RBD (T579N) protein  Binds to receptor DPP4 and MERS-CoV RBD-specific neutralizing mAbs (hMS-1, m336, m337, and m338)  Montanide; Alum  I.M. or S.C.  BALB/c mice; hDPP4-Tg mice  Elicits neutralizing Abs (> 1:3 × 103) in mice against live MERS-CoV (EMC2012 strain)  N/A  Protects all vaccinated hDPP4-Tg mice from challenge of MERS-CoV (EMC2012 strain, 104 TCID50)  Du et al., 2016a