 |
562-572 |
RB |
denotes |
terminally |
T11033 |
573-588 |
VBG |
denotes |
differentiating |
T11034 |
589-596 |
NNS |
denotes |
neurons |
T11035 |
596-597 |
. |
denotes |
. |
T11036 |
597-660 |
sentence |
denotes |
Conditional inactivation of Atrx in other tissues is underway. |
T11037 |
598-609 |
JJ |
denotes |
Conditional |
T11038 |
610-622 |
NN |
denotes |
inactivation |
T11039 |
648-650 |
VBZ |
denotes |
is |
T11040 |
623-625 |
IN |
denotes |
of |
T11041 |
626-630 |
NN |
denotes |
Atrx |
T11042 |
631-633 |
IN |
denotes |
in |
T11043 |
634-639 |
JJ |
denotes |
other |
T11044 |
640-647 |
NNS |
denotes |
tissues |
T11045 |
651-659 |
JJ |
denotes |
underway |
T11046 |
659-660 |
. |
denotes |
. |
T11047 |
660-849 |
sentence |
denotes |
Here we have shown that animal-wide disruption of the Atrx gene causes a severe embryonic-lethal phenotype, revealing an essential role for Atrx in the formation of the murine trophoblast. |
T11048 |
661-665 |
RB |
denotes |
Here |
T11049 |
674-679 |
VBN |
denotes |
shown |
T11050 |
666-668 |
PRP |
denotes |
we |
T11051 |
669-673 |
VBP |
denotes |
have |
T11052 |
680-684 |
IN |
denotes |
that |
T11053 |
725-731 |
VBZ |
denotes |
causes |
T11054 |
685-691 |
NN |
denotes |
animal |
T11055 |
692-696 |
JJ |
denotes |
wide |
T11056 |
691-692 |
HYPH |
denotes |
- |
T11057 |
697-707 |
NN |
denotes |
disruption |
T11058 |
708-710 |
IN |
denotes |
of |
T11059 |
711-714 |
DT |
denotes |
the |
T11060 |
720-724 |
NN |
denotes |
gene |
T11061 |
715-719 |
NN |
denotes |
Atrx |
T11062 |
732-733 |
DT |
denotes |
a |
T11063 |
758-767 |
NN |
denotes |
phenotype |
T11064 |
734-740 |
JJ |
denotes |
severe |
T11065 |
741-750 |
JJ |
denotes |
embryonic |
T11066 |
751-757 |
JJ |
denotes |
lethal |
T11067 |
750-751 |
HYPH |
denotes |
- |
T11068 |
767-769 |
, |
denotes |
, |
T11069 |
769-778 |
VBG |
denotes |
revealing |
T11070 |
779-781 |
DT |
denotes |
an |
T11071 |
792-796 |
NN |
denotes |
role |
T11072 |
782-791 |
JJ |
denotes |
essential |
T11073 |
797-800 |
IN |
denotes |
for |
T11074 |
801-805 |
NN |
denotes |
Atrx |
T11075 |
806-808 |
IN |
denotes |
in |
T11076 |
809-812 |
DT |
denotes |
the |
T11077 |
813-822 |
NN |
denotes |
formation |
T11078 |
823-825 |
IN |
denotes |
of |
T11079 |
826-829 |
DT |
denotes |
the |
T11080 |
837-848 |
NN |
denotes |
trophoblast |
T11081 |
830-836 |
JJ |
denotes |
murine |
T11082 |
848-849 |
. |
denotes |
. |
T11083 |
849-980 |
sentence |
denotes |
In addition, Atrx appears to escape imprinted X-chromosome inactivation in the extraembryonic tissues of some carrier female mice. |
T11084 |
850-852 |
IN |
denotes |
In |
T11085 |
868-875 |
VBZ |
denotes |
appears |
T11086 |
853-861 |
NN |
denotes |
addition |
T11087 |
861-863 |
, |
denotes |
, |
T11088 |
863-867 |
NN |
denotes |
Atrx |
T11089 |
876-878 |
TO |
denotes |
to |
T11090 |
879-885 |
VB |
denotes |
escape |
T11091 |
886-895 |
VBN |
denotes |
imprinted |
T11092 |
898-908 |
NN |
denotes |
chromosome |
T11093 |
896-897 |
NN |
denotes |
X |
T11094 |
897-898 |
HYPH |
denotes |
- |
T11095 |
909-921 |
NN |
denotes |
inactivation |
T11096 |
922-924 |
IN |
denotes |
in |
T11097 |
925-928 |
DT |
denotes |
the |
T11098 |
944-951 |
NNS |
denotes |
tissues |
T11099 |
929-943 |
JJ |
denotes |
extraembryonic |
T11100 |
952-954 |
IN |
denotes |
of |
T11101 |
955-959 |
DT |
denotes |
some |
T11102 |
975-979 |
NNS |
denotes |
mice |
T11103 |
960-967 |
NN |
denotes |
carrier |
T11104 |
968-974 |
JJ |
denotes |
female |
T11105 |
979-980 |
. |
denotes |
. |
R6773 |
T10927 |
T10928 |
compound |
ATR,X |
R6774 |
T10928 |
T10930 |
compound |
X,syndrome |
R6775 |
T10929 |
T10928 |
punct |
-,X |
R6776 |
T10930 |
T10931 |
nsubj |
syndrome,is |
R6777 |
T10932 |
T10933 |
det |
the,disease |
R6778 |
T10933 |
T10931 |
attr |
disease,is |
R6779 |
T10934 |
T10933 |
amod |
first,disease |
R6780 |
T10935 |
T10933 |
amod |
human,disease |
R6781 |
T10936 |
T10933 |
amod |
genetic,disease |
R6782 |
T10937 |
T10933 |
acl |
known,disease |
R6783 |
T10938 |
T10939 |
aux |
to,caused |
R6784 |
T10939 |
T10937 |
xcomp |
caused,known |
R6785 |
T10940 |
T10939 |
auxpass |
be,caused |
R6786 |
T10941 |
T10939 |
agent |
by,caused |
R6787 |
T10942 |
T10941 |
pobj |
mutations,by |
R6788 |
T10943 |
T10942 |
prep |
in,mutations |
R6789 |
T10944 |
T10945 |
det |
a,factor |
R6790 |
T10945 |
T10943 |
pobj |
factor,in |
R6791 |
T10946 |
T10947 |
compound |
chromatin,remodelling |
R6792 |
T10947 |
T10945 |
compound |
remodelling,factor |
R6793 |
T10948 |
T10931 |
punct |
.,is |
R6794 |
T10950 |
T10951 |
prep |
At,know |
R6795 |
T10952 |
T10950 |
amod |
present,At |
R6796 |
T10953 |
T10951 |
nsubj |
we,know |
R6797 |
T10954 |
T10951 |
aux |
do,know |
R6798 |
T10955 |
T10951 |
neg |
not,know |
R6799 |
T10956 |
T10957 |
advmod |
how,influences |
R6800 |
T10957 |
T10951 |
advcl |
influences,know |
R6801 |
T10958 |
T10957 |
nsubj |
ATRX,influences |
R6802 |
T10959 |
T10960 |
compound |
gene,expression |
R6803 |
T10960 |
T10957 |
dobj |
expression,influences |
R6804 |
T10961 |
T10957 |
cc |
or,influences |
R6805 |
T10962 |
T10963 |
det |
what,effect |
R6806 |
T10963 |
T10964 |
dep |
effect,has |
R6807 |
T10964 |
T10957 |
conj |
has,influences |
R6808 |
T10965 |
T10964 |
nsubj |
it,has |
R6809 |
T10966 |
T10964 |
prep |
on,has |
R6810 |
T10967 |
T10968 |
compound |
cell,behaviour |
R6811 |
T10968 |
T10966 |
pobj |
behaviour,on |
R6812 |
T10969 |
T10951 |
punct |
.,know |
R6813 |
T10971 |
T10972 |
advmod |
Nevertheless,noted |
R6814 |
T10973 |
T10972 |
punct |
", ",noted |
R6815 |
T10974 |
T10972 |
nsubj |
we,noted |
R6816 |
T10975 |
T10972 |
aux |
have,noted |
R6817 |
T10976 |
T10972 |
advmod |
previously,noted |
R6818 |
T10977 |
T10978 |
mark |
that,are |
R6819 |
T10978 |
T10972 |
ccomp |
are,noted |
R6820 |
T10979 |
T10978 |
nsubj |
none,are |
R6821 |
T10980 |
T10979 |
prep |
of,none |
R6822 |
T10981 |
T10982 |
det |
the,mutations |
R6823 |
T10982 |
T10980 |
pobj |
mutations,of |
R6824 |
T10983 |
T10982 |
amod |
natural,mutations |
R6825 |
T10984 |
T10982 |
acl |
causing,mutations |
R6826 |
T10985 |
T10986 |
compound |
ATR,X |
R6827 |
T10986 |
T10988 |
compound |
X,syndrome |
R6828 |
T10987 |
T10986 |
punct |
-,X |
R6829 |
T10988 |
T10984 |
dobj |
syndrome,causing |
R6830 |
T10989 |
T10978 |
attr |
nulls,are |
R6831 |
T10990 |
T10972 |
punct |
", ",noted |
R6832 |
T10991 |
T10992 |
dep |
which,suggests |
R6833 |
T10992 |
T10972 |
advcl |
suggests,noted |
R6834 |
T10993 |
T10994 |
mark |
that,plays |
R6835 |
T10994 |
T10992 |
ccomp |
plays,suggests |
R6836 |
T10995 |
T10994 |
nsubj |
it,plays |
R6837 |
T10996 |
T10997 |
det |
a,role |
R6838 |
T10997 |
T10994 |
dobj |
role,plays |
R6839 |
T10998 |
T10997 |
amod |
critical,role |
R6840 |
T10999 |
T10994 |
prep |
in,plays |
R6841 |
T11000 |
T11001 |
amod |
normal,development |
R6842 |
T11001 |
T10999 |
pobj |
development,in |
R6843 |
T11002 |
T10972 |
punct |
.,noted |
R6844 |
T11004 |
T11005 |
nsubj |
Results,shows |
R6845 |
T11006 |
T11004 |
prep |
of,Results |
R6846 |
T11007 |
T11008 |
amod |
conditional,inactivation |
R6847 |
T11008 |
T11006 |
pobj |
inactivation,of |
R6848 |
T11009 |
T11008 |
prep |
of,inactivation |
R6849 |
T11010 |
T11009 |
pobj |
Atrx,of |
R6850 |
T11011 |
T11008 |
prep |
in,inactivation |
R6851 |
T11012 |
T11013 |
det |
the,forebrain |
R6852 |
T11013 |
T11011 |
pobj |
forebrain,in |
R6853 |
T11014 |
T11013 |
amod |
developing,forebrain |
R6854 |
T11015 |
T11013 |
compound |
mouse,forebrain |
R6855 |
T11016 |
T11004 |
punct |
", ",Results |
R6856 |
T11017 |
T11004 |
prep |
based,Results |
R6857 |
T11018 |
T11017 |
prep |
on,based |
R6858 |
T11019 |
T11020 |
det |
the,allele |
R6859 |
T11020 |
T11018 |
pobj |
allele,on |
R6860 |
T11021 |
T11020 |
compound |
Atrx flox,allele |
R6861 |
T11022 |
T11020 |
acl |
described,allele |
R6862 |
T11023 |
T11022 |
advmod |
here,described |
R6863 |
T11024 |
T11005 |
punct |
", ",shows |
R6864 |
T11025 |
T11026 |
mark |
that,exerts |
R6865 |
T11026 |
T11005 |
ccomp |
exerts,shows |
R6866 |
T11027 |
T11026 |
nsubj |
Atrx,exerts |
R6867 |
T11028 |
T11029 |
det |
a,effect |
R6868 |
T11029 |
T11026 |
dobj |
effect,exerts |
R6869 |
T11030 |
T11029 |
amod |
major,effect |
R6870 |
T11031 |
T11026 |
prep |
on,exerts |
R6871 |
T11032 |
T11033 |
advmod |
terminally,differentiating |
R6872 |
T11033 |
T11034 |
amod |
differentiating,neurons |
R6873 |
T11034 |
T11031 |
pobj |
neurons,on |
R6874 |
T11035 |
T11005 |
punct |
.,shows |
R6875 |
T11037 |
T11038 |
amod |
Conditional,inactivation |
R6876 |
T11038 |
T11039 |
nsubj |
inactivation,is |
R6877 |
T11040 |
T11038 |
prep |
of,inactivation |
R6878 |
T11041 |
T11040 |
pobj |
Atrx,of |
R6879 |
T11042 |
T11038 |
prep |
in,inactivation |
R6880 |
T11043 |
T11044 |
amod |
other,tissues |
R6881 |
T11044 |
T11042 |
pobj |
tissues,in |
R6882 |
T11045 |
T11039 |
acomp |
underway,is |
R6883 |
T11046 |
T11039 |
punct |
.,is |
R6884 |
T11048 |
T11049 |
advmod |
Here,shown |
R6885 |
T11050 |
T11049 |
nsubj |
we,shown |
R6886 |
T11051 |
T11049 |
aux |
have,shown |
R6887 |
T11052 |
T11053 |
mark |
that,causes |
R6888 |
T11053 |
T11049 |
ccomp |
causes,shown |
R6889 |
T11054 |
T11055 |
npadvmod |
animal,wide |
R6890 |
T11055 |
T11057 |
amod |
wide,disruption |
R6891 |
T11056 |
T11055 |
punct |
-,wide |
R6892 |
T11057 |
T11053 |
nsubj |
disruption,causes |
R6893 |
T11058 |
T11057 |
prep |
of,disruption |
R6894 |
T11059 |
T11060 |
det |
the,gene |
R6895 |
T11060 |
T11058 |
pobj |
gene,of |
R6896 |
T11061 |
T11060 |
compound |
Atrx,gene |
R6897 |
T11062 |
T11063 |
det |
a,phenotype |
R6898 |
T11063 |
T11053 |
dobj |
phenotype,causes |
R6899 |
T11064 |
T11063 |
amod |
severe,phenotype |
R6900 |
T11065 |
T11066 |
amod |
embryonic,lethal |
R6901 |
T11066 |
T11063 |
amod |
lethal,phenotype |
R6902 |
T11067 |
T11066 |
punct |
-,lethal |
R6903 |
T11068 |
T11049 |
punct |
", ",shown |
R6904 |
T11069 |
T11049 |
advcl |
revealing,shown |
R6905 |
T11070 |
T11071 |
det |
an,role |
R6906 |
T11071 |
T11069 |
dobj |
role,revealing |
R6907 |
T11072 |
T11071 |
amod |
essential,role |
R6908 |
T11073 |
T11071 |
prep |
for,role |
R6909 |
T11074 |
T11073 |
pobj |
Atrx,for |
R6910 |
T11075 |
T11071 |
prep |
in,role |
R6911 |
T11076 |
T11077 |
det |
the,formation |
R6912 |
T11077 |
T11075 |
pobj |
formation,in |
R6913 |
T11078 |
T11077 |
prep |
of,formation |
R6914 |
T11079 |
T11080 |
det |
the,trophoblast |
R6915 |
T11080 |
T11078 |
pobj |
trophoblast,of |
R6916 |
T11081 |
T11080 |
amod |
murine,trophoblast |
R6917 |
T11082 |
T11049 |
punct |
.,shown |
R6918 |
T11084 |
T11085 |
prep |
In,appears |
R6919 |
T11086 |
T11084 |
pobj |
addition,In |
R6920 |
T11087 |
T11085 |
punct |
", ",appears |
R6921 |
T11088 |
T11085 |
nsubj |
Atrx,appears |
R6922 |
T11089 |
T11090 |
aux |
to,escape |
R6923 |
T11090 |
T11085 |
xcomp |
escape,appears |
R6924 |
T11091 |
T11092 |
amod |
imprinted,chromosome |
R6925 |
T11092 |
T11095 |
compound |
chromosome,inactivation |
R6926 |
T11093 |
T11092 |
compound |
X,chromosome |
R6927 |
T11094 |
T11092 |
punct |
-,chromosome |
R6928 |
T11095 |
T11090 |
dobj |
inactivation,escape |
R6929 |
T11096 |
T11090 |
prep |
in,escape |
R6930 |
T11097 |
T11098 |
det |
the,tissues |
R6931 |
T11098 |
T11096 |
pobj |
tissues,in |
R6932 |
T11099 |
T11098 |
amod |
extraembryonic,tissues |
R6933 |
T11100 |
T11098 |
prep |
of,tissues |
R6934 |
T11101 |
T11102 |
det |
some,mice |
R6935 |
T11102 |
T11100 |
pobj |
mice,of |
R6936 |
T11103 |
T11102 |
nmod |
carrier,mice |
R6937 |
T11104 |
T11103 |
amod |
female,carrier |
R6938 |
T11105 |
T11085 |
punct |
.,appears |