oronaviruses HCoV genera Coronaviruses Discovery Cellular receptor Natural host(s) α‐Coronaviruses HCoV‐229E 1966 Human aminopeptidase N (CD13) Bats HCoV‐NL63 2004 ACE2 Palm civets, bats β‐Coronaviruses HCoV‐OC43 1967 9‐O‐Acetylated sialic acid Cattle HCoV‐HKU1 2005 9‐O‐Acetylated sialic acid Mice SARS‐CoV‐1 2003 ACE2