In June 2012, the second major human coronavirus, MERS-CoV revealed itself in a sputum sample from a 60-year-old Saudi man who died of overwhelming bilateral pneumonia and renal failure [9]; this Betacoronavirus also has a close relationship with two bat-CoVs (HKU4 and HKU5), and camelids are thought to serve as the intermediates between infected vespertilionid bats and humans [10].