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An additional observation suggesting that M-opsin and S-opsin are controlled by different mechanisms comes from in vitro experiments [31,32]. While explanted P3 retinas express S-opsin and M-opsin with normal kinetics, explanted P0 retinas express only S-opsin [32]. The factor(s) controlling these differences are unknown, but may be intrinsic, as cocultures of older and younger retinas, conditioned media from older retinas, and addition of a variety of small molecules were unable to promote the expression of M-opsin in the P0-initiated cultures 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