R.B., E.S.G., N.S.L., F.S.C. and L.A.-H. conceived and designed the experiments. R.B., B.A.H., R.M.M., S.D.N., M.B. and O.S. performed the experiments. R.B., B.A.H., R.M.M., S.D.N., E.S.G., N.S.L., F.S.C., L.A.-H. and D.A.N. analyzed the data. R.B., E.S.G., N.S.L., F.S.C., L.A.-H. and D.A.N. wrote the paper.