In phylogenies based on the aa sequences of the entire coding sequences and the NS3 genes, DYPV formed a highly distinct lineage, confirming that it represents a novel pestivirus. Notably, the DYPV lineage occupied the basal position within the Artiodactylous lineage, with those pestiviruses present in rodents and bats forming more divergent lineages. In the phylogenies based on the aa sequences of the RdRp gene and putative RNA methyltransferase gene (VP2; Fig. 5), LSPV also formed a distinct lineage within the genus Coltivirus. Interestingly, however, the virus was most closely related to Shelly headland identified in ticks (I. holocyclus) sampled from the long-nosed bandicoot (a marsupial) in Australia (Harvey et al. 2018).