Acknowledgments We greatly appreciate Youyu He (Shanghai Center for Bioinformation Technology) in helping us to blast the insertion sequences against the viral sequence database. Disclosure statement No potential conflict of interest was reported by the author(s). ORCID Xiaojun Li http://orcid.org/0000-0002-5780-0880 Feng Gao http://orcid.org/0000-0001-8903-0203