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These authors contributed equally: Yanan Cao, Lin Li

Supplementary information
Supplementary Information accompanies the paper at (10.1038/s41421-020-0147-1).

Acknowledgements
This work is supported by the National Basic Research Program of China (973 Program, 2015CB553601); the National Natural Science Foundation of China (81530020, 81522032).

Author contributions
Y.C. and W.W. conceived the study; L.L., Z.F, S.W., P.H., X.S., F.W., X.H., and Y.C. performed the bioinformatics analysis. G.N. contributed to the discussion of the results. Y.C. and L.L. wrote the manuscript; Y.C. and W.W. revised the manuscript.

Conflict of interest
The authors declare that they have no conflict of interest.