The pleiotropic effects of lec mutations, represented in Fig. 1, suggest that the LEC genes serve as central regulators of embryo and seed development. This prediction was confirmed by showing that LEC1 and FUS3 encode putative transcription factors that regulate critical embryonic processes (12, 19, 20). Here we present evidence suggesting that LEC2 likely encodes a transcriptional regulator with a significant role in embryogenesis.