Conflict of interest: WJH and YJT declare that they are involved in the licensing of mAbs 1A9, 1G10, 4B12 and 2B2 to commercial companies as research or diagnostic reagents. The other authors have declared that no competing interests exist. Authors’ contributions: Zheng Z – Designed experiments, performed experiments, analysed and organised data, wrote the manuscript. Monteil, VM - Designed experiments, performed experiments, analysed and organised data, wrote the manuscript. Maurer-Stroh, S - Designed experiments, performed experiments, analysed and organised data, wrote the manuscript. Yew CW - Designed experiments, performed experiments. Leong C - Designed experiments, performed experiments. Mohd-Ismail NK - Designed experiments, performed experiments, analysed and organised data, wrote the manuscript. Cheyyatraivendran Arularasu S - Designed experiments, performed experiments. Chow VTK – Analysed and organised data. Lin RTP – Analysed and organised data. Mirazimi A– Designed experiments, performed experiments, analysed and organised data, wrote the manuscript. Hong WJ- Analysed and organised data. Tan YJ- Designed experiments, performed experiments, analysed and organised data, wrote the manuscript.