J.A.H., W.B.L., J.F.C., C.J.L.-A., and E.P.S. conceived and designed research; J.A.H., W.B.L., and E.P.S. prepared figures; J.A.H., W.B.L., J.F.C., C.J.L.-A., and E.P.S. drafted manuscript; J.A.H., W.B.L., J.F.C., and C.J.L.-A. edited and revised manuscript; J.A.H., W.B.L., J.F.C., C.J.L.-A., and E.P.S. approved final version of manuscript.