To further characterize both viruses, primers were designed based on the sequences obtained here and those of related viruses described previously. In this manner we were able to recover the complete genomes of DYPV from 1-Dongyang and the ticks (A. javanense) collected from this animal. The genetic features of DYPV are described in Fig. 2a. Notably, viruses obtained from the pangolin and ticks were closely related to each other, although still exhibited almost 2 per cent nt sequence difference. In addition, both viruses were genetically and phylogenetically distinct from known viruses of the genus Pestivirus, with >43 per cent nt and >48 per cent aa differences (Supplementary Table S5). As with classical pestiviruses, the genome of DYPV encodes twelve proteins. All known cleavage sites of the NS3 protease were observed in DYPV (Supplementary Fig. S3). In addition, the NS4A-NS4B cleavage site, located at Position L2426/A2427 in BVDV1 and L2336/A2337 in CSFV, appeared at Position L2315/K2316 site in DYPV (Supplementary Fig.S3).