Funding. YS was supported by a Fondation Méditerranée Infection doctoral position. This work was supported by the French Government under the Investissements d'avenir (Investments for the Future) program managed by the Agence Nationale de la Recherche (reference: Méditerranée Infection 10-IAHU-03) and the Région Provence-Alpes-Côte d'Azur and the European funding FEDER IHU PRIMMI.