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Supplementary information
is available for this paper at 10.1038/s41598-020-61700-0.

Author contributions
R.B., E.S.G., N.S.L., F.S.C. and L.A.-H. conceived and designed the experiments. R.B., B.A.H., R.M.M., S.D.N., M.B. and O.S. performed the experiments. R.B., B.A.H., R.M.M., S.D.N., E.S.G., N.S.L., F.S.C., L.A.-H. and D.A.N. analyzed the data. R.B., E.S.G., N.S.L., F.S.C., L.A.-H. and D.A.N. wrote the paper.

Data availability
All data generated or analyzed during this study are included in this published article and its supplementary information files, or they have been placed in public repositories.

Competing interests
The authors declare no competing interests.