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Id	Subject	Object	Predicate	Lexical cue
T4145	0-9	NN	denotes	Evolution
T4146	10-12	IN	denotes	of
T4147	13-16	DT	denotes	the
T4148	29-37	NNS	denotes	segments
T4149	17-28	JJ	denotes	chromosomal
T4150	38-48	VBG	denotes	containing
T4151	49-52	DT	denotes	the
T4152	58-63	NNS	denotes	genes
T4153	53-57	NN	denotes	TACC
T4154	63-159	sentence	denotes	The phylogenetic tree of the FGFR genes closely resembles that of the vertebrate TACC1-3 genes.
T4155	64-67	DT	denotes	The
T4156	81-85	NN	denotes	tree
T4157	68-80	JJ	denotes	phylogenetic
T4158	112-121	VBZ	denotes	resembles
T4159	86-88	IN	denotes	of
T4160	89-92	DT	denotes	the
T4161	98-103	NNS	denotes	genes
T4162	93-97	NN	denotes	FGFR
T4163	104-111	RB	denotes	closely
T4164	122-126	DT	denotes	that
T4165	127-129	IN	denotes	of
T4166	130-133	DT	denotes	the
T4167	153-158	NNS	denotes	genes
T4168	134-144	NN	denotes	vertebrate
T4169	145-150	NN	denotes	TACC1
T4170	150-151	HYPH	denotes	-
T4171	151-152	CD	denotes	3
T4172	158-159	.	denotes	.
T4173	159-435	sentence	denotes	Recently, detailed analyses of the chromosomal regions containing the FGFR gene family in humans, mouse and the arthopod D. melanogaster have revealed the conservation of paralogous chromosomal segments between these organisms (Fig. 2, [13], Table 1 [see Additional file 1]).
T4174	160-168	RB	denotes	Recently
T4175	302-310	VBN	denotes	revealed
T4176	168-170	,	denotes	,
T4177	170-178	JJ	denotes	detailed
T4178	179-187	NNS	denotes	analyses
T4179	188-190	IN	denotes	of
T4180	191-194	DT	denotes	the
T4181	207-214	NNS	denotes	regions
T4182	195-206	JJ	denotes	chromosomal
T4183	215-225	VBG	denotes	containing
T4184	226-229	DT	denotes	the
T4185	240-246	NN	denotes	family
T4186	230-234	NN	denotes	FGFR
T4187	235-239	NN	denotes	gene
T4188	247-249	IN	denotes	in
T4189	250-256	NNS	denotes	humans
T4190	256-258	,	denotes	,
T4191	258-263	NN	denotes	mouse
T4192	264-267	CC	denotes	and
T4193	268-271	DT	denotes	the
T4194	272-280	NN	denotes	arthopod
T4195	281-283	NNP	denotes	D.
T4196	284-296	NNP	denotes	melanogaster
T4197	297-301	VBP	denotes	have
T4198	311-314	DT	denotes	the
T4199	315-327	NN	denotes	conservation
T4200	328-330	IN	denotes	of
T4201	331-341	JJ	denotes	paralogous
T4202	354-362	NNS	denotes	segments
T4203	342-353	JJ	denotes	chromosomal
T4204	363-370	IN	denotes	between
T4205	371-376	DT	denotes	these
T4206	377-386	NNS	denotes	organisms
T4207	387-388	-LRB-	denotes	(
T4208	402-407	NN	denotes	Table
T4209	388-392	NN	denotes	Fig.
T4210	393-394	CD	denotes	2
T4211	394-396	,	denotes	,
T4212	396-397	-LRB-	denotes	[
T4213	397-399	CD	denotes	13
T4214	399-400	-RRB-	denotes	]
T4215	400-402	,	denotes	,
T4216	408-409	CD	denotes	1
T4217	410-411	-LRB-	denotes	[
T4218	411-414	VB	denotes	see
T4219	415-425	JJ	denotes	Additional
T4220	426-430	NN	denotes	file
T4221	431-432	CD	denotes	1
T4222	432-433	-RRB-	denotes	]
T4223	433-434	-RRB-	denotes	)
T4224	434-435	.	denotes	.
T4225	435-744	sentence	denotes	This has provided further support that an ancient chromosomal segment was duplicated twice during vertebrate evolution, with the first duplication that gave rise to the human chromosome 4p16/5q32-ter and human chromosome 8p/10q23-ter ancestors occurring in the early stages after the invertebrate divergence.
T4226	436-440	DT	denotes	This
T4227	445-453	VBN	denotes	provided
T4228	441-444	VBZ	denotes	has
T4229	454-461	JJ	denotes	further
T4230	462-469	NN	denotes	support
T4231	470-474	IN	denotes	that
T4232	510-520	VBN	denotes	duplicated
T4233	475-477	DT	denotes	an
T4234	498-505	NN	denotes	segment
T4235	478-485	JJ	denotes	ancient
T4236	486-497	JJ	denotes	chromosomal
T4237	506-509	VBD	denotes	was
T4238	521-526	RB	denotes	twice
T4239	527-533	IN	denotes	during
T4240	534-544	NN	denotes	vertebrate
T4241	545-554	NN	denotes	evolution
T4242	554-556	,	denotes	,
T4243	556-560	IN	denotes	with
T4244	561-564	DT	denotes	the
T4245	571-582	NN	denotes	duplication
T4246	565-570	JJ	denotes	first
T4247	680-689	VBG	denotes	occurring
T4248	583-587	WDT	denotes	that
T4249	588-592	VBD	denotes	gave
T4250	593-597	NN	denotes	rise
T4251	598-600	IN	denotes	to
T4252	601-604	DT	denotes	the
T4253	670-679	NNS	denotes	ancestors
T4254	605-610	JJ	denotes	human
T4255	611-621	NN	denotes	chromosome
T4256	632-635	NN	denotes	ter
T4257	622-626	NN	denotes	4p16
T4258	627-631	NN	denotes	5q32
T4259	626-627	HYPH	denotes	/
T4260	631-632	HYPH	denotes	-
T4261	636-639	CC	denotes	and
T4262	640-645	JJ	denotes	human
T4263	646-656	NN	denotes	chromosome
T4264	666-669	NN	denotes	ter
T4265	657-659	NN	denotes	8p
T4266	660-665	NN	denotes	10q23
T4267	659-660	HYPH	denotes	/
T4268	665-666	HYPH	denotes	-
T4269	690-692	IN	denotes	in
T4270	693-696	DT	denotes	the
T4271	703-709	NNS	denotes	stages
T4272	697-702	JJ	denotes	early
T4273	710-715	IN	denotes	after
T4274	716-719	DT	denotes	the
T4275	733-743	NN	denotes	divergence
T4276	720-732	NN	denotes	invertebrate
T4277	743-744	.	denotes	.
T4278	744-962	sentence	denotes	This suggests that the ancestral FGFR-TACC gene pair most probably arose prior to the initial duplication and subsequent divergence of these paralogous chromosomal segments, estimated to have occurred 687 ± 155.7 MYA.
T4279	745-749	DT	denotes	This
T4280	750-758	VBZ	denotes	suggests
T4281	759-763	IN	denotes	that
T4282	812-817	VBD	denotes	arose
T4283	764-767	DT	denotes	the
T4284	793-797	NN	denotes	pair
T4285	768-777	JJ	denotes	ancestral
T4286	778-782	NN	denotes	FGFR
T4287	783-787	NN	denotes	TACC
T4288	782-783	HYPH	denotes	-
T4289	788-792	NN	denotes	gene
T4290	798-802	RBS	denotes	most
T4291	803-811	RB	denotes	probably
T4292	818-823	JJ	denotes	prior
T4293	824-826	IN	denotes	to
T4294	827-830	DT	denotes	the
T4295	839-850	NN	denotes	duplication
T4296	831-838	JJ	denotes	initial
T4297	851-854	CC	denotes	and
T4298	855-865	JJ	denotes	subsequent
T4299	866-876	NN	denotes	divergence
T4300	877-879	IN	denotes	of
T4301	880-885	DT	denotes	these
T4302	909-917	NNS	denotes	segments
T4303	886-896	JJ	denotes	paralogous
T4304	897-908	JJ	denotes	chromosomal
T4305	917-919	,	denotes	,
T4306	919-928	VBN	denotes	estimated
T4307	929-931	TO	denotes	to
T4308	937-945	VBN	denotes	occurred
T4309	932-936	VB	denotes	have
T4310	946-949	CD	denotes	687
T4311	952-957	CD	denotes	155.7
T4312	950-951	SYM	denotes	±
T4313	958-961	RB	denotes	MYA
T4314	961-962	.	denotes	.
T4315	962-1086	sentence	denotes	This has raised the suggestion that a fourth TACC gene in vertebrates would reside in the same chromosomal region as FGFR4.
T4316	963-967	DT	denotes	This
T4317	972-978	VBN	denotes	raised
T4318	968-971	VBZ	denotes	has
T4319	979-982	DT	denotes	the
T4320	983-993	NN	denotes	suggestion
T4321	994-998	IN	denotes	that
T4322	1039-1045	VB	denotes	reside
T4323	999-1000	DT	denotes	a
T4324	1013-1017	NN	denotes	gene
T4325	1001-1007	JJ	denotes	fourth
T4326	1008-1012	NN	denotes	TACC
T4327	1018-1020	IN	denotes	in
T4328	1021-1032	NNS	denotes	vertebrates
T4329	1033-1038	MD	denotes	would
T4330	1046-1048	IN	denotes	in
T4331	1049-1052	DT	denotes	the
T4332	1070-1076	NN	denotes	region
T4333	1053-1057	JJ	denotes	same
T4334	1058-1069	JJ	denotes	chromosomal
T4335	1077-1079	IN	denotes	as
T4336	1080-1085	NN	denotes	FGFR4
T4337	1085-1086	.	denotes	.
T4338	1086-1190	sentence	denotes	Indeed this hypothesis has been used in support for the RHAMM gene as a member of the TACC family [10].
T4339	1087-1093	RB	denotes	Indeed
T4340	1119-1123	VBN	denotes	used
T4341	1094-1098	DT	denotes	this
T4342	1099-1109	NN	denotes	hypothesis
T4343	1110-1113	VBZ	denotes	has
T4344	1114-1118	VBN	denotes	been
T4345	1124-1126	IN	denotes	in
T4346	1127-1134	NN	denotes	support
T4347	1135-1138	IN	denotes	for
T4348	1139-1142	DT	denotes	the
T4349	1149-1153	NN	denotes	gene
T4350	1143-1148	NN	denotes	RHAMM
T4351	1154-1156	IN	denotes	as
T4352	1157-1158	DT	denotes	a
T4353	1159-1165	NN	denotes	member
T4354	1166-1168	IN	denotes	of
T4355	1169-1172	DT	denotes	the
T4356	1178-1184	NN	denotes	family
T4357	1173-1177	NN	denotes	TACC
T4358	1185-1186	-LRB-	denotes	[
T4359	1186-1188	CD	denotes	10
T4360	1188-1189	-RRB-	denotes	]
T4361	1189-1190	.	denotes	.
T4362	1190-1265	sentence	denotes	Human RHAMM maps to chromosome 5q32 in a region bounded by GPX3 and NKX2E.
T4363	1191-1196	JJ	denotes	Human
T4364	1197-1202	NN	denotes	RHAMM
T4365	1203-1207	VBZ	denotes	maps
T4366	1208-1210	IN	denotes	to
T4367	1211-1221	NN	denotes	chromosome
T4368	1222-1226	NN	denotes	5q32
T4369	1227-1229	IN	denotes	in
T4370	1230-1231	DT	denotes	a
T4371	1232-1238	NN	denotes	region
T4372	1239-1246	VBN	denotes	bounded
T4373	1247-1249	IN	denotes	by
T4374	1250-1254	NN	denotes	GPX3
T4375	1255-1258	CC	denotes	and
T4376	1259-1264	NN	denotes	NKX2E
T4377	1264-1265	.	denotes	.
T4378	1265-1360	sentence	denotes	These loci separate two clusters of genes on human chromosome 5 that are paralogous with 4p16.
T4379	1266-1271	DT	denotes	These
T4380	1272-1276	NNS	denotes	loci
T4381	1277-1285	VBP	denotes	separate
T4382	1286-1289	CD	denotes	two
T4383	1290-1298	NNS	denotes	clusters
T4384	1299-1301	IN	denotes	of
T4385	1302-1307	NNS	denotes	genes
T4386	1308-1310	IN	denotes	on
T4387	1311-1316	JJ	denotes	human
T4388	1317-1327	NN	denotes	chromosome
T4389	1328-1329	CD	denotes	5
T4390	1330-1334	WDT	denotes	that
T4391	1335-1338	VBP	denotes	are
T4392	1339-1349	JJ	denotes	paralogous
T4393	1350-1354	IN	denotes	with
T4394	1355-1359	CD	denotes	4p16
T4395	1359-1360	.	denotes	.
T4396	1360-1575	sentence	denotes	Interestingly, these three clusters are located on different chromosomes in mouse and rat (Fig. 2), further suggesting that this cluster of genes was transposed into this region after the primate/rodent divergence.
T4397	1361-1374	RB	denotes	Interestingly
T4398	1401-1408	VBN	denotes	located
T4399	1374-1376	,	denotes	,
T4400	1376-1381	DT	denotes	these
T4482	2489-2490	``	denotes	"
T4483	2499-2500	''	denotes	"
T4484	2501-2508	JJ	denotes	compact
T4485	2509-2519	NN	denotes	vertebrate
T4486	2527-2536	VBN	denotes	sequenced
T4487	2537-2539	IN	denotes	to
T4488	2540-2544	NN	denotes	date
T4489	2544-2546	,	denotes	,
T4490	2546-2548	NNP	denotes	T.
T4491	2549-2557	NNP	denotes	rubripes
T4492	2558-2559	-LRB-	denotes	[
T4493	2559-2561	CD	denotes	14
T4494	2561-2562	-RRB-	denotes	]
T4495	2562-2563	.	denotes	.
T4496	2563-2739	sentence	denotes	Although not fully assembled, examination of the genome of T. rubripes confirmed the presence of chromosomal segments paralogous to those found in higher vertebrates (Fig. 2).
T4497	2564-2572	IN	denotes	Although
T4498	2583-2592	VBN	denotes	assembled
T4499	2573-2576	RB	denotes	not
T4500	2577-2582	RB	denotes	fully
T4501	2635-2644	VBD	denotes	confirmed
T4502	2592-2594	,	denotes	,
T4503	2594-2605	NN	denotes	examination
T4504	2606-2608	IN	denotes	of
T4505	2609-2612	DT	denotes	the
T4506	2613-2619	NN	denotes	genome
T4507	2620-2622	IN	denotes	of
T4508	2623-2625	NNP	denotes	T.
T4509	2626-2634	NNP	denotes	rubripes
T4510	2645-2648	DT	denotes	the
T4511	2649-2657	NN	denotes	presence
T4512	2658-2660	IN	denotes	of
T4513	2661-2672	JJ	denotes	chromosomal
T4514	2673-2681	NNS	denotes	segments
T4515	2682-2692	JJ	denotes	paralogous
T4516	2693-2695	IN	denotes	to
T4517	2696-2701	DT	denotes	those
T4518	2702-2707	VBN	denotes	found
T4519	2708-2710	IN	denotes	in
T4520	2711-2717	JJR	denotes	higher
T4521	2718-2729	NNS	denotes	vertebrates
T4522	2730-2731	-LRB-	denotes	(
T4523	2731-2735	NN	denotes	Fig.
T4524	2736-2737	CD	denotes	2
T4525	2737-2738	-RRB-	denotes	)
T4526	2738-2739	.	denotes	.
T4527	2739-2896	sentence	denotes	For instance, the orthologues of GPRK2L and RGS12 are found on T. rubripes scaffold 290 (emb\|CAAB01000290.1), and within 300 kb of each other in human 4p16.
T4528	2740-2743	IN	denotes	For
T4529	2794-2799	VBN	denotes	found
T4530	2744-2752	NN	denotes	instance
T4531	2752-2754	,	denotes	,
T4532	2754-2757	DT	denotes	the
T4533	2758-2769	NNS	denotes	orthologues
T4534	2770-2772	IN	denotes	of
T4535	2773-2779	NN	denotes	GPRK2L
T4536	2780-2783	CC	denotes	and
T4537	2784-2789	NN	denotes	RGS12
T4538	2790-2793	VBP	denotes	are
T4539	2800-2802	IN	denotes	on
T4540	2803-2805	NNP	denotes	T.
T4541	2806-2814	NNP	denotes	rubripes
T4542	2815-2823	NN	denotes	scaffold
T4543	2824-2827	CD	denotes	290
T4544	2828-2829	-LRB-	denotes	(
T4545	2829-2847	NN	denotes	emb\|CAAB01000290.1
T4546	2847-2848	-RRB-	denotes	)
T4547	2848-2850	,	denotes	,
T4548	2850-2853	CC	denotes	and
T4549	2854-2860	IN	denotes	within
T4550	2861-2864	CD	denotes	300
T4551	2865-2867	NN	denotes	kb
T4552	2868-2870	IN	denotes	of
T4553	2871-2875	DT	denotes	each
T4554	2876-2881	JJ	denotes	other
T4555	2882-2884	IN	denotes	in
T4556	2885-2890	JJ	denotes	human
T4557	2891-2895	CD	denotes	4p16
T4558	2895-2896	.	denotes	.
T4559	2896-3024	sentence	denotes	The T. rubripes orthologues of FGFR3, LETM1 and WHSC1 are located on the same 166 kb genomic scaffold 251 (emb\|CAAB01000166.1).
T4560	2897-2900	DT	denotes	The
T4561	2913-2924	NNS	denotes	orthologues
T4562	2901-2903	NNP	denotes	T.
T4563	2904-2912	NNP	denotes	rubripes
T4564	2955-2962	VBN	denotes	located
T4565	2925-2927	IN	denotes	of
T4566	2928-2933	NN	denotes	FGFR3
T4567	2933-2935	,	denotes	,
T4568	2935-2940	NN	denotes	LETM1
T4569	2941-2944	CC	denotes	and
T4570	2945-2950	NN	denotes	WHSC1
T4571	2951-2954	VBP	denotes	are
T4572	2963-2965	IN	denotes	on
T4573	2966-2969	DT	denotes	the
T4574	2990-2998	NN	denotes	scaffold
T4575	2970-2974	JJ	denotes	same
T4576	2975-2978	CD	denotes	166
T4577	2979-2981	NN	denotes	kb
T4578	2982-2989	JJ	denotes	genomic
T4579	2999-3002	CD	denotes	251
T4580	3003-3004	-LRB-	denotes	(
T4581	3004-3022	NN	denotes	emb\|CAAB01000166.1
T4582	3022-3023	-RRB-	denotes	)
T4583	3023-3024	.	denotes	.
T4584	3024-3136	sentence	denotes	Significantly, the three human orthologues of these genes are also located within 300 kb of each other on 4p16.
T4585	3025-3038	RB	denotes	Significantly
T4586	3092-3099	VBN	denotes	located
T4587	3038-3040	,	denotes	,
T4588	3040-3043	DT	denotes	the
T4589	3056-3067	NNS	denotes	orthologues
T4590	3044-3049	CD	denotes	three
T4591	3050-3055	JJ	denotes	human
T4592	3068-3070	IN	denotes	of
T4593	3071-3076	DT	denotes	these
T4594	3077-3082	NNS	denotes	genes
T4595	3083-3086	VBP	denotes	are
T4596	3087-3091	RB	denotes	also
T4597	3100-3106	IN	denotes	within
T4598	3107-3110	CD	denotes	300
T4599	3111-3113	NN	denotes	kb
T4600	3114-3116	IN	denotes	of
T4601	3117-3121	DT	denotes	each
T4602	3122-3127	JJ	denotes	other
T4603	3128-3130	IN	denotes	on
T4604	3131-3135	NN	denotes	4p16
T4605	3135-3136	.	denotes	.
T4606	3136-3228	sentence	denotes	Furthermore, TACC3 and FGFRL map to the overlapping scaffolds 1184/4669 (emb\|CAAB01004668).
T4607	3137-3148	RB	denotes	Furthermore
T4608	3166-3169	VBP	denotes	map
T4609	3148-3150	,	denotes	,
T4610	3150-3155	NN	denotes	TACC3
T4611	3156-3159	CC	denotes	and
T4612	3160-3165	NN	denotes	FGFRL
T4613	3170-3172	IN	denotes	to
T4614	3173-3176	DT	denotes	the
T4615	3189-3198	NNS	denotes	scaffolds
T4616	3177-3188	VBG	denotes	overlapping
T4617	3199-3203	CD	denotes	1184
T4618	3204-3208	CD	denotes	4669
T4619	3203-3204	HYPH	denotes	/
T4620	3209-3210	-LRB-	denotes	(
T4621	3210-3226	NN	denotes	emb\|CAAB01004668
T4622	3226-3227	-RRB-	denotes	)
T4623	3227-3228	.	denotes	.
T4624	3228-3477	sentence	denotes	Similarly, elements of these gene clusters, extending from HMP19 to GPRK6 in human chromosome 5q34-ter are also found in the pufferfish, with the T. rubripes orthologues of NSD1, FGFR4 and a RAB-like gene mapping on scaffold 407 (emb\|CAAB01000407).
T4625	3229-3238	RB	denotes	Similarly
T4626	3341-3346	VBN	denotes	found
T4627	3238-3240	,	denotes	,
T4628	3240-3248	NNS	denotes	elements
T4629	3249-3251	IN	denotes	of
T4630	3252-3257	DT	denotes	these
T4631	3263-3271	NNS	denotes	clusters
T4632	3258-3262	NN	denotes	gene
T4633	3271-3273	,	denotes	,
T4634	3273-3282	VBG	denotes	extending
T4635	3283-3287	IN	denotes	from
T4636	3288-3293	NN	denotes	HMP19
T4637	3294-3296	IN	denotes	to
T4638	3297-3302	NN	denotes	GPRK6
T4639	3303-3305	IN	denotes	in
T4640	3306-3311	JJ	denotes	human
T4641	3312-3322	NN	denotes	chromosome
T4642	3328-3331	NN	denotes	ter
T4643	3323-3327	NN	denotes	5q34
T4644	3327-3328	HYPH	denotes	-
T4645	3332-3335	VBP	denotes	are
T4646	3336-3340	RB	denotes	also
T4647	3347-3349	IN	denotes	in
T4648	3350-3353	DT	denotes	the
T4649	3354-3364	NN	denotes	pufferfish
T4650	3364-3366	,	denotes	,
T4651	3366-3370	IN	denotes	with
T4652	3371-3374	DT	denotes	the
T4653	3387-3398	NNS	denotes	orthologues
T4654	3375-3377	NNP	denotes	T.
T4655	3378-3386	NNP	denotes	rubripes
T4656	3399-3401	IN	denotes	of
T4657	3402-3406	NN	denotes	NSD1
T4658	3406-3408	,	denotes	,
T4659	3408-3413	NN	denotes	FGFR4
T4660	3414-3417	CC	denotes	and
T4661	3418-3419	DT	denotes	a
T4662	3424-3428	JJ	denotes	like
T4663	3420-3423	NN	denotes	RAB
T4664	3423-3424	HYPH	denotes	-
T4665	3434-3441	NN	denotes	mapping
T4666	3429-3433	NN	denotes	gene
T4667	3442-3444	IN	denotes	on
T4668	3445-3453	NN	denotes	scaffold
T4669	3454-3457	CD	denotes	407
T4670	3458-3459	-LRB-	denotes	(
T4671	3459-3475	NN	denotes	emb\|CAAB01000407
T4672	3475-3476	-RRB-	denotes	)
T4673	3476-3477	.	denotes	.
T4674	3477-3574	sentence	denotes	However, there is no evidence for a gene corresponding to a TACC4 gene in any of these clusters.
T4675	3478-3485	RB	denotes	However
T4676	3493-3495	VBZ	denotes	is
T4677	3485-3487	,	denotes	,
T4678	3487-3492	EX	denotes	there
T4679	3496-3498	DT	denotes	no
T4680	3499-3507	NN	denotes	evidence
T4681	3508-3511	IN	denotes	for
T4682	3512-3513	DT	denotes	a
T4683	3514-3518	NN	denotes	gene
T4684	3519-3532	VBG	denotes	corresponding
T4685	3533-3535	IN	denotes	to
T4686	3536-3537	DT	denotes	a
T4687	3544-3548	NN	denotes	gene
T4688	3538-3543	NN	denotes	TACC4
T4689	3549-3551	IN	denotes	in
T4690	3552-3555	DT	denotes	any
T4691	3556-3558	IN	denotes	of
T4692	3559-3564	DT	denotes	these
T4693	3565-3573	NNS	denotes	clusters
T4694	3573-3574	.	denotes	.
T4695	3574-3701	sentence	denotes	As noted above, phylogenetic analysis of the TACC sequences indicate that there are two TACC1 related genes in the pufferfish.
T4696	3575-3577	IN	denotes	As
T4697	3578-3583	VBN	denotes	noted
T4698	3635-3643	VBP	denotes	indicate
T4699	3584-3589	RB	denotes	above
T4700	3589-3591	,	denotes	,
T4701	3591-3603	JJ	denotes	phylogenetic
T4702	3604-3612	NN	denotes	analysis
T4703	3613-3615	IN	denotes	of
T4704	3616-3619	DT	denotes	the
T4705	3625-3634	NNS	denotes	sequences
T4706	3620-3624	NN	denotes	TACC
T4707	3644-3648	IN	denotes	that
T4708	3655-3658	VBP	denotes	are
T4709	3649-3654	EX	denotes	there
T4710	3659-3662	CD	denotes	two
T4711	3677-3682	NNS	denotes	genes
T4712	3663-3668	NN	denotes	TACC1
T4713	3669-3676	VBN	denotes	related
T4714	3683-3685	IN	denotes	in
T4715	3686-3689	DT	denotes	the
T4716	3690-3700	NN	denotes	pufferfish
T4717	3700-3701	.	denotes	.
T4718	3701-3860	sentence	denotes	trTACC1B is located on the 180 kb scaffold 191 (emb\|CAAB01000191.1), which also contains the orthologues of several genes located in human chromosome 8p21-11.
T4719	3702-3710	NN	denotes	trTACC1B
T4720	3714-3721	VBN	denotes	located
T4721	3711-3713	VBZ	denotes	is
T4722	3722-3724	IN	denotes	on
T4723	3725-3728	DT	denotes	the
T4724	3736-3744	NN	denotes	scaffold
T4725	3729-3732	CD	denotes	180
T4726	3733-3735	NN	denotes	kb
T4727	3745-3748	CD	denotes	191
T4728	3749-3750	-LRB-	denotes	(
T4729	3750-3768	NN	denotes	emb\|CAAB01000191.1
T4730	3768-3769	-RRB-	denotes	)
T4731	3769-3771	,	denotes	,
T4732	3771-3776	WDT	denotes	which
T4733	3782-3790	VBZ	denotes	contains
T4734	3777-3781	RB	denotes	also
T4735	3791-3794	DT	denotes	the
T4736	3795-3806	NNS	denotes	orthologues
T4737	3807-3809	IN	denotes	of
T4738	3810-3817	JJ	denotes	several
T4739	3818-3823	NNS	denotes	genes
T4740	3824-3831	VBN	denotes	located
T4741	3832-3834	IN	denotes	in
T4742	3835-3840	JJ	denotes	human
T4743	3852-3856	NN	denotes	8p21
T4744	3841-3851	NN	denotes	chromosome
T4745	3856-3857	HYPH	denotes	-
T4746	3857-3859	CD	denotes	11
T4747	3859-3860	.	denotes	.
T4748	3860-3977	sentence	denotes	Thus, this scaffold represents the more "developed" TACC1 chromosomal segment that is evident in higher vertebrates.
T4749	3861-3865	RB	denotes	Thus
T4750	3881-3891	VBZ	denotes	represents
T4751	3865-3867	,	denotes	,
T4752	3867-3871	DT	denotes	this
T4753	3872-3880	NN	denotes	scaffold
T4754	3892-3895	DT	denotes	the
T4755	3931-3938	NN	denotes	segment
T4756	3896-3900	RBR	denotes	more
T4757	3902-3911	VBN	denotes	developed
T4758	3901-3902	``	denotes	"
T4759	3911-3912	''	denotes	"
T4760	3913-3918	NN	denotes	TACC1
T4761	3919-3930	JJ	denotes	chromosomal
T4762	3939-3943	WDT	denotes	that
T4763	3944-3946	VBZ	denotes	is
T4764	3947-3954	JJ	denotes	evident
T4765	3955-3957	IN	denotes	in
T4766	3958-3964	JJR	denotes	higher
T4767	3965-3976	NNS	denotes	vertebrates
T4768	3976-3977	.	denotes	.
T4769	3977-4071	sentence	denotes	On the other hand, the trTACC1A gene is located in the 396 kb scaffold 12 (emb\|CAAB010012.1).
T4770	3978-3980	IN	denotes	On
T4771	4018-4025	VBN	denotes	located
T4772	3981-3984	DT	denotes	the
T4773	3991-3995	NN	denotes	hand
T4774	3985-3990	JJ	denotes	other
T4775	3995-3997	,	denotes	,
T4776	3997-4000	DT	denotes	the
T4777	4010-4014	NN	denotes	gene
T4778	4001-4009	NN	denotes	trTACC1A
T4779	4015-4017	VBZ	denotes	is
T4780	4026-4028	IN	denotes	in
T4781	4029-4032	DT	denotes	the
T4782	4040-4048	NN	denotes	scaffold
T4783	4033-4036	CD	denotes	396
T4784	4037-4039	NN	denotes	kb
T4785	4049-4051	CD	denotes	12
T4786	4052-4053	-LRB-	denotes	(
T4787	4053-4069	NN	denotes	emb\|CAAB010012.1
T4788	4069-4070	-RRB-	denotes	)
T4789	4070-4071	.	denotes	.
T4790	4071-4269	sentence	denotes	This scaffold also contains the T. rubripes orthologues of MSX1, STX18, D4S234E and the predicted gene LOC118711, in addition to sequences with homology to LOXL, EVC, LOC159291, and the LDB family.
T4791	4072-4076	DT	denotes	This
T4792	4077-4085	NN	denotes	scaffold
T4793	4091-4099	VBZ	denotes	contains
T4794	4086-4090	RB	denotes	also
T4795	4100-4103	DT	denotes	the
T4796	4116-4127	NNS	denotes	orthologues
T4797	4104-4106	NNP	denotes	T.
T4798	4107-4115	NNP	denotes	rubripes
T4799	4128-4130	IN	denotes	of
T4800	4131-4135	NN	denotes	MSX1
T4801	4135-4137	,	denotes	,
T4802	4137-4142	NN	denotes	STX18
T4803	4142-4144	,	denotes	,
T4804	4144-4151	NN	denotes	D4S234E
T4805	4152-4155	CC	denotes	and
T4806	4156-4159	DT	denotes	the
T4807	4170-4174	NN	denotes	gene
T4808	4160-4169	VBN	denotes	predicted
T4809	4175-4184	NN	denotes	LOC118711
T4810	4184-4186	,	denotes	,
T4811	4186-4188	IN	denotes	in
T4812	4189-4197	NN	denotes	addition
T4813	4198-4200	IN	denotes	to
T4814	4201-4210	NNS	denotes	sequences
T4815	4211-4215	IN	denotes	with
T4816	4216-4224	NN	denotes	homology
T4817	4225-4227	IN	denotes	to
T4818	4228-4232	NN	denotes	LOXL
T4819	4232-4234	,	denotes	,
T4820	4234-4237	NN	denotes	EVC
T4821	4237-4239	,	denotes	,
T4822	4239-4248	NN	denotes	LOC159291
T4823	4248-4250	,	denotes	,
T4824	4250-4253	CC	denotes	and
T4825	4254-4257	DT	denotes	the
T4826	4262-4268	NN	denotes	family
T4827	4258-4261	NN	denotes	LDB
T4828	4268-4269	.	denotes	.
T4829	4269-4567	sentence	denotes	Thus, scaffold 12 contains genes found in the regions of human chromosome 4 and 10 that also contain the loci for TACC3 and TACC2, respectively, and may therefore more closely resemble the genomic organization resulting from the initial duplication of the ancestral paralogous chromosomal segment.
T4830	4270-4274	RB	denotes	Thus
T4831	4288-4296	VBZ	denotes	contains
T4832	4274-4276	,	denotes	,
T4833	4276-4284	NN	denotes	scaffold
T4834	4285-4287	CD	denotes	12
T4835	4297-4302	NNS	denotes	genes
T4836	4303-4308	VBN	denotes	found
T4837	4309-4311	IN	denotes	in
T4838	4312-4315	DT	denotes	the
T4839	4316-4323	NNS	denotes	regions
T4840	4324-4326	IN	denotes	of
T4841	4327-4332	JJ	denotes	human
T4842	4344-4345	CD	denotes	4
T4843	4333-4343	NN	denotes	chromosome
T4844	4346-4349	CC	denotes	and
T4845	4350-4352	CD	denotes	10
T4846	4353-4357	WDT	denotes	that
T4847	4363-4370	VBP	denotes	contain
T4848	4358-4362	RB	denotes	also
T4849	4371-4374	DT	denotes	the
T4850	4375-4379	NNS	denotes	loci
T4851	4380-4383	IN	denotes	for
T4852	4384-4389	NN	denotes	TACC3
T4853	4390-4393	CC	denotes	and
T4854	4394-4399	NN	denotes	TACC2
T4855	4399-4401	,	denotes	,
T4856	4401-4413	RB	denotes	respectively
T4857	4413-4415	,	denotes	,
T4858	4415-4418	CC	denotes	and
T4859	4419-4422	MD	denotes	may
T4860	4446-4454	VB	denotes	resemble
T4861	4423-4432	RB	denotes	therefore
T4862	4433-4437	RBR	denotes	more
T4863	4438-4445	RB	denotes	closely
T4864	4455-4458	DT	denotes	the
T4865	4467-4479	NN	denotes	organization
T4866	4459-4466	JJ	denotes	genomic
T4867	4480-4489	VBG	denotes	resulting
T4868	4490-4494	IN	denotes	from
T4869	4495-4498	DT	denotes	the
T4870	4507-4518	NN	denotes	duplication
T4871	4499-4506	JJ	denotes	initial
T4872	4519-4521	IN	denotes	of
T4873	4522-4525	DT	denotes	the
T4874	4559-4566	NN	denotes	segment
T4875	4526-4535	JJ	denotes	ancestral
T4876	4536-4546	JJ	denotes	paralogous
T4877	4547-4558	JJ	denotes	chromosomal
T4878	4566-4567	.	denotes	.
T4879	4567-4697	sentence	denotes	Conserved paralogous clusters may result from the initial clustering of the genes in a relatively small ancestral genomic contig.
T4880	4568-4577	VBN	denotes	Conserved
T4881	4589-4597	NNS	denotes	clusters
T4882	4578-4588	JJ	denotes	paralogous
T4883	4602-4608	VB	denotes	result
T4884	4598-4601	MD	denotes	may
T4885	4609-4613	IN	denotes	from
T4886	4614-4617	DT	denotes	the
T4887	4626-4636	NN	denotes	clustering
T4888	4618-4625	JJ	denotes	initial
T4889	4637-4639	IN	denotes	of
T4890	4640-4643	DT	denotes	the
T4891	4644-4649	NNS	denotes	genes
T4892	4650-4652	IN	denotes	in
T4893	4653-4654	DT	denotes	a
T4894	4690-4696	NN	denotes	contig
T4895	4655-4665	RB	denotes	relatively
T4896	4666-4671	JJ	denotes	small
T4897	4672-4681	JJ	denotes	ancestral
T4898	4682-4689	JJ	denotes	genomic
T4899	4696-4697	.	denotes	.
T4900	4697-4903	sentence	denotes	Some evidence for the existence of "protoclusters" that could correspond to the paralogous chromosomal segments noted in higher vertebrates is present in the genome of the urochordate C. intestinalis [11].
T4901	4698-4702	DT	denotes	Some
T4902	4703-4711	NN	denotes	evidence
T4903	4838-4840	VBZ	denotes	is
T4904	4712-4715	IN	denotes	for
T4905	4716-4719	DT	denotes	the
T4906	4720-4729	NN	denotes	existence
T4907	4730-4732	IN	denotes	of
T4908	4733-4734	``	denotes	"
T4909	4734-4747	NNS	denotes	protoclusters
T4910	4747-4748	''	denotes	"
T4911	4749-4753	WDT	denotes	that
T4912	4760-4770	VB	denotes	correspond
T4913	4754-4759	MD	denotes	could
T4914	4771-4773	IN	denotes	to
T4915	4774-4777	DT	denotes	the
T4916	4801-4809	NNS	denotes	segments
T4917	4778-4788	JJ	denotes	paralogous
T4918	4789-4800	JJ	denotes	chromosomal
T4919	4810-4815	VBN	denotes	noted
T4920	4816-4818	IN	denotes	in
T4921	4819-4825	JJR	denotes	higher
T4922	4826-4837	NNS	denotes	vertebrates
T4923	4841-4848	JJ	denotes	present
T4924	4849-4851	IN	denotes	in
T4925	4852-4855	DT	denotes	the
T4926	4856-4862	NN	denotes	genome
T4927	4863-4865	IN	denotes	of
T4928	4866-4869	DT	denotes	the
T4929	4870-4881	NN	denotes	urochordate
T4930	4882-4884	NNP	denotes	C.
T4931	4885-4897	NNP	denotes	intestinalis
T4932	4898-4899	-LRB-	denotes	[
T4933	4899-4901	CD	denotes	11
T4934	4901-4902	-RRB-	denotes	]
T4935	4902-4903	.	denotes	.
T4936	4903-5171	sentence	denotes	For instance, the orthologues of FGFR, and WHSC1, carboxypeptidase Z and FLJ25359 cluster within an 85 kb region of the C. intestinalis genome and the human orthologues are still maintained in paralogous segments of 4p16, 8p and 10q (Fig. 3, [see Additional file 1]).
T4937	4904-4907	IN	denotes	For
T4938	4986-4993	VBP	denotes	cluster
T4939	4908-4916	NN	denotes	instance
T4940	4916-4918	,	denotes	,
T4941	4918-4921	DT	denotes	the
T4942	4922-4933	NNS	denotes	orthologues
T4943	4934-4936	IN	denotes	of
T4944	4937-4941	NN	denotes	FGFR
T4945	4941-4943	,	denotes	,
T4946	4943-4946	CC	denotes	and
T4947	4947-4952	NN	denotes	WHSC1
T4948	4952-4954	,	denotes	,
T4949	4954-4970	NN	denotes	carboxypeptidase
T4950	4971-4972	NN	denotes	Z
T4951	4973-4976	CC	denotes	and
T4952	4977-4985	NN	denotes	FLJ25359
T4953	4994-5000	IN	denotes	within
T4954	5001-5003	DT	denotes	an
T4955	5010-5016	NN	denotes	region
T4956	5004-5006	CD	denotes	85
T4957	5007-5009	NN	denotes	kb
T4958	5017-5019	IN	denotes	of
T4959	5020-5023	DT	denotes	the
T4960	5040-5046	NN	denotes	genome
T4961	5024-5026	NNP	denotes	C.
T4962	5027-5039	NNP	denotes	intestinalis
T4963	5047-5050	CC	denotes	and
T4964	5051-5054	DT	denotes	the
T4965	5061-5072	NNS	denotes	orthologues
T4966	5055-5060	JJ	denotes	human
T4967	5083-5093	VBN	denotes	maintained
T4968	5073-5076	VBP	denotes	are
T4969	5077-5082	RB	denotes	still
T4970	5094-5096	IN	denotes	in
T4971	5097-5107	JJ	denotes	paralogous
T4972	5108-5116	NNS	denotes	segments
T4973	5117-5119	IN	denotes	of
T4974	5120-5124	NN	denotes	4p16
T4975	5124-5126	,	denotes	,
T4976	5126-5128	NN	denotes	8p
T4977	5129-5132	CC	denotes	and
T4978	5133-5136	NN	denotes	10q
T4979	5137-5138	-LRB-	denotes	(
T4980	5138-5142	NN	denotes	Fig.
T4981	5143-5144	CD	denotes	3
T4982	5144-5146	,	denotes	,
T4983	5146-5147	-LRB-	denotes	[
T4984	5147-5150	VB	denotes	see
T4985	5151-5161	JJ	denotes	Additional
T4986	5162-5166	NN	denotes	file
T4987	5167-5168	CD	denotes	1
T4988	5168-5169	-RRB-	denotes	]
T4989	5169-5170	-RRB-	denotes	)
T4990	5170-5171	.	denotes	.
T4991	5171-5539	sentence	denotes	However, it should be noted that no clusters of genes from the vertebrate paralogous segments are locate close to the TACC or RHAMM genes of C. intestinalis, indicating that the formation of the much larger paralogous segments encompassing the FGFR-TACC genes formed later in evolutionary time, or conversely have been subject to extensive rearrangement in tunicates.
T4992	5172-5179	RB	denotes	However
T4993	5194-5199	VBN	denotes	noted
T4994	5179-5181	,	denotes	,
T4995	5181-5183	PRP	denotes	it
T4996	5184-5190	MD	denotes	should
T4997	5191-5193	VB	denotes	be
T4998	5200-5204	IN	denotes	that
T4999	5270-5276	VBN	denotes	locate
T5000	5205-5207	DT	denotes	no
T5001	5208-5216	NNS	denotes	clusters
T5002	5217-5219	IN	denotes	of
T5003	5220-5225	NNS	denotes	genes
T5004	5226-5230	IN	denotes	from
T5005	5231-5234	DT	denotes	the
T5006	5257-5265	NNS	denotes	segments
T5007	5235-5245	NN	denotes	vertebrate
T5008	5246-5256	JJ	denotes	paralogous
T5009	5266-5269	VBP	denotes	are
T5010	5277-5282	RB	denotes	close
T5011	5283-5285	IN	denotes	to
T5012	5286-5289	DT	denotes	the
T5013	5304-5309	NNS	denotes	genes
T5014	5290-5294	NN	denotes	TACC
T5015	5295-5297	CC	denotes	or
T5016	5298-5303	NN	denotes	RHAMM
T5017	5310-5312	IN	denotes	of
T5018	5313-5315	NNP	denotes	C.
T5019	5316-5328	NNP	denotes	intestinalis
T5020	5328-5330	,	denotes	,
T5021	5330-5340	VBG	denotes	indicating
T5022	5341-5345	IN	denotes	that
T5023	5432-5438	VBD	denotes	formed
T5024	5346-5349	DT	denotes	the
T5025	5350-5359	NN	denotes	formation
T5026	5360-5362	IN	denotes	of
T5027	5363-5366	DT	denotes	the
T5028	5390-5398	NNS	denotes	segments
T5029	5367-5371	RB	denotes	much
T5030	5372-5378	JJR	denotes	larger
T5031	5379-5389	JJ	denotes	paralogous
T5032	5399-5411	VBG	denotes	encompassing
T5033	5412-5415	DT	denotes	the
T5034	5426-5431	NNS	denotes	genes
T5035	5416-5420	NN	denotes	FGFR
T5036	5421-5425	NN	denotes	TACC
T5037	5420-5421	HYPH	denotes	-
T5038	5439-5444	RB	denotes	later
T5039	5445-5447	IN	denotes	in
T5040	5448-5460	JJ	denotes	evolutionary
T5041	5461-5465	NN	denotes	time
T5042	5465-5467	,	denotes	,
T5043	5467-5469	CC	denotes	or
T5044	5470-5480	RB	denotes	conversely
T5045	5486-5490	VBN	denotes	been
T5046	5481-5485	VBP	denotes	have
T5047	5491-5498	JJ	denotes	subject
T5048	5499-5501	IN	denotes	to
T5049	5502-5511	JJ	denotes	extensive
T5050	5512-5525	NN	denotes	rearrangement
T5051	5526-5528	IN	denotes	in
T5052	5529-5538	NNS	denotes	tunicates
T5053	5538-5539	.	denotes	.
T5054	5539-5896	sentence	denotes	In combination with the examination of the T. rubripes genome, this also provides additional evidence that either the second round of duplication of the chromosomal segment that contained the FGFR3/4 ancestor did not include a TACC gene, or that such a gene was lost very early in vertebrate evolution, prior to the divergence of the Gnanthostome lineages.
T5055	5540-5542	IN	denotes	In
T5056	5613-5621	VBZ	denotes	provides
T5057	5543-5554	NN	denotes	combination
T5058	5555-5559	IN	denotes	with
T5059	5560-5563	DT	denotes	the
T5060	5564-5575	NN	denotes	examination
T5061	5576-5578	IN	denotes	of
T5062	5579-5582	DT	denotes	the
T5063	5595-5601	NN	denotes	genome
T5064	5583-5585	NNP	denotes	T.
T5065	5586-5594	NNP	denotes	rubripes
T5066	5601-5603	,	denotes	,
T5067	5603-5607	DT	denotes	this
T5068	5608-5612	RB	denotes	also
T5069	5622-5632	JJ	denotes	additional
T5070	5633-5641	NN	denotes	evidence
T5071	5642-5646	IN	denotes	that
T5072	5757-5764	VB	denotes	include
T5073	5647-5653	CC	denotes	either
T5074	5654-5657	DT	denotes	the
T5075	5665-5670	NN	denotes	round
T5076	5658-5664	JJ	denotes	second
T5077	5671-5673	IN	denotes	of
T5078	5674-5685	NN	denotes	duplication
T5079	5686-5688	IN	denotes	of
T5080	5689-5692	DT	denotes	the
T5081	5705-5712	NN	denotes	segment
T5082	5693-5704	JJ	denotes	chromosomal
T5083	5713-5717	WDT	denotes	that
T5084	5718-5727	VBD	denotes	contained
T5085	5728-5731	DT	denotes	the
T5086	5740-5748	NN	denotes	ancestor
T5087	5732-5737	NN	denotes	FGFR3
T5088	5737-5738	HYPH	denotes	/
T5089	5738-5739	CD	denotes	4
T5090	5749-5752	VBD	denotes	did
T5091	5753-5756	RB	denotes	not
T5092	5765-5766	DT	denotes	a
T5093	5772-5776	NN	denotes	gene
T5094	5767-5771	NN	denotes	TACC
T5095	5776-5778	,	denotes	,
T5096	5778-5780	CC	denotes	or
T5097	5781-5785	IN	denotes	that
T5098	5802-5806	VBN	denotes	lost
T5099	5786-5790	PDT	denotes	such
T5100	5793-5797	NN	denotes	gene
T5101	5791-5792	DT	denotes	a
T5102	5798-5801	VBD	denotes	was
T5103	5807-5811	RB	denotes	very
T5104	5812-5817	RB	denotes	early
T5105	5818-5820	IN	denotes	in
T5106	5821-5831	NN	denotes	vertebrate
T5107	5832-5841	NN	denotes	evolution
T5108	5841-5843	,	denotes	,
T5109	5843-5848	RB	denotes	prior
T5110	5849-5851	IN	denotes	to
T5111	5852-5855	DT	denotes	the
T5112	5856-5866	NN	denotes	divergence
T5113	5867-5869	IN	denotes	of
T5114	5870-5873	DT	denotes	the
T5115	5887-5895	NNS	denotes	lineages
T5116	5874-5886	NNP	denotes	Gnanthostome
T5117	5895-5896	.	denotes	.
T5118	5896-6177	sentence	denotes	However, the final resolution of the initial evolution of these paralogous segment will await the sequencing of the amphioxus and lamprey genomes, which only have one FGFR gene, and therefore should only contain one copy of the other corresponding genes in this conserved segment.
T5119	5897-5904	RB	denotes	However
T5161	6131-6144	VBG	denotes	corresponding
T5162	6151-6153	IN	denotes	in
T5163	6154-6158	DT	denotes	this
T5164	6169-6176	NN	denotes	segment
T5165	6159-6168	VBN	denotes	conserved
T5166	6176-6177	.	denotes	.
T12611	1586-1592	JJ	denotes	Linear
T12612	1593-1605	NN	denotes	organization
T12613	1606-1608	IN	denotes	of
T12614	1609-1613	NN	denotes	gene
T12615	1614-1622	NNS	denotes	clusters
T12616	1623-1632	VBG	denotes	centering
T12617	1633-1637	IN	denotes	upon
T12618	1638-1641	DT	denotes	the
T12619	1654-1658	NNS	denotes	loci
T12620	1642-1653	JJ	denotes	chromosomal
T12621	1659-1661	IN	denotes	of
T12622	1662-1665	DT	denotes	the
T12623	1671-1676	NNS	denotes	genes
T12624	1666-1670	NN	denotes	FGFR
T12625	1677-1679	IN	denotes	in
T12626	1680-1686	NNS	denotes	humans
T12627	1686-1687	.	denotes	.
T12628	1687-1974	sentence	denotes	Paralogous genes present in at least two of the four loci are shown, with the exception of the region between GPX3 and NKX2E on chromosome 5, which appears to represent a series of intervening genes inserted after duplication of the 4p16/5q32-35 clusters, and genes mentioned in Fig. 3.
T12629	1688-1698	JJ	denotes	Paralogous
T12630	1699-1704	NNS	denotes	genes
T12631	1750-1755	VBN	denotes	shown
T12632	1705-1712	JJ	denotes	present
T12633	1713-1715	IN	denotes	in
T12634	1716-1718	RB	denotes	at
T12635	1725-1728	CD	denotes	two
T12636	1719-1724	RBS	denotes	least
T12637	1729-1731	IN	denotes	of
T12638	1732-1735	DT	denotes	the
T12639	1741-1745	NNS	denotes	loci
T12640	1736-1740	CD	denotes	four
T12641	1746-1749	VBP	denotes	are
T12642	1755-1757	,	denotes	,
T12643	1757-1761	IN	denotes	with
T12644	1762-1765	DT	denotes	the
T12645	1766-1775	NN	denotes	exception
T12646	1776-1778	IN	denotes	of
T12647	1779-1782	DT	denotes	the
T12648	1783-1789	NN	denotes	region
T12649	1790-1797	IN	denotes	between
T12650	1798-1802	NN	denotes	GPX3
T12651	1803-1806	CC	denotes	and
T12652	1807-1812	NN	denotes	NKX2E
T12653	1813-1815	IN	denotes	on
T12654	1816-1826	NN	denotes	chromosome
T12655	1827-1828	CD	denotes	5
T12656	1828-1830	,	denotes	,
T12657	1830-1835	WDT	denotes	which
T12658	1836-1843	VBZ	denotes	appears
T12659	1844-1846	TO	denotes	to
T12660	1847-1856	VB	denotes	represent
T12661	1857-1858	DT	denotes	a
T12662	1859-1865	NN	denotes	series
T12663	1866-1868	IN	denotes	of
T12664	1869-1880	VBG	denotes	intervening
T12665	1881-1886	NNS	denotes	genes
T12666	1887-1895	VBN	denotes	inserted
T12667	1896-1901	IN	denotes	after
T12668	1902-1913	NN	denotes	duplication
T12669	1914-1916	IN	denotes	of
T12670	1917-1920	DT	denotes	the
T12671	1934-1942	NNS	denotes	clusters
T12672	1921-1925	NN	denotes	4p16
T12673	1926-1930	NN	denotes	5q32
T12674	1925-1926	HYPH	denotes	/
T12675	1930-1931	HYPH	denotes	-
T12676	1931-1933	CD	denotes	35
T12677	1942-1944	,	denotes	,
T12678	1944-1947	CC	denotes	and
T12679	1948-1953	NNS	denotes	genes
T12680	1954-1963	VBN	denotes	mentioned
T12681	1964-1966	IN	denotes	in
T12682	1967-1971	NN	denotes	Fig.
T12683	1972-1973	CD	denotes	3
T12684	1973-1974	.	denotes	.
T12685	1974-2044	sentence	denotes	Corresponding syntenic mouse chromosomal regions (mm*) are indicated.
T12686	1975-1988	VBG	denotes	Corresponding
T12687	2016-2023	NNS	denotes	regions
T12688	1989-1997	JJ	denotes	syntenic
T12689	1998-2003	NN	denotes	mouse
T12690	2004-2015	JJ	denotes	chromosomal
T12691	2034-2043	VBN	denotes	indicated
T12692	2024-2025	-LRB-	denotes	(
T12693	2025-2027	NNP	denotes	mm
T12694	2027-2028	SYM	denotes	*
T12695	2028-2029	-RRB-	denotes	)
T12696	2030-2033	VBP	denotes	are
T12697	2043-2044	.	denotes	.
T12698	2044-2152	sentence	denotes	Takifugu rubripes scaffolds are shown (TR*) that contain more than one homologous gene from these clusters.
T12699	2045-2053	NNP	denotes	Takifugu
T12700	2054-2062	NNP	denotes	rubripes
T12701	2063-2072	NNS	denotes	scaffolds
T12702	2077-2082	VBN	denotes	shown
T12703	2073-2076	VBP	denotes	are
T12704	2083-2084	-LRB-	denotes	(
T12705	2084-2086	NN	denotes	TR
T12706	2086-2087	SYM	denotes	*
T12707	2087-2088	-RRB-	denotes	)
T12708	2089-2093	WDT	denotes	that
T12709	2094-2101	VBP	denotes	contain
T12710	2102-2106	JJR	denotes	more
T12711	2112-2115	CD	denotes	one
T12712	2107-2111	IN	denotes	than
T12713	2127-2131	NN	denotes	gene
T12714	2116-2126	JJ	denotes	homologous
T12715	2132-2136	IN	denotes	from
T12716	2137-2142	DT	denotes	these
T12717	2143-2151	NNS	denotes	clusters
T12718	2151-2152	.	denotes	.
T12719	2152-2245	sentence	denotes	Further details on the location of paralogous genes can be found in [see Additional file 1].
T12720	2153-2160	JJ	denotes	Further
T12721	2161-2168	NNS	denotes	details
T12722	2212-2217	VBN	denotes	found
T12723	2169-2171	IN	denotes	on
T12724	2172-2175	DT	denotes	the
T12725	2176-2184	NN	denotes	location
T12726	2185-2187	IN	denotes	of
T12727	2188-2198	JJ	denotes	paralogous
T12728	2199-2204	NNS	denotes	genes
T12729	2205-2208	MD	denotes	can
T12730	2209-2211	VB	denotes	be
T12731	2218-2220	IN	denotes	in
T12732	2221-2222	-LRB-	denotes	[
T12733	2222-2225	VB	denotes	see
T12734	2226-2236	JJ	denotes	Additional
T12735	2237-2241	NN	denotes	file
T12736	2242-2243	CD	denotes	1
T12737	2243-2244	-RRB-	denotes	]
T12738	2244-2245	.	denotes	.
T12814	6188-6197	NN	denotes	Formation
T12815	6198-6200	IN	denotes	of
T12816	6201-6214	NNS	denotes	protoclusters
T12817	6215-6217	IN	denotes	in
T12818	6218-6226	NNP	denotes	Takifugu
T12819	6227-6235	NNP	denotes	rubripes
T12820	6236-6239	CC	denotes	and
T12821	6240-6245	NNP	denotes	Ciona
T12822	6246-6258	NNP	denotes	intestinalis
T12823	6258-6260	:	denotes	:
T12824	6260-6261	-LRB-	denotes	(
T12825	6261-6262	LS	denotes	A
T12826	6260-6361	sentence	denotes	(A): Structure of the genomic scaffolds containing the Takifugu rubripes trTACC1A and trTACC1B genes.
T12827	6265-6274	NN	denotes	Structure
T12828	6262-6263	-RRB-	denotes	)
T12829	6263-6265	:	denotes	:
T12830	6275-6277	IN	denotes	of
T12831	6278-6281	DT	denotes	the
T12832	6290-6299	NNS	denotes	scaffolds
T12833	6282-6289	JJ	denotes	genomic
T12834	6300-6310	VBG	denotes	containing
T12835	6311-6314	DT	denotes	the
T12836	6355-6360	NNS	denotes	genes
T12837	6315-6323	NNP	denotes	Takifugu
T12838	6324-6332	NNS	denotes	rubripes
T12839	6333-6341	NN	denotes	trTACC1A
T12840	6342-6345	CC	denotes	and
T12841	6346-6354	NN	denotes	trTACC1B
T12842	6360-6361	.	denotes	.
T12843	6361-6520	sentence	denotes	Scaffold 12, the site for the trTACC1A gene contains genes found with either homologues or orthologues on the distal long arm of human chromosome 10 and 4p16.
T12844	6362-6370	NN	denotes	Scaffold
T12845	6406-6414	VBZ	denotes	contains
T12846	6371-6373	CD	denotes	12
T12847	6373-6375	,	denotes	,
T12848	6375-6378	DT	denotes	the
T12849	6379-6383	NN	denotes	site
T12850	6384-6387	IN	denotes	for
T12851	6388-6391	DT	denotes	the
T12852	6401-6405	NN	denotes	gene
T12853	6392-6400	NN	denotes	trTACC1A
T12854	6415-6420	NNS	denotes	genes
T12855	6421-6426	VBN	denotes	found
T12856	6427-6431	IN	denotes	with
T12857	6432-6438	CC	denotes	either
T12858	6439-6449	NNS	denotes	homologues
T12859	6450-6452	CC	denotes	or
T12860	6453-6464	NNS	denotes	orthologues
T12861	6465-6467	IN	denotes	on
T12862	6468-6471	DT	denotes	the
T12863	6484-6487	NN	denotes	arm
T12864	6472-6478	JJ	denotes	distal
T12865	6479-6483	JJ	denotes	long
T12866	6488-6490	IN	denotes	of
T12867	6491-6496	JJ	denotes	human
T12868	6515-6519	NN	denotes	4p16
T12869	6497-6507	NN	denotes	chromosome
T12870	6508-6510	CD	denotes	10
T12871	6511-6514	CC	denotes	and
T12872	6519-6520	.	denotes	.
T12873	6520-6654	sentence	denotes	This scaffold, therefore, has some of the characteristics of the predicted immediate ancestor of the TACC1/TACC2 chromosomal segment.
T12874	6521-6525	DT	denotes	This
T12875	6526-6534	NN	denotes	scaffold
T12876	6547-6550	VBZ	denotes	has
T12877	6534-6536	,	denotes	,
T12878	6536-6545	RB	denotes	therefore
T12879	6545-6547	,	denotes	,
T12880	6551-6555	DT	denotes	some
T12881	6556-6558	IN	denotes	of
T12882	6559-6562	DT	denotes	the
T12883	6563-6578	NNS	denotes	characteristics
T12884	6579-6581	IN	denotes	of
T12885	6582-6585	DT	denotes	the
T12886	6606-6614	NN	denotes	ancestor
T12887	6586-6595	VBN	denotes	predicted
T12888	6596-6605	JJ	denotes	immediate
T12889	6615-6617	IN	denotes	of
T12890	6618-6621	DT	denotes	the
T12891	6646-6653	NN	denotes	segment
T12892	6622-6627	NN	denotes	TACC1
T12893	6628-6633	NN	denotes	TACC2
T12894	6627-6628	HYPH	denotes	/
T12895	6634-6645	JJ	denotes	chromosomal
T12896	6653-6654	.	denotes	.
T12897	6654-6780	sentence	denotes	trTACC1B is found on scaffold 191, which contains orthologues of genes found in the proximal short arm of human chromosome 8.
T12898	6655-6663	NN	denotes	trTACC1B
T12899	6667-6672	VBN	denotes	found
T12900	6664-6666	VBZ	denotes	is
T12901	6673-6675	IN	denotes	on
T12902	6676-6684	NN	denotes	scaffold
T12903	6685-6688	CD	denotes	191
T12904	6688-6690	,	denotes	,
T12905	6690-6695	WDT	denotes	which
T12906	6696-6704	VBZ	denotes	contains
T12907	6705-6716	NNS	denotes	orthologues
T12908	6717-6719	IN	denotes	of
T12909	6720-6725	NNS	denotes	genes
T12910	6726-6731	VBN	denotes	found
T12911	6732-6734	IN	denotes	in
T12912	6735-6738	DT	denotes	the
T12913	6754-6757	NN	denotes	arm
T12914	6739-6747	JJ	denotes	proximal
T12915	6748-6753	JJ	denotes	short
T12916	6758-6760	IN	denotes	of
T12917	6761-6766	JJ	denotes	human
T12918	6767-6777	NN	denotes	chromosome
T12919	6778-6779	CD	denotes	8
T12920	6779-6780	.	denotes	.
T12921	6780-6889	sentence	denotes	(B): Ciona intestinalis clusters containing genes found in paralogous segments on human 8, 4p16, 10q and 5q.
T12922	6781-6782	-LRB-	denotes	(
T12923	6782-6783	LS	denotes	B
T12924	6805-6813	NNS	denotes	clusters
T12925	6783-6784	-RRB-	denotes	)
T12926	6784-6786	:	denotes	:
T12927	6786-6791	NNP	denotes	Ciona
T12928	6792-6804	NNP	denotes	intestinalis
T12929	6814-6824	VBG	denotes	containing
T12930	6825-6830	NNS	denotes	genes
T12931	6831-6836	VBN	denotes	found
T12932	6837-6839	IN	denotes	in
T12933	6840-6850	JJ	denotes	paralogous
T12934	6851-6859	NNS	denotes	segments
T12935	6860-6862	IN	denotes	on
T12936	6863-6868	JJ	denotes	human
T12937	6872-6876	NN	denotes	4p16
T12938	6869-6870	CD	denotes	8
T12939	6870-6872	,	denotes	,
T12940	6876-6878	,	denotes	,
T12941	6878-6881	NN	denotes	10q
T12942	6882-6885	CC	denotes	and
T12943	6886-6888	NN	denotes	5q
T12944	6888-6889	.	denotes	.
T12994	6900-6907	JJ	denotes	Genomic
T12995	6908-6917	NN	denotes	structure
T12996	6918-6920	IN	denotes	of
T12997	6921-6924	DT	denotes	the
T12998	6930-6935	NNS	denotes	genes
T12999	6925-6929	NN	denotes	TACC
T13000	6935-6936	.	denotes	.
T13001	6936-7022	sentence	denotes	Conserved regions known to bind protein factors in all human TACC proteins are shown.
T13002	6937-6946	VBN	denotes	Conserved
T13003	6947-6954	NNS	denotes	regions
T13004	7016-7021	VBN	denotes	shown
T13005	6955-6960	VBN	denotes	known
T13006	6961-6963	TO	denotes	to
T13007	6964-6968	VB	denotes	bind
T13008	6969-6976	NN	denotes	protein
T13009	6977-6984	NNS	denotes	factors
T13010	6985-6987	IN	denotes	in
T13011	6988-6991	DT	denotes	all
T13012	7003-7011	NN	denotes	proteins
T13013	6992-6997	JJ	denotes	human
T13014	6998-7002	NN	denotes	TACC
T13015	7012-7015	VBP	denotes	are
T13016	7021-7022	.	denotes	.
T13017	7022-7106	sentence	denotes	The SDP repeat of human TACC1-3 is known to bind GAS41 ([3,15] and data not shown).
T13018	7023-7026	DT	denotes	The
T13019	7031-7037	NN	denotes	repeat
T13020	7027-7030	NN	denotes	SDP
T13021	7058-7063	VBN	denotes	known
T13022	7038-7040	IN	denotes	of
T13023	7041-7046	JJ	denotes	human
T13024	7047-7052	NN	denotes	TACC1
T13025	7052-7053	HYPH	denotes	-
T13026	7053-7054	CD	denotes	3
T13027	7055-7057	VBZ	denotes	is
T13028	7064-7066	TO	denotes	to
T13029	7067-7071	VB	denotes	bind
T13030	7072-7077	NN	denotes	GAS41
T13031	7078-7079	-LRB-	denotes	(
T13032	7082-7084	CD	denotes	15
T13033	7079-7080	-LRB-	denotes	[
T13034	7080-7081	CD	denotes	3
T13035	7081-7082	,	denotes	,
T13036	7084-7085	-RRB-	denotes	]
T13037	7086-7089	CC	denotes	and
T13038	7090-7094	NNS	denotes	data
T13039	7099-7104	VBN	denotes	shown
T13040	7095-7098	RB	denotes	not
T13041	7104-7105	-RRB-	denotes	)
T13042	7105-7106	.	denotes	.
T13043	7106-7202	sentence	denotes	The TACC domain binds ch-TOG and members of the Aurora kinases in all species examined to date.
T13044	7107-7110	DT	denotes	The
T13045	7116-7122	NN	denotes	domain
T13046	7111-7115	NN	denotes	TACC
T13047	7123-7128	VBZ	denotes	binds
T13048	7129-7131	NN	denotes	ch
T13049	7132-7135	NN	denotes	TOG
T13050	7131-7132	HYPH	denotes	-
T13051	7136-7139	CC	denotes	and
T13052	7140-7147	NNS	denotes	members
T13053	7148-7150	IN	denotes	of
T13054	7151-7154	DT	denotes	the
T13055	7162-7169	NNS	denotes	kinases
T13056	7155-7161	NNP	denotes	Aurora
T13057	7170-7172	IN	denotes	in
T13058	7173-7176	DT	denotes	all
T13059	7177-7184	NNS	denotes	species
T13060	7185-7193	VBN	denotes	examined
T13061	7194-7196	IN	denotes	to
T13062	7197-7201	NN	denotes	date
T13063	7201-7202	.	denotes	.
T13064	7202-7278	sentence	denotes	This motif is characteristically encoded by the 3' exons of the TACC genes.
T13065	7203-7207	DT	denotes	This
T13066	7208-7213	NN	denotes	motif
T13067	7236-7243	VBN	denotes	encoded
T13068	7214-7216	VBZ	denotes	is
T13069	7217-7235	RB	denotes	characteristically
T13070	7244-7246	IN	denotes	by
T13071	7247-7250	DT	denotes	the
T13072	7254-7259	NNS	denotes	exons
T13073	7251-7252	CD	denotes	3
T13074	7252-7253	SYM	denotes	'
T13075	7260-7262	IN	denotes	of
T13076	7263-7266	DT	denotes	the
T13077	7272-7277	NNS	denotes	genes
T13078	7267-7271	NN	denotes	TACC
T13079	7277-7278	.	denotes	.
T13080	7278-7334	sentence	denotes	The size of the largest isoform for each gene is shown.
T13081	7279-7282	DT	denotes	The
T13082	7283-7287	NN	denotes	size
T13083	7328-7333	VBN	denotes	shown
T13084	7288-7290	IN	denotes	of
T13085	7291-7294	DT	denotes	the
T13086	7303-7310	NN	denotes	isoform
T13087	7295-7302	JJS	denotes	largest
T13088	7311-7314	IN	denotes	for
T13089	7315-7319	DT	denotes	each
T13090	7320-7324	NN	denotes	gene
T13091	7325-7327	VBZ	denotes	is
T13092	7333-7334	.	denotes	.
T4441	2271-2273	IN	denotes	of
T4442	2274-2278	NN	denotes	gene
T4443	2279-2284	NN	denotes	order
T4444	2285-2288	MD	denotes	can
T4445	2289-2293	RB	denotes	also
T4446	2353-2362	VBD	denotes	attempted
T4447	2302-2307	NNS	denotes	clues
T4448	2308-2310	IN	denotes	to
T4449	2311-2314	DT	denotes	the
T4450	2315-2324	NN	denotes	evolution
T4451	2325-2327	IN	denotes	of
T4452	2328-2332	NN	denotes	gene
T4453	2333-2343	NN	denotes	regulation
T4454	2343-2345	,	denotes	,
T4455	2345-2347	PRP	denotes	we
T4456	2348-2352	RB	denotes	next
T4457	2363-2365	TO	denotes	to
T4458	2366-2371	VB	denotes	trace
T4459	2372-2375	DT	denotes	the
T4460	2376-2385	NN	denotes	evolution
T4461	2386-2388	IN	denotes	of
T4462	2389-2394	DT	denotes	these
T4463	2406-2414	NNS	denotes	segments
T4464	2395-2405	JJ	denotes	paralogous
T4465	2415-2417	IN	denotes	by
T4466	2418-2427	VBG	denotes	examining
T4467	2428-2431	DT	denotes	the
T4468	2432-2438	NN	denotes	genome
T4469	2439-2441	IN	denotes	of
T4470	2442-2445	DT	denotes	the
T4471	2446-2454	NN	denotes	tunicate
T4401	1388-1396	NNS	denotes	clusters
T4402	1382-1387	CD	denotes	three
T4403	1397-1400	VBP	denotes	are
T4404	1409-1411	IN	denotes	on
T4405	1412-1421	JJ	denotes	different
T4406	1422-1433	NNS	denotes	chromosomes
T4407	1434-1436	IN	denotes	in
T4408	1437-1442	NN	denotes	mouse
T4409	1443-1446	CC	denotes	and
T4410	1447-1450	NN	denotes	rat
T4411	1451-1452	-LRB-	denotes	(
T4412	1452-1456	NN	denotes	Fig.
T4413	1457-1458	CD	denotes	2
T4414	1458-1459	-RRB-	denotes	)
T4415	1459-1461	,	denotes	,
T4416	1461-1468	RB	denotes	further
T4417	1469-1479	VBG	denotes	suggesting
T4418	1480-1484	IN	denotes	that
T4419	1511-1521	VBN	denotes	transposed
T4420	1485-1489	DT	denotes	this
T4421	1490-1497	NN	denotes	cluster
T4422	1498-1500	IN	denotes	of
T4423	1501-1506	NNS	denotes	genes
T4424	1507-1510	VBD	denotes	was
T4425	1522-1526	IN	denotes	into
T4426	1527-1531	DT	denotes	this
T4427	1532-1538	NN	denotes	region
T4428	1539-1544	IN	denotes	after
T4429	1545-1548	DT	denotes	the
T4430	1564-1574	NN	denotes	divergence
T4431	1549-1556	NN	denotes	primate
T4432	1557-1563	NN	denotes	rodent
T4433	1556-1557	HYPH	denotes	/
T4434	1574-1575	.	denotes	.
T4435	1575-1576	sentence	denotes
T4436	2246-2563	sentence	denotes	Because the conservation of gene order can also provide clues to the evolution of gene regulation, we next attempted to trace the evolution of these paralogous segments by examining the genome of the tunicate C. intestinalis [11] and the most "primitive" compact vertebrate genome sequenced to date, T. rubripes [14].
T4437	2246-2253	IN	denotes	Because
T4438	2294-2301	VB	denotes	provide
T4439	2254-2257	DT	denotes	the
T4440	2258-2270	NN	denotes	conservation
T4472	2455-2457	NNP	denotes	C.
T4473	2458-2470	NNP	denotes	intestinalis
T4474	2471-2472	-LRB-	denotes	[
T4475	2472-2474	CD	denotes	11
T4476	2474-2475	-RRB-	denotes	]
T4477	2476-2479	CC	denotes	and
T4478	2480-2483	DT	denotes	the
T4479	2520-2526	NN	denotes	genome
T4480	2484-2488	JJS	denotes	most
T4481	2490-2499	JJ	denotes	primitive
T5120	5985-5990	VB	denotes	await
T5121	5904-5906	,	denotes	,
T5122	5906-5909	DT	denotes	the
T5123	5916-5926	NN	denotes	resolution
T5124	5910-5915	JJ	denotes	final
T5125	5927-5929	IN	denotes	of
T5126	5930-5933	DT	denotes	the
T5127	5942-5951	NN	denotes	evolution
T5128	5934-5941	JJ	denotes	initial
T5129	5952-5954	IN	denotes	of
T5130	5955-5960	DT	denotes	these
T5131	5972-5979	NN	denotes	segment
T5132	5961-5971	JJ	denotes	paralogous
T5133	5980-5984	MD	denotes	will
T5134	5991-5994	DT	denotes	the
T5135	5995-6005	NN	denotes	sequencing
T5136	6006-6008	IN	denotes	of
T5137	6009-6012	DT	denotes	the
T5138	6035-6042	NNS	denotes	genomes
T5139	6013-6022	NN	denotes	amphioxus
T5140	6023-6026	CC	denotes	and
T5141	6027-6034	NN	denotes	lamprey
T5142	6042-6044	,	denotes	,
T5143	6044-6049	WDT	denotes	which
T5144	6055-6059	VBP	denotes	have
T5145	6050-6054	RB	denotes	only
T5146	6060-6063	CD	denotes	one
T5147	6069-6073	NN	denotes	gene
T5148	6064-6068	NN	denotes	FGFR
T5149	6073-6075	,	denotes	,
T5150	6075-6078	CC	denotes	and
T5151	6079-6088	RB	denotes	therefore
T5152	6101-6108	VB	denotes	contain
T5153	6089-6095	MD	denotes	should
T5154	6096-6100	RB	denotes	only
T5155	6109-6112	CD	denotes	one
T5156	6113-6117	NN	denotes	copy
T5157	6118-6120	IN	denotes	of
T5158	6121-6124	DT	denotes	the
T5159	6145-6150	NNS	denotes	genes
T5160	6125-6130	JJ	denotes	other
R8159	T12611	T12612	amod	Linear,organization
R8160	T12613	T12612	prep	of,organization
R8161	T12614	T12615	compound	gene,clusters
R8162	T12615	T12613	pobj	clusters,of
R8163	T12616	T12612	acl	centering,organization
R8164	T12617	T12616	prep	upon,centering
R8165	T12618	T12619	det	the,loci
R8166	T12619	T12617	pobj	loci,upon
R8167	T12620	T12619	amod	chromosomal,loci
R8168	T12621	T12619	prep	of,loci
R8169	T12622	T12623	det	the,genes
R8170	T12623	T12621	pobj	genes,of
R8171	T12624	T12623	compound	FGFR,genes
R8172	T12625	T12619	prep	in,loci
R8173	T12626	T12625	pobj	humans,in
R8174	T12627	T12612	punct	.,organization
R8175	T12629	T12630	amod	Paralogous,genes
R8176	T12630	T12631	nsubjpass	genes,shown
R8177	T12632	T12630	amod	present,genes
R8178	T12633	T12632	prep	in,present
R8179	T12634	T12635	advmod	at,two
R8180	T12635	T12633	pobj	two,in
R8181	T12636	T12635	advmod	least,two
R8182	T12637	T12635	prep	of,two
R8183	T12638	T12639	det	the,loci
R8184	T12639	T12637	pobj	loci,of
R8185	T12640	T12639	nummod	four,loci
R8186	T12641	T12631	auxpass	are,shown
R8187	T12642	T12631	punct	", ",shown
R8188	T12643	T12631	prep	with,shown
R8189	T12644	T12645	det	the,exception
R8190	T12645	T12643	pobj	exception,with
R8191	T12646	T12645	prep	of,exception
R8192	T12647	T12648	det	the,region
R8193	T12648	T12646	pobj	region,of
R8194	T12649	T12648	prep	between,region
R8195	T12650	T12649	pobj	GPX3,between
R8196	T12651	T12650	cc	and,GPX3
R8197	T12652	T12650	conj	NKX2E,GPX3
R8198	T12653	T12650	prep	on,GPX3
R8199	T12654	T12653	pobj	chromosome,on
R8200	T12655	T12654	nummod	5,chromosome
R8201	T12656	T12648	punct	", ",region
R8202	T12657	T12658	dep	which,appears
R8203	T12658	T12648	relcl	appears,region
R8204	T12659	T12660	aux	to,represent
R8205	T12660	T12658	xcomp	represent,appears
R8206	T12661	T12662	det	a,series
R8207	T12662	T12660	dobj	series,represent
R8208	T12663	T12662	prep	of,series
R8209	T12664	T12665	amod	intervening,genes
R8210	T12665	T12663	pobj	genes,of
R8211	T12666	T12662	acl	inserted,series
R8212	T12667	T12666	prep	after,inserted
R8213	T12668	T12667	pobj	duplication,after
R8214	T12669	T12668	prep	of,duplication
R8215	T12670	T12671	det	the,clusters
R8216	T12671	T12669	pobj	clusters,of
R8217	T12672	T12673	nmod	4p16,5q32
R8218	T12673	T12671	nmod	5q32,clusters
R8219	T12674	T12673	punct	/,5q32
R8220	T12675	T12673	punct	-,5q32
R8221	T12676	T12673	nummod	35,5q32
R8222	T12677	T12648	punct	", ",region
R8223	T12678	T12648	cc	and,region
R8224	T12679	T12648	conj	genes,region
R8225	T12680	T12679	acl	mentioned,genes
R8226	T12681	T12680	prep	in,mentioned
R8227	T12682	T12681	pobj	Fig.,in
R8228	T12683	T12682	nummod	3,Fig.
R8229	T12684	T12631	punct	.,shown
R8230	T12686	T12687	amod	Corresponding,regions
R8231	T12687	T12691	nsubjpass	regions,indicated
R8232	T12688	T12687	amod	syntenic,regions
R8233	T12689	T12687	nmod	mouse,regions
R8234	T12690	T12687	amod	chromosomal,regions
R8235	T12692	T12693	punct	(,mm
R8236	T12693	T12687	parataxis	mm,regions
R8237	T12694	T12693	punct	*,mm
R8238	T12695	T12693	punct	),mm
R8239	T12696	T12691	auxpass	are,indicated
R8240	T12697	T12691	punct	.,indicated
R8241	T12699	T12700	compound	Takifugu,rubripes
R8242	T12700	T12701	compound	rubripes,scaffolds
R8243	T12701	T12702	nsubjpass	scaffolds,shown
R8244	T12703	T12702	auxpass	are,shown
R8245	T12704	T12705	punct	(,TR
R8246	T12705	T12702	parataxis	TR,shown
R8247	T12706	T12705	punct	*,TR
R8248	T12707	T12705	punct	),TR
R8249	T12708	T12709	dep	that,contain
R8250	T12709	T12702	ccomp	contain,shown
R8251	T12710	T12711	amod	more,one
R8252	T12711	T12713	nummod	one,gene
R8253	T12712	T12711	quantmod	than,one
R8254	T12713	T12709	dobj	gene,contain
R8255	T12714	T12713	amod	homologous,gene
R8256	T12715	T12713	prep	from,gene
R8257	T12716	T12717	det	these,clusters
R8258	T12717	T12715	pobj	clusters,from
R8259	T12718	T12702	punct	.,shown
R8260	T12720	T12721	amod	Further,details
R8261	T12721	T12722	nsubjpass	details,found
R8262	T12723	T12721	prep	on,details
R8263	T12724	T12725	det	the,location
R8264	T12725	T12723	pobj	location,on
R8265	T12726	T12725	prep	of,location
R8266	T12727	T12728	amod	paralogous,genes
R8267	T12728	T12726	pobj	genes,of
R8268	T12729	T12722	aux	can,found
R8269	T12730	T12722	auxpass	be,found
R8270	T12731	T12722	prep	in,found
R8271	T12732	T12733	punct	[,see
R8272	T12733	T12731	pcomp	see,in
R8273	T12734	T12735	amod	Additional,file
R8274	T12735	T12733	dobj	file,see
R8275	T12736	T12735	nummod	1,file
R8276	T12737	T12733	punct	],see
R8277	T12738	T12722	punct	.,found
R8282	T12815	T12814	prep	of,Formation
R8283	T12816	T12815	pobj	protoclusters,of
R8284	T12817	T12814	prep	in,Formation
R8285	T12818	T12819	compound	Takifugu,rubripes
R8286	T12819	T12817	pobj	rubripes,in
R8287	T12820	T12819	cc	and,rubripes
R8288	T12821	T12822	compound	Ciona,intestinalis
R8289	T12822	T12819	conj	intestinalis,rubripes
R8290	T12823	T12814	punct	: ,Formation
R8291	T12824	T12825	punct	(,A
R8292	T12825	T12827	meta	A,Structure
R8293	T12828	T12825	punct	),A
R8294	T12829	T12827	punct	: ,Structure
R8295	T12830	T12827	prep	of,Structure
R8296	T12831	T12832	det	the,scaffolds
R8297	T12832	T12830	pobj	scaffolds,of
R8298	T12833	T12832	amod	genomic,scaffolds
R8299	T12834	T12832	acl	containing,scaffolds
R8300	T12835	T12836	det	the,genes
R8301	T12836	T12834	dobj	genes,containing
R8302	T12837	T12838	nmod	Takifugu,rubripes
R8303	T12838	T12836	nmod	rubripes,genes
R8304	T12839	T12836	nmod	trTACC1A,genes
R8305	T12840	T12839	cc	and,trTACC1A
R8306	T12841	T12839	conj	trTACC1B,trTACC1A
R8307	T12842	T12827	punct	.,Structure
R8308	T12844	T12845	nsubj	Scaffold,contains
R8309	T12846	T12844	nummod	12,Scaffold
R8310	T12847	T12844	punct	", ",Scaffold
R8311	T12848	T12849	det	the,site
R8312	T12849	T12844	appos	site,Scaffold
R8313	T12850	T12849	prep	for,site
R8314	T12851	T12852	det	the,gene
R8315	T12852	T12850	pobj	gene,for
R8316	T12853	T12852	compound	trTACC1A,gene
R8317	T12854	T12845	dobj	genes,contains
R8318	T12855	T12854	acl	found,genes
R8319	T12856	T12855	prep	with,found
R8320	T12857	T12858	preconj	either,homologues
R8321	T12858	T12856	pobj	homologues,with
R8322	T12859	T12858	cc	or,homologues
R8323	T12860	T12858	conj	orthologues,homologues
R8324	T12861	T12855	prep	on,found
R8325	T12862	T12863	det	the,arm
R8326	T12863	T12861	pobj	arm,on
R8327	T12864	T12863	amod	distal,arm
R8328	T12865	T12863	amod	long,arm
R8329	T12866	T12863	prep	of,arm
R8330	T12867	T12868	amod	human,4p16
R8331	T12868	T12866	pobj	4p16,of
R8332	T12869	T12868	nmod	chromosome,4p16
R8333	T12870	T12868	nummod	10,4p16
R8334	T12871	T12868	cc	and,4p16
R8335	T12872	T12845	punct	.,contains
R8336	T12874	T12875	det	This,scaffold
R8337	T12875	T12876	nsubj	scaffold,has
R8338	T12877	T12876	punct	", ",has
R8339	T12878	T12876	advmod	therefore,has
R8340	T12879	T12876	punct	", ",has
R8341	T12880	T12876	dobj	some,has
R8342	T12881	T12880	prep	of,some
R8343	T12882	T12883	det	the,characteristics
R8344	T12883	T12881	pobj	characteristics,of
R8345	T12884	T12883	prep	of,characteristics
R8346	T12885	T12886	det	the,ancestor
R8347	T12886	T12884	pobj	ancestor,of
R8348	T12887	T12886	amod	predicted,ancestor
R8349	T12888	T12886	amod	immediate,ancestor
R8350	T12889	T12886	prep	of,ancestor
R8351	T12890	T12891	det	the,segment
R8352	T12891	T12889	pobj	segment,of
R8353	T12892	T12893	nmod	TACC1,TACC2
R8354	T12893	T12891	nmod	TACC2,segment
R8355	T12894	T12893	punct	/,TACC2
R8356	T12895	T12891	amod	chromosomal,segment
R8357	T12896	T12876	punct	.,has
R8358	T12898	T12899	nsubjpass	trTACC1B,found
R8359	T12900	T12899	auxpass	is,found
R8360	T12901	T12899	prep	on,found
R8361	T12902	T12901	pobj	scaffold,on
R8362	T12903	T12902	nummod	191,scaffold
R8363	T12904	T12902	punct	", ",scaffold
R8364	T12905	T12906	dep	which,contains
R8365	T12906	T12902	relcl	contains,scaffold
R8366	T12907	T12906	dobj	orthologues,contains
R8367	T12908	T12907	prep	of,orthologues
R8368	T12909	T12908	pobj	genes,of
R8369	T12910	T12909	acl	found,genes
R8370	T12911	T12910	prep	in,found
R8371	T12912	T12913	det	the,arm
R8372	T12913	T12911	pobj	arm,in
R8373	T12914	T12913	amod	proximal,arm
R8374	T12915	T12913	amod	short,arm
R8375	T12916	T12913	prep	of,arm
R8376	T12917	T12918	amod	human,chromosome
R8377	T12918	T12916	pobj	chromosome,of
R8378	T12919	T12918	nummod	8,chromosome
R8379	T12920	T12899	punct	.,found
R8380	T12922	T12923	punct	(,B
R8381	T12923	T12924	meta	B,clusters
R8382	T12925	T12923	punct	),B
R8383	T12926	T12924	punct	: ,clusters
R8384	T12927	T12928	compound	Ciona,intestinalis
R8385	T12928	T12924	compound	intestinalis,clusters
R8386	T12929	T12924	acl	containing,clusters
R8387	T12930	T12929	dobj	genes,containing
R8388	T12931	T12930	acl	found,genes
R8389	T12932	T12931	prep	in,found
R8390	T12933	T12934	amod	paralogous,segments
R8391	T12934	T12932	pobj	segments,in
R8392	T12935	T12931	prep	on,found
R8393	T12936	T12937	amod	human,4p16
R8394	T12937	T12935	pobj	4p16,on
R8395	T12938	T12937	nummod	8,4p16
R8396	T12939	T12937	punct	", ",4p16
R8397	T12940	T12937	punct	", ",4p16
R8398	T12941	T12937	conj	10q,4p16
R8399	T12942	T12941	cc	and,10q
R8400	T12943	T12941	conj	5q,10q
R8401	T12944	T12924	punct	.,clusters
R8402	T12994	T12995	amod	Genomic,structure
R8403	T12996	T12995	prep	of,structure
R8404	T12997	T12998	det	the,genes
R8405	T12998	T12996	pobj	genes,of
R8406	T12999	T12998	compound	TACC,genes
R8407	T13000	T12995	punct	.,structure
R8408	T13002	T13003	amod	Conserved,regions
R8409	T13003	T13004	nsubjpass	regions,shown
R8410	T13005	T13003	acl	known,regions
R8411	T13006	T13007	aux	to,bind
R8412	T13007	T13005	xcomp	bind,known
R8413	T13008	T13009	compound	protein,factors
R8414	T13009	T13007	dobj	factors,bind
R8415	T13010	T13007	prep	in,bind
R8416	T13011	T13012	det	all,proteins
R8417	T13012	T13010	pobj	proteins,in
R8418	T13013	T13012	amod	human,proteins
R8419	T13014	T13012	compound	TACC,proteins
R8420	T13015	T13004	auxpass	are,shown
R8421	T13016	T13004	punct	.,shown
R8422	T13018	T13019	det	The,repeat
R8423	T13019	T13021	nsubjpass	repeat,known
R8424	T13020	T13019	compound	SDP,repeat
R8425	T13022	T13019	prep	of,repeat
R8426	T13023	T13024	amod	human,TACC1
R8427	T13024	T13022	pobj	TACC1,of
R8428	T13025	T13024	punct	-,TACC1
R8429	T13026	T13024	nummod	3,TACC1
R8430	T13027	T13021	auxpass	is,known
R8431	T13028	T13029	aux	to,bind
R8432	T13029	T13021	xcomp	bind,known
R8433	T13030	T13029	dobj	GAS41,bind
R8434	T13031	T13032	punct	(,15
R8435	T13032	T13021	parataxis	15,known
R8436	T13033	T13032	punct	[,15
R8437	T13034	T13032	nummod	3,15
R8438	T13035	T13032	punct	",",15
R8439	T13036	T13032	punct	],15
R8440	T13037	T13032	cc	and,15
R8441	T13038	T13039	nsubj	data,shown
R8442	T13039	T13032	conj	shown,15
R8443	T13040	T13039	neg	not,shown
R8444	T13041	T13032	punct	),15
R8445	T13042	T13021	punct	.,known
R8446	T13044	T13045	det	The,domain
R8447	T13045	T13047	nsubj	domain,binds
R8448	T13046	T13045	compound	TACC,domain
R8449	T13048	T13049	compound	ch,TOG
R8450	T13049	T13047	dobj	TOG,binds
R8451	T13050	T13049	punct	-,TOG
R8452	T13051	T13049	cc	and,TOG
R8453	T13052	T13049	conj	members,TOG
R8454	T13053	T13052	prep	of,members
R8455	T13054	T13055	det	the,kinases
R8456	T13055	T13053	pobj	kinases,of
R8457	T13056	T13055	compound	Aurora,kinases
R8458	T13057	T13047	prep	in,binds
R8459	T13058	T13059	det	all,species
R8460	T13059	T13057	pobj	species,in
R8461	T13060	T13059	acl	examined,species
R8462	T13061	T13060	prep	to,examined
R8463	T13062	T13061	pobj	date,to
R8464	T13063	T13047	punct	.,binds
R8465	T13065	T13066	det	This,motif
R8466	T13066	T13067	nsubjpass	motif,encoded
R8467	T13068	T13067	auxpass	is,encoded
R8468	T13069	T13067	advmod	characteristically,encoded
R8469	T13070	T13067	agent	by,encoded
R8470	T13071	T13072	det	the,exons
R8471	T13072	T13070	pobj	exons,by
R8472	T13073	T13072	nummod	3,exons
R8473	T13074	T13073	punct	',3
R8474	T13075	T13072	prep	of,exons
R8475	T13076	T13077	det	the,genes
R8476	T13077	T13075	pobj	genes,of
R8477	T13078	T13077	compound	TACC,genes
R8478	T13079	T13067	punct	.,encoded
R8479	T13081	T13082	det	The,size
R8480	T13082	T13083	nsubjpass	size,shown
R8481	T13084	T13082	prep	of,size
R8482	T13085	T13086	det	the,isoform
R8483	T13086	T13084	pobj	isoform,of
R8484	T13087	T13086	amod	largest,isoform
R8485	T13088	T13086	prep	for,isoform
R8486	T13089	T13090	det	each,gene
R8487	T13090	T13088	pobj	gene,for
R8488	T13091	T13083	auxpass	is,shown
R8489	T13092	T13083	punct	.,shown
R2443	T4146	T4145	prep	of,Evolution
R2444	T4147	T4148	det	the,segments
R2445	T4148	T4146	pobj	segments,of
R2446	T4149	T4148	amod	chromosomal,segments
R2447	T4150	T4148	acl	containing,segments
R2448	T4151	T4152	det	the,genes
R2449	T4152	T4150	dobj	genes,containing
R2450	T4153	T4152	compound	TACC,genes
R2451	T4155	T4156	det	The,tree
R2452	T4156	T4158	nsubj	tree,resembles
R2453	T4157	T4156	amod	phylogenetic,tree
R2454	T4159	T4156	prep	of,tree
R2455	T4160	T4161	det	the,genes
R2456	T4161	T4159	pobj	genes,of
R2457	T4162	T4161	compound	FGFR,genes
R2458	T4163	T4158	advmod	closely,resembles
R2459	T4164	T4158	dobj	that,resembles
R2460	T4165	T4164	prep	of,that
R2461	T4166	T4167	det	the,genes
R2462	T4167	T4165	pobj	genes,of
R2463	T4168	T4167	nmod	vertebrate,genes
R2464	T4169	T4167	nmod	TACC1,genes
R2465	T4170	T4169	punct	-,TACC1
R2466	T4171	T4169	nummod	3,TACC1
R2467	T4172	T4158	punct	.,resembles
R2468	T4174	T4175	advmod	Recently,revealed
R2469	T4176	T4175	punct	", ",revealed
R2470	T4177	T4178	amod	detailed,analyses
R2471	T4178	T4175	nsubj	analyses,revealed
R2472	T4179	T4178	prep	of,analyses
R2473	T4180	T4181	det	the,regions
R2474	T4181	T4179	pobj	regions,of
R2475	T4182	T4181	amod	chromosomal,regions
R2476	T4183	T4181	acl	containing,regions
R2477	T4184	T4185	det	the,family
R2478	T4185	T4183	dobj	family,containing
R2479	T4186	T4185	compound	FGFR,family
R2480	T4187	T4185	compound	gene,family
R2481	T4188	T4183	prep	in,containing
R2482	T4189	T4188	pobj	humans,in
R2483	T4190	T4189	punct	", ",humans
R2484	T4191	T4189	conj	mouse,humans
R2485	T4192	T4191	cc	and,mouse
R2486	T4193	T4194	det	the,arthopod
R2487	T4194	T4191	conj	arthopod,mouse
R2488	T4195	T4196	compound	D.,melanogaster
R2489	T4196	T4194	appos	melanogaster,arthopod
R2490	T4197	T4175	aux	have,revealed
R2491	T4198	T4199	det	the,conservation
R2492	T4199	T4175	dobj	conservation,revealed
R2493	T4200	T4199	prep	of,conservation
R2494	T4201	T4202	amod	paralogous,segments
R2495	T4202	T4200	pobj	segments,of
R2496	T4203	T4202	amod	chromosomal,segments
R2497	T4204	T4199	prep	between,conservation
R2498	T4205	T4206	det	these,organisms
R2499	T4206	T4204	pobj	organisms,between
R2500	T4207	T4208	punct	(,Table
R2501	T4208	T4175	parataxis	Table,revealed
R2502	T4209	T4208	dep	Fig.,Table
R2503	T4210	T4209	nummod	2,Fig.
R2504	T4211	T4208	punct	", ",Table
R2505	T4212	T4208	punct	[,Table
R2506	T4213	T4208	dep	13,Table
R2507	T4214	T4208	punct	],Table
R2508	T4215	T4208	punct	", ",Table
R2509	T4216	T4208	nummod	1,Table
R2510	T4217	T4208	punct	[,Table
R2511	T4218	T4208	advcl	see,Table
R2512	T4219	T4220	amod	Additional,file
R2513	T4220	T4218	dobj	file,see
R2514	T4221	T4220	nummod	1,file
R2515	T4222	T4208	punct	],Table
R2516	T4223	T4208	punct	),Table
R2517	T4224	T4175	punct	.,revealed
R2518	T4226	T4227	nsubj	This,provided
R2519	T4228	T4227	aux	has,provided
R2520	T4229	T4230	amod	further,support
R2521	T4230	T4227	dobj	support,provided
R2522	T4231	T4232	mark	that,duplicated
R2523	T4232	T4230	acl	duplicated,support
R2524	T4233	T4234	det	an,segment
R2525	T4234	T4232	nsubjpass	segment,duplicated
R2526	T4235	T4234	amod	ancient,segment
R2527	T4236	T4234	amod	chromosomal,segment
R2528	T4237	T4232	auxpass	was,duplicated
R2529	T4238	T4232	advmod	twice,duplicated
R2530	T4239	T4232	prep	during,duplicated
R2531	T4240	T4241	compound	vertebrate,evolution
R2532	T4241	T4239	pobj	evolution,during
R2533	T4242	T4232	punct	", ",duplicated
R2534	T4243	T4232	prep	with,duplicated
R2535	T4244	T4245	det	the,duplication
R2536	T4245	T4247	nsubj	duplication,occurring
R2537	T4246	T4245	amod	first,duplication
R2538	T4247	T4243	pcomp	occurring,with
R2539	T4248	T4249	dep	that,gave
R2540	T4249	T4245	relcl	gave,duplication
R2541	T4250	T4249	dobj	rise,gave
R2542	T4251	T4249	prep	to,gave
R2543	T4252	T4253	det	the,ancestors
R2544	T4253	T4251	pobj	ancestors,to
R2545	T4254	T4255	amod	human,chromosome
R2546	T4255	T4256	nmod	chromosome,ter
R2547	T4256	T4253	nmod	ter,ancestors
R2548	T4257	T4258	nmod	4p16,5q32
R2549	T4258	T4256	nmod	5q32,ter
R2550	T4259	T4258	punct	/,5q32
R2551	T4260	T4256	punct	-,ter
R2552	T4261	T4256	cc	and,ter
R2553	T4262	T4263	amod	human,chromosome
R2554	T4263	T4264	compound	chromosome,ter
R2555	T4264	T4256	conj	ter,ter
R2556	T4265	T4266	compound	8p,10q23
R2557	T4266	T4264	compound	10q23,ter
R2558	T4267	T4266	punct	/,10q23
R2559	T4268	T4264	punct	-,ter
R2560	T4269	T4247	prep	in,occurring
R2561	T4270	T4271	det	the,stages
R2562	T4271	T4269	pobj	stages,in
R2563	T4272	T4271	amod	early,stages
R2564	T4273	T4271	prep	after,stages
R2565	T4274	T4275	det	the,divergence
R2566	T4275	T4273	pobj	divergence,after
R2567	T4276	T4275	compound	invertebrate,divergence
R2568	T4277	T4227	punct	.,provided
R2569	T4279	T4280	nsubj	This,suggests
R2570	T4281	T4282	mark	that,arose
R2571	T4282	T4280	ccomp	arose,suggests
R2572	T4283	T4284	det	the,pair
R2573	T4284	T4282	nsubj	pair,arose
R2574	T4285	T4284	amod	ancestral,pair
R2575	T4286	T4287	compound	FGFR,TACC
R2576	T4287	T4284	compound	TACC,pair
R2577	T4288	T4287	punct	-,TACC
R2578	T4289	T4284	compound	gene,pair
R2579	T4290	T4291	advmod	most,probably
R2580	T4291	T4282	advmod	probably,arose
R2581	T4292	T4293	amod	prior,to
R2582	T4293	T4282	prep	to,arose
R2583	T4294	T4295	det	the,duplication
R2584	T4295	T4293	pobj	duplication,to
R2585	T4296	T4295	amod	initial,duplication
R2586	T4297	T4295	cc	and,duplication
R2587	T4298	T4299	amod	subsequent,divergence
R2588	T4299	T4295	conj	divergence,duplication
R2589	T4300	T4295	prep	of,duplication
R2590	T4301	T4302	det	these,segments
R2591	T4302	T4300	pobj	segments,of
R2592	T4303	T4302	amod	paralogous,segments
R2593	T4304	T4302	amod	chromosomal,segments
R2594	T4305	T4295	punct	", ",duplication
R2595	T4306	T4295	acl	estimated,duplication
R2596	T4307	T4308	aux	to,occurred
R2597	T4308	T4306	xcomp	occurred,estimated
R2598	T4309	T4308	aux	have,occurred
R2599	T4310	T4311	quantmod	687,155.7
R2600	T4311	T4313	npadvmod	155.7,MYA
R2601	T4312	T4311	punct	±,155.7
R2602	T4313	T4308	advmod	MYA,occurred
R2603	T4314	T4280	punct	.,suggests
R2604	T4316	T4317	nsubj	This,raised
R2605	T4318	T4317	aux	has,raised
R2606	T4319	T4320	det	the,suggestion
R2607	T4320	T4317	dobj	suggestion,raised
R2608	T4321	T4322	mark	that,reside
R2609	T4322	T4320	acl	reside,suggestion
R2610	T4323	T4324	det	a,gene
R2611	T4324	T4322	nsubj	gene,reside
R2612	T4325	T4324	amod	fourth,gene
R2613	T4326	T4324	compound	TACC,gene
R2614	T4327	T4324	prep	in,gene
R2615	T4328	T4327	pobj	vertebrates,in
R2616	T4329	T4322	aux	would,reside
R2617	T4330	T4322	prep	in,reside
R2618	T4331	T4332	det	the,region
R2619	T4332	T4330	pobj	region,in
R2620	T4333	T4332	amod	same,region
R2621	T4334	T4332	amod	chromosomal,region
R2622	T4335	T4332	prep	as,region
R2623	T4336	T4335	pobj	FGFR4,as
R2624	T4337	T4317	punct	.,raised
R2625	T4339	T4340	advmod	Indeed,used
R2626	T4341	T4342	det	this,hypothesis
R2627	T4342	T4340	nsubjpass	hypothesis,used
R2628	T4343	T4340	aux	has,used
R2629	T4344	T4340	auxpass	been,used
R2630	T4345	T4340	prep	in,used
R2631	T4346	T4345	pobj	support,in
R2632	T4347	T4346	prep	for,support
R2633	T4348	T4349	det	the,gene
R2634	T4349	T4347	pobj	gene,for
R2635	T4350	T4349	compound	RHAMM,gene
R2636	T4351	T4349	prep	as,gene
R2637	T4352	T4353	det	a,member
R2638	T4353	T4351	pobj	member,as
R2639	T4354	T4353	prep	of,member
R2640	T4355	T4356	det	the,family
R2641	T4356	T4354	pobj	family,of
R2642	T4357	T4356	compound	TACC,family
R2643	T4358	T4359	punct	[,10
R2644	T4359	T4340	parataxis	10,used
R2645	T4360	T4359	punct	],10
R2646	T4361	T4340	punct	.,used
R2647	T4363	T4364	amod	Human,RHAMM
R2648	T4364	T4365	nsubj	RHAMM,maps
R2649	T4366	T4365	prep	to,maps
R2650	T4367	T4368	compound	chromosome,5q32
R2651	T4368	T4366	pobj	5q32,to
R2652	T4369	T4365	prep	in,maps
R2653	T4370	T4371	det	a,region
R2654	T4371	T4369	pobj	region,in
R2655	T4372	T4371	acl	bounded,region
R2656	T4373	T4372	agent	by,bounded
R2657	T4374	T4373	pobj	GPX3,by
R2658	T4375	T4374	cc	and,GPX3
R2659	T4376	T4374	conj	NKX2E,GPX3
R2660	T4377	T4365	punct	.,maps
R2661	T4379	T4380	det	These,loci
R2662	T4380	T4381	nsubj	loci,separate
R2663	T4382	T4383	nummod	two,clusters
R2664	T4383	T4381	dobj	clusters,separate
R2665	T4384	T4383	prep	of,clusters
R2666	T4385	T4384	pobj	genes,of
R2667	T4386	T4383	prep	on,clusters
R2668	T4387	T4388	amod	human,chromosome
R2669	T4388	T4386	pobj	chromosome,on
R2670	T4389	T4388	nummod	5,chromosome
R2671	T4390	T4391	dep	that,are
R2672	T4391	T4383	relcl	are,clusters
R2713	T4434	T4398	punct	.,located
R2715	T4438	T4446	advcl	provide,attempted
R2716	T4439	T4440	det	the,conservation
R2717	T4440	T4438	nsubj	conservation,provide
R2718	T4441	T4440	prep	of,conservation
R2719	T4442	T4443	compound	gene,order
R2720	T4443	T4441	pobj	order,of
R2721	T4444	T4438	aux	can,provide
R2722	T4445	T4438	advmod	also,provide
R2723	T4447	T4438	dobj	clues,provide
R2724	T4448	T4447	prep	to,clues
R2725	T4449	T4450	det	the,evolution
R2726	T4450	T4448	pobj	evolution,to
R2727	T4451	T4450	prep	of,evolution
R2728	T4452	T4453	compound	gene,regulation
R2729	T4453	T4451	pobj	regulation,of
R2730	T4454	T4446	punct	", ",attempted
R2731	T4455	T4446	nsubj	we,attempted
R2732	T4456	T4446	advmod	next,attempted
R2733	T4457	T4458	aux	to,trace
R2734	T4458	T4446	xcomp	trace,attempted
R2735	T4459	T4460	det	the,evolution
R2736	T4460	T4458	dobj	evolution,trace
R2737	T4461	T4460	prep	of,evolution
R2738	T4462	T4463	det	these,segments
R2739	T4463	T4461	pobj	segments,of
R2740	T4464	T4463	amod	paralogous,segments
R2741	T4465	T4458	prep	by,trace
R2742	T4466	T4465	pcomp	examining,by
R2743	T4467	T4468	det	the,genome
R2744	T4468	T4466	dobj	genome,examining
R2745	T4469	T4468	prep	of,genome
R2746	T4470	T4471	det	the,tunicate
R2747	T4471	T4469	pobj	tunicate,of
R2748	T4472	T4473	compound	C.,intestinalis
R2749	T4473	T4471	appos	intestinalis,tunicate
R2750	T4474	T4475	punct	[,11
R2751	T4475	T4471	parataxis	11,tunicate
R2752	T4476	T4475	punct	],11
R2753	T4477	T4468	cc	and,genome
R2754	T4478	T4479	det	the,genome
R2755	T4479	T4468	conj	genome,genome
R2756	T4480	T4481	amod	most,primitive
R2757	T4481	T4479	amod	primitive,genome
R2758	T4482	T4481	punct	"""",primitive
R2759	T4483	T4481	punct	"""",primitive
R2760	T4484	T4479	amod	compact,genome
R2761	T4485	T4479	compound	vertebrate,genome
R2762	T4486	T4479	acl	sequenced,genome
R2763	T4487	T4486	prep	to,sequenced
R2764	T4488	T4487	pobj	date,to
R2765	T4489	T4479	punct	", ",genome
R2766	T4490	T4491	compound	T.,rubripes
R2767	T4491	T4479	appos	rubripes,genome
R2768	T4492	T4493	punct	[,14
R2769	T4493	T4479	parataxis	14,genome
R2770	T4494	T4493	punct	],14
R2771	T4495	T4446	punct	.,attempted
R2772	T4497	T4498	mark	Although,assembled
R2773	T4498	T4501	advcl	assembled,confirmed
R2774	T4499	T4498	neg	not,assembled
R2775	T4500	T4498	advmod	fully,assembled
R2776	T4502	T4501	punct	", ",confirmed
R2777	T4503	T4501	nsubj	examination,confirmed
R2778	T4504	T4503	prep	of,examination
R2779	T4505	T4506	det	the,genome
R2780	T4506	T4504	pobj	genome,of
R2781	T4507	T4506	prep	of,genome
R2782	T4508	T4509	compound	T.,rubripes
R2783	T4509	T4507	pobj	rubripes,of
R2784	T4510	T4511	det	the,presence
R2785	T4511	T4501	dobj	presence,confirmed
R2786	T4512	T4511	prep	of,presence
R2787	T4513	T4514	amod	chromosomal,segments
R2788	T4514	T4512	pobj	segments,of
R2789	T4515	T4514	amod	paralogous,segments
R2790	T4516	T4515	prep	to,paralogous
R2791	T4517	T4516	pobj	those,to
R2792	T4518	T4517	acl	found,those
R2793	T4519	T4518	prep	in,found
R2794	T4520	T4521	amod	higher,vertebrates
R2795	T4521	T4519	pobj	vertebrates,in
R2796	T4522	T4523	punct	(,Fig.
R2797	T4523	T4501	parataxis	Fig.,confirmed
R2798	T4524	T4523	nummod	2,Fig.
R2799	T4525	T4523	punct	),Fig.
R2800	T4526	T4501	punct	.,confirmed
R2801	T4528	T4529	prep	For,found
R2802	T4530	T4528	pobj	instance,For
R2803	T4531	T4529	punct	", ",found
R2804	T4532	T4533	det	the,orthologues
R2805	T4533	T4529	nsubjpass	orthologues,found
R2806	T4534	T4533	prep	of,orthologues
R2807	T4535	T4534	pobj	GPRK2L,of
R2808	T4536	T4535	cc	and,GPRK2L
R2809	T4537	T4535	conj	RGS12,GPRK2L
R2810	T4538	T4529	auxpass	are,found
R2811	T4539	T4529	prep	on,found
R2812	T4540	T4541	compound	T.,rubripes
R2813	T4541	T4542	compound	rubripes,scaffold
R2814	T4542	T4539	pobj	scaffold,on
R2815	T4543	T4542	nummod	290,scaffold
R2816	T4544	T4542	punct	(,scaffold
R2817	T4545	T4542	appos	emb\|CAAB01000290.1,scaffold
R2818	T4546	T4542	punct	),scaffold
R2819	T4547	T4539	punct	", ",on
R2820	T4548	T4539	cc	and,on
R2821	T4549	T4539	conj	within,on
R2822	T4550	T4551	nummod	300,kb
R2823	T4551	T4549	pobj	kb,within
R2824	T4552	T4551	prep	of,kb
R2825	T4553	T4554	det	each,other
R2826	T4554	T4552	pobj	other,of
R2827	T4555	T4551	prep	in,kb
R2828	T4556	T4557	amod	human,4p16
R2829	T4557	T4555	pobj	4p16,in
R2830	T4558	T4529	punct	.,found
R2831	T4560	T4561	det	The,orthologues
R2832	T4561	T4564	nsubjpass	orthologues,located
R2833	T4562	T4563	compound	T.,rubripes
R2834	T4563	T4561	compound	rubripes,orthologues
R2835	T4565	T4561	prep	of,orthologues
R2836	T4566	T4565	pobj	FGFR3,of
R2837	T4567	T4561	punct	", ",orthologues
R2838	T4568	T4561	appos	LETM1,orthologues
R2839	T4569	T4568	cc	and,LETM1
R2840	T4570	T4568	conj	WHSC1,LETM1
R2841	T4571	T4564	auxpass	are,located
R2842	T4572	T4564	prep	on,located
R2843	T4573	T4574	det	the,scaffold
R2844	T4574	T4572	pobj	scaffold,on
R2845	T4575	T4574	amod	same,scaffold
R2846	T4576	T4577	nummod	166,kb
R2847	T4577	T4574	nmod	kb,scaffold
R2848	T4578	T4574	amod	genomic,scaffold
R2849	T4579	T4574	nummod	251,scaffold
R2850	T4580	T4574	punct	(,scaffold
R2851	T4581	T4574	appos	emb\|CAAB01000166.1,scaffold
R2852	T4582	T4574	punct	),scaffold
R2853	T4583	T4564	punct	.,located
R2854	T4585	T4586	advmod	Significantly,located
R2855	T4587	T4586	punct	", ",located
R2856	T4588	T4589	det	the,orthologues
R2857	T4589	T4586	nsubjpass	orthologues,located
R2858	T4590	T4589	nummod	three,orthologues
R2859	T4591	T4589	amod	human,orthologues
R2860	T4592	T4589	prep	of,orthologues
R2861	T4593	T4594	det	these,genes
R2862	T4594	T4592	pobj	genes,of
R2863	T4595	T4586	auxpass	are,located
R2864	T4596	T4586	advmod	also,located
R2865	T4597	T4586	prep	within,located
R2866	T4598	T4599	nummod	300,kb
R2867	T4599	T4597	pobj	kb,within
R2868	T4600	T4599	prep	of,kb
R2869	T4601	T4602	det	each,other
R2870	T4602	T4600	pobj	other,of
R2871	T4603	T4586	prep	on,located
R2872	T4604	T4603	pobj	4p16,on
R2873	T4605	T4586	punct	.,located
R2874	T4607	T4608	advmod	Furthermore,map
R2875	T4609	T4608	punct	", ",map
R2876	T4610	T4608	nsubj	TACC3,map
R2877	T4611	T4610	cc	and,TACC3
R2878	T4612	T4610	conj	FGFRL,TACC3
R2879	T4613	T4608	prep	to,map
R2880	T4614	T4615	det	the,scaffolds
R2881	T4615	T4613	pobj	scaffolds,to
R2882	T4616	T4615	amod	overlapping,scaffolds
R2883	T4617	T4618	nummod	1184,4669
R2884	T4618	T4615	appos	4669,scaffolds
R2885	T4619	T4618	punct	/,4669
R2886	T4620	T4618	punct	(,4669
R2887	T4621	T4618	appos	emb\|CAAB01004668,4669
R2888	T4622	T4618	punct	),4669
R2889	T4623	T4608	punct	.,map
R2890	T4625	T4626	advmod	Similarly,found
R2891	T4627	T4626	punct	", ",found
R2892	T4628	T4626	nsubjpass	elements,found
R2893	T4629	T4628	prep	of,elements
R2894	T4630	T4631	det	these,clusters
R2895	T4631	T4629	pobj	clusters,of
R2896	T4632	T4631	compound	gene,clusters
R2897	T4633	T4628	punct	", ",elements
R2898	T4634	T4628	acl	extending,elements
R2899	T4635	T4634	prep	from,extending
R2900	T4636	T4635	pobj	HMP19,from
R2901	T4637	T4634	prep	to,extending
R2902	T4638	T4637	pobj	GPRK6,to
R2903	T4639	T4634	prep	in,extending
R2904	T4640	T4641	amod	human,chromosome
R2905	T4641	T4642	compound	chromosome,ter
R2906	T4642	T4639	pobj	ter,in
R2907	T4643	T4642	compound	5q34,ter
R2908	T4644	T4642	punct	-,ter
R2909	T4645	T4626	auxpass	are,found
R2910	T4646	T4626	advmod	also,found
R2911	T4647	T4626	prep	in,found
R2912	T4648	T4649	det	the,pufferfish
R2913	T4649	T4647	pobj	pufferfish,in
R2914	T4650	T4626	punct	", ",found
R2915	T4651	T4626	prep	with,found
R2916	T4652	T4653	det	the,orthologues
R2917	T4653	T4651	pobj	orthologues,with
R2918	T4654	T4655	compound	T.,rubripes
R2919	T4655	T4653	compound	rubripes,orthologues
R2920	T4656	T4653	prep	of,orthologues
R2921	T4657	T4656	pobj	NSD1,of
R2922	T4658	T4653	punct	", ",orthologues
R2923	T4659	T4653	appos	FGFR4,orthologues
R2924	T4660	T4653	cc	and,orthologues
R2925	T4661	T4662	advmod	a,like
R2926	T4662	T4665	amod	like,mapping
R2927	T4663	T4662	npadvmod	RAB,like
R2928	T4664	T4662	punct	-,like
R2929	T4665	T4653	conj	mapping,orthologues
R2930	T4666	T4665	compound	gene,mapping
R2931	T4667	T4653	prep	on,orthologues
R2932	T4668	T4667	pobj	scaffold,on
R2933	T4669	T4668	nummod	407,scaffold
R2934	T4670	T4668	punct	(,scaffold
R2935	T4671	T4668	appos	emb\|CAAB01000407,scaffold
R2936	T4672	T4668	punct	),scaffold
R2937	T4673	T4626	punct	.,found
R2938	T4675	T4676	advmod	However,is
R2939	T4677	T4676	punct	", ",is
R2940	T4678	T4676	expl	there,is
R2941	T4679	T4680	det	no,evidence
R2942	T4680	T4676	attr	evidence,is
R2943	T4681	T4680	prep	for,evidence
R2944	T4682	T4683	det	a,gene
R2945	T4683	T4681	pobj	gene,for
R2946	T4684	T4683	acl	corresponding,gene
R2947	T4685	T4684	prep	to,corresponding
R2948	T4686	T4687	det	a,gene
R2949	T4687	T4685	pobj	gene,to
R2950	T4688	T4687	compound	TACC4,gene
R2951	T4689	T4676	prep	in,is
R2952	T4690	T4689	pobj	any,in
R2953	T4691	T4690	prep	of,any
R2954	T4692	T4693	det	these,clusters
R2955	T4693	T4691	pobj	clusters,of
R2956	T4694	T4676	punct	.,is
R2957	T4696	T4697	mark	As,noted
R2958	T4697	T4698	advcl	noted,indicate
R2959	T4699	T4697	advmod	above,noted
R2960	T4700	T4698	punct	", ",indicate
R2961	T4701	T4702	amod	phylogenetic,analysis
R2962	T4702	T4698	nsubj	analysis,indicate
R2963	T4703	T4702	prep	of,analysis
R2964	T4704	T4705	det	the,sequences
R2965	T4705	T4703	pobj	sequences,of
R2966	T4706	T4705	compound	TACC,sequences
R2967	T4707	T4708	mark	that,are
R2968	T4708	T4698	ccomp	are,indicate
R2969	T4709	T4708	expl	there,are
R2970	T4710	T4711	nummod	two,genes
R2971	T4711	T4708	attr	genes,are
R2972	T4712	T4713	npadvmod	TACC1,related
R2973	T4713	T4711	amod	related,genes
R2974	T4714	T4708	prep	in,are
R2975	T4715	T4716	det	the,pufferfish
R2976	T4716	T4714	pobj	pufferfish,in
R2977	T4717	T4698	punct	.,indicate
R2978	T4719	T4720	nsubjpass	trTACC1B,located
R2979	T4721	T4720	auxpass	is,located
R2980	T4722	T4720	prep	on,located
R2981	T4723	T4724	det	the,scaffold
R2982	T4724	T4722	pobj	scaffold,on
R2983	T4725	T4726	nummod	180,kb
R2984	T4726	T4724	compound	kb,scaffold
R2985	T4727	T4724	nummod	191,scaffold
R2986	T4728	T4724	punct	(,scaffold
R2987	T4729	T4724	appos	emb\|CAAB01000191.1,scaffold
R2988	T4730	T4724	punct	),scaffold
R2989	T4731	T4724	punct	", ",scaffold
R2990	T4732	T4733	dep	which,contains
R2991	T4733	T4724	relcl	contains,scaffold
R2992	T4734	T4733	advmod	also,contains
R2993	T4735	T4736	det	the,orthologues
R2994	T4736	T4733	dobj	orthologues,contains
R2995	T4737	T4736	prep	of,orthologues
R2996	T4738	T4739	amod	several,genes
R2997	T4739	T4737	pobj	genes,of
R2998	T4740	T4739	acl	located,genes
R2999	T4741	T4740	prep	in,located
R3000	T4742	T4743	amod	human,8p21
R3001	T4743	T4741	pobj	8p21,in
R3002	T4744	T4743	compound	chromosome,8p21
R3003	T4745	T4743	punct	-,8p21
R3004	T4746	T4743	nummod	11,8p21
R3005	T4747	T4720	punct	.,located
R3006	T4749	T4750	advmod	Thus,represents
R3007	T4751	T4750	punct	", ",represents
R3008	T4752	T4753	det	this,scaffold
R3009	T4753	T4750	nsubj	scaffold,represents
R3010	T4754	T4755	det	the,segment
R3011	T4755	T4750	dobj	segment,represents
R3012	T4756	T4757	advmod	more,developed
R3013	T4757	T4755	amod	developed,segment
R3014	T4758	T4757	punct	"""",developed
R3015	T4759	T4757	punct	"""",developed
R3016	T4760	T4755	nmod	TACC1,segment
R3017	T4761	T4755	amod	chromosomal,segment
R3018	T4762	T4763	dep	that,is
R3019	T4763	T4755	relcl	is,segment
R3020	T4764	T4763	acomp	evident,is
R3021	T4765	T4763	prep	in,is
R3022	T4766	T4767	amod	higher,vertebrates
R3023	T4767	T4765	pobj	vertebrates,in
R3024	T4768	T4750	punct	.,represents
R3025	T4770	T4771	prep	On,located
R3026	T4772	T4773	det	the,hand
R3027	T4773	T4770	pobj	hand,On
R3028	T4774	T4773	amod	other,hand
R3029	T4775	T4771	punct	", ",located
R3030	T4776	T4777	det	the,gene
R3031	T4777	T4771	nsubjpass	gene,located
R3032	T4778	T4777	compound	trTACC1A,gene
R3033	T4779	T4771	auxpass	is,located
R3034	T4780	T4771	prep	in,located
R3035	T4781	T4782	det	the,scaffold
R3036	T4782	T4780	pobj	scaffold,in
R3037	T4783	T4784	nummod	396,kb
R3038	T4784	T4782	compound	kb,scaffold
R3039	T4785	T4782	nummod	12,scaffold
R3040	T4786	T4782	punct	(,scaffold
R3041	T4787	T4782	appos	emb\|CAAB010012.1,scaffold
R3042	T4788	T4782	punct	),scaffold
R3043	T4789	T4771	punct	.,located
R3044	T4791	T4792	det	This,scaffold
R3045	T4792	T4793	nsubj	scaffold,contains
R3046	T4794	T4793	advmod	also,contains
R3047	T4795	T4796	det	the,orthologues
R3048	T4796	T4793	dobj	orthologues,contains
R3049	T4797	T4798	compound	T.,rubripes
R3050	T4798	T4796	compound	rubripes,orthologues
R3051	T4799	T4796	prep	of,orthologues
R3052	T4800	T4799	pobj	MSX1,of
R3053	T4801	T4800	punct	", ",MSX1
R3054	T4802	T4800	conj	STX18,MSX1
R3055	T4803	T4802	punct	", ",STX18
R3056	T4804	T4802	conj	D4S234E,STX18
R3057	T4805	T4804	cc	and,D4S234E
R3058	T4806	T4807	det	the,gene
R3059	T4807	T4804	conj	gene,D4S234E
R3060	T4808	T4807	amod	predicted,gene
R3061	T4809	T4807	appos	LOC118711,gene
R3062	T4810	T4793	punct	", ",contains
R3063	T4811	T4793	prep	in,contains
R3064	T4812	T4811	pobj	addition,in
R3065	T4813	T4812	prep	to,addition
R3066	T4814	T4813	pobj	sequences,to
R3067	T4815	T4814	prep	with,sequences
R3068	T4816	T4815	pobj	homology,with
R3069	T4817	T4816	prep	to,homology
R3070	T4818	T4817	pobj	LOXL,to
R3071	T4819	T4818	punct	", ",LOXL
R3072	T4820	T4818	conj	EVC,LOXL
R3073	T4821	T4820	punct	", ",EVC
R3074	T4822	T4820	conj	LOC159291,EVC
R3075	T4823	T4822	punct	", ",LOC159291
R3076	T4824	T4822	cc	and,LOC159291
R3077	T4825	T4826	det	the,family
R3078	T4826	T4822	conj	family,LOC159291
R3079	T4827	T4826	compound	LDB,family
R3080	T4828	T4793	punct	.,contains
R3081	T4830	T4831	advmod	Thus,contains
R3082	T4832	T4831	punct	", ",contains
R3083	T4833	T4831	nsubj	scaffold,contains
R3084	T4834	T4833	nummod	12,scaffold
R3085	T4835	T4831	dobj	genes,contains
R3086	T4836	T4835	acl	found,genes
R3087	T4837	T4836	prep	in,found
R3088	T4838	T4839	det	the,regions
R3089	T4839	T4837	pobj	regions,in
R3090	T4840	T4839	prep	of,regions
R3091	T4841	T4842	amod	human,4
R3092	T4842	T4840	pobj	4,of
R3093	T4843	T4842	nmod	chromosome,4
R3094	T4844	T4842	cc	and,4
R3095	T4845	T4842	conj	10,4
R3096	T4846	T4847	dep	that,contain
R3097	T4847	T4835	relcl	contain,genes
R3098	T4848	T4847	advmod	also,contain
R3099	T4849	T4850	det	the,loci
R3100	T4850	T4847	dobj	loci,contain
R3101	T4851	T4850	prep	for,loci
R3102	T4852	T4851	pobj	TACC3,for
R3103	T4853	T4852	cc	and,TACC3
R3104	T4854	T4852	conj	TACC2,TACC3
R3105	T4855	T4847	punct	", ",contain
R3106	T4856	T4847	advmod	respectively,contain
R3107	T4857	T4831	punct	", ",contains
R3108	T4858	T4831	cc	and,contains
R3109	T4859	T4860	aux	may,resemble
R3110	T4860	T4831	conj	resemble,contains
R3111	T4861	T4860	advmod	therefore,resemble
R3112	T4862	T4863	advmod	more,closely
R3113	T4863	T4860	advmod	closely,resemble
R3114	T4864	T4865	det	the,organization
R3115	T4865	T4860	dobj	organization,resemble
R3116	T4866	T4865	amod	genomic,organization
R3117	T4867	T4865	acl	resulting,organization
R3118	T4868	T4867	prep	from,resulting
R3119	T4869	T4870	det	the,duplication
R3120	T4870	T4868	pobj	duplication,from
R3121	T4871	T4870	amod	initial,duplication
R3122	T4872	T4870	prep	of,duplication
R3123	T4873	T4874	det	the,segment
R3124	T4874	T4872	pobj	segment,of
R3125	T4875	T4874	amod	ancestral,segment
R3126	T4876	T4874	amod	paralogous,segment
R3127	T4877	T4874	amod	chromosomal,segment
R3128	T4878	T4831	punct	.,contains
R3129	T4880	T4881	amod	Conserved,clusters
R3130	T4881	T4883	nsubj	clusters,result
R3131	T4882	T4881	amod	paralogous,clusters
R3132	T4884	T4883	aux	may,result
R3133	T4885	T4883	prep	from,result
R3134	T4886	T4887	det	the,clustering
R3135	T4887	T4885	pobj	clustering,from
R3136	T4888	T4887	amod	initial,clustering
R3137	T4889	T4887	prep	of,clustering
R3138	T4890	T4891	det	the,genes
R3139	T4891	T4889	pobj	genes,of
R3140	T4892	T4887	prep	in,clustering
R3141	T4893	T4894	det	a,contig
R3142	T4894	T4892	pobj	contig,in
R3143	T4895	T4896	advmod	relatively,small
R3144	T4896	T4894	amod	small,contig
R3145	T4897	T4894	amod	ancestral,contig
R3146	T4898	T4894	amod	genomic,contig
R3147	T4899	T4883	punct	.,result
R3148	T4901	T4902	det	Some,evidence
R3149	T4902	T4903	nsubj	evidence,is
R3150	T4904	T4902	prep	for,evidence
R3151	T4905	T4906	det	the,existence
R3152	T4906	T4904	pobj	existence,for
R3153	T4907	T4906	prep	of,existence
R3154	T4908	T4909	punct	"""",protoclusters
R3155	T4909	T4907	pobj	protoclusters,of
R3156	T4910	T4909	punct	"""",protoclusters
R3157	T4911	T4912	dep	that,correspond
R3158	T4912	T4909	relcl	correspond,protoclusters
R3159	T4913	T4912	aux	could,correspond
R3160	T4914	T4912	prep	to,correspond
R3161	T4915	T4916	det	the,segments
R3162	T4916	T4914	pobj	segments,to
R3163	T4917	T4916	amod	paralogous,segments
R3164	T4918	T4916	amod	chromosomal,segments
R3165	T4919	T4916	acl	noted,segments
R3166	T4920	T4919	prep	in,noted
R3167	T4921	T4922	amod	higher,vertebrates
R3168	T4922	T4920	pobj	vertebrates,in
R3169	T4923	T4903	acomp	present,is
R3170	T4924	T4903	prep	in,is
R3171	T4925	T4926	det	the,genome
R3172	T4926	T4924	pobj	genome,in
R3173	T4927	T4926	prep	of,genome
R3174	T4928	T4929	det	the,urochordate
R3175	T4929	T4927	pobj	urochordate,of
R3176	T4930	T4931	compound	C.,intestinalis
R3177	T4931	T4929	appos	intestinalis,urochordate
R3178	T4932	T4933	punct	[,11
R3179	T4933	T4903	parataxis	11,is
R3180	T4934	T4933	punct	],11
R3181	T4935	T4903	punct	.,is
R3182	T4937	T4938	prep	For,cluster
R3183	T4939	T4937	pobj	instance,For
R3184	T4940	T4938	punct	", ",cluster
R3185	T4941	T4942	det	the,orthologues
R3186	T4942	T4938	nsubj	orthologues,cluster
R3187	T4943	T4942	prep	of,orthologues
R3188	T4944	T4943	pobj	FGFR,of
R3189	T4945	T4944	punct	", ",FGFR
R3190	T4946	T4944	cc	and,FGFR
R3191	T4947	T4944	conj	WHSC1,FGFR
R3192	T4948	T4942	punct	", ",orthologues
R3193	T4949	T4950	compound	carboxypeptidase,Z
R3194	T4950	T4942	appos	Z,orthologues
R3195	T4951	T4950	cc	and,Z
R3196	T4952	T4950	conj	FLJ25359,Z
R3197	T4953	T4938	prep	within,cluster
R3198	T4954	T4955	det	an,region
R3199	T4955	T4953	pobj	region,within
R3200	T4956	T4957	nummod	85,kb
R3201	T4957	T4955	compound	kb,region
R3202	T4958	T4955	prep	of,region
R3203	T4959	T4960	det	the,genome
R3204	T4960	T4958	pobj	genome,of
R3205	T4961	T4962	compound	C.,intestinalis
R3206	T4962	T4960	compound	intestinalis,genome
R3207	T4963	T4938	cc	and,cluster
R3208	T4964	T4965	det	the,orthologues
R3209	T4965	T4967	nsubjpass	orthologues,maintained
R3210	T4966	T4965	amod	human,orthologues
R3211	T4967	T4938	conj	maintained,cluster
R3212	T4968	T4967	auxpass	are,maintained
R3213	T4969	T4967	advmod	still,maintained
R3214	T4970	T4967	prep	in,maintained
R3215	T4971	T4972	amod	paralogous,segments
R3216	T4972	T4970	pobj	segments,in
R3217	T4973	T4972	prep	of,segments
R3218	T4974	T4973	pobj	4p16,of
R3219	T4975	T4974	punct	", ",4p16
R3220	T4976	T4974	conj	8p,4p16
R3221	T4977	T4976	cc	and,8p
R3222	T4978	T4976	conj	10q,8p
R3223	T4979	T4980	punct	(,Fig.
R3224	T4980	T4967	parataxis	Fig.,maintained
R3225	T4981	T4980	nummod	3,Fig.
R3226	T4982	T4980	punct	", ",Fig.
R3227	T4983	T4980	punct	[,Fig.
R3228	T4984	T4980	advcl	see,Fig.
R3229	T4985	T4986	amod	Additional,file
R3230	T4986	T4984	dobj	file,see
R3231	T4987	T4986	nummod	1,file
R3232	T4988	T4980	punct	],Fig.
R3233	T4989	T4980	punct	),Fig.
R3234	T4990	T4938	punct	.,cluster
R3235	T4992	T4993	advmod	However,noted
R3236	T4994	T4993	punct	", ",noted
R3237	T4995	T4993	nsubjpass	it,noted
R3238	T4996	T4993	aux	should,noted
R3239	T4997	T4993	auxpass	be,noted
R3240	T4998	T4999	mark	that,locate
R3241	T4999	T4993	ccomp	locate,noted
R3242	T5000	T5001	det	no,clusters
R3243	T5001	T4999	nsubjpass	clusters,locate
R3244	T5002	T5001	prep	of,clusters
R3245	T5003	T5002	pobj	genes,of
R3246	T5004	T5003	prep	from,genes
R3247	T5005	T5006	det	the,segments
R3248	T5006	T5004	pobj	segments,from
R3249	T5007	T5006	nmod	vertebrate,segments
R3250	T5008	T5006	amod	paralogous,segments
R3251	T5009	T4999	auxpass	are,locate
R3252	T5010	T4999	advmod	close,locate
R3253	T5011	T5010	prep	to,close
R3254	T5012	T5013	det	the,genes
R3255	T5013	T5011	pobj	genes,to
R3256	T5014	T5013	nmod	TACC,genes
R3257	T5015	T5014	cc	or,TACC
R3258	T5016	T5014	conj	RHAMM,TACC
R3259	T5017	T5013	prep	of,genes
R3260	T5018	T5019	compound	C.,intestinalis
R3261	T5019	T5017	pobj	intestinalis,of
R3262	T5020	T4999	punct	", ",locate
R3263	T5021	T4999	advcl	indicating,locate
R3264	T5022	T5023	mark	that,formed
R3265	T5023	T5021	ccomp	formed,indicating
R3266	T5024	T5025	det	the,formation
R3267	T5025	T5023	nsubj	formation,formed
R3268	T5026	T5025	prep	of,formation
R3269	T5027	T5028	det	the,segments
R3270	T5028	T5026	pobj	segments,of
R3271	T5029	T5030	advmod	much,larger
R3272	T5030	T5028	amod	larger,segments
R3273	T5031	T5028	amod	paralogous,segments
R3274	T5032	T5028	acl	encompassing,segments
R3275	T5033	T5034	det	the,genes
R3276	T5034	T5032	dobj	genes,encompassing
R3277	T5035	T5036	compound	FGFR,TACC
R3278	T5036	T5034	compound	TACC,genes
R3279	T5037	T5036	punct	-,TACC
R3280	T5038	T5023	advmod	later,formed
R3281	T5039	T5038	prep	in,later
R3282	T5040	T5041	amod	evolutionary,time
R3283	T5041	T5039	pobj	time,in
R3284	T5042	T5023	punct	", ",formed
R3285	T5043	T5023	cc	or,formed
R3286	T5044	T5045	advmod	conversely,been
R3287	T5045	T5023	conj	been,formed
R3288	T5046	T5045	aux	have,been
R3289	T5047	T5045	acomp	subject,been
R3290	T5048	T5047	prep	to,subject
R3291	T5049	T5050	amod	extensive,rearrangement
R3292	T5050	T5048	pobj	rearrangement,to
R3293	T5051	T5045	prep	in,been
R3294	T5052	T5051	pobj	tunicates,in
R3295	T5053	T4993	punct	.,noted
R3296	T5055	T5056	prep	In,provides
R3297	T5057	T5055	pobj	combination,In
R3298	T5058	T5057	prep	with,combination
R3299	T5059	T5060	det	the,examination
R3300	T5060	T5058	pobj	examination,with
R3301	T5061	T5060	prep	of,examination
R3302	T5062	T5063	det	the,genome
R3303	T5063	T5061	pobj	genome,of
R3304	T5064	T5065	compound	T.,rubripes
R3305	T5065	T5063	compound	rubripes,genome
R3306	T5066	T5056	punct	", ",provides
R3307	T5067	T5056	nsubj	this,provides
R3308	T5068	T5056	advmod	also,provides
R3309	T5069	T5070	amod	additional,evidence
R3310	T5070	T5056	dobj	evidence,provides
R3311	T5071	T5072	mark	that,include
R3312	T5072	T5070	advcl	include,evidence
R3313	T5073	T5072	preconj	either,include
R3314	T5074	T5075	det	the,round
R3315	T5075	T5072	nsubj	round,include
R3316	T5076	T5075	amod	second,round
R3317	T5077	T5075	prep	of,round
R3318	T5078	T5077	pobj	duplication,of
R3319	T5079	T5078	prep	of,duplication
R3320	T5080	T5081	det	the,segment
R3321	T5081	T5079	pobj	segment,of
R3322	T5082	T5081	amod	chromosomal,segment
R3323	T5083	T5084	dep	that,contained
R3324	T5084	T5081	relcl	contained,segment
R3325	T5085	T5086	det	the,ancestor
R3326	T5086	T5084	dobj	ancestor,contained
R3327	T5087	T5086	nmod	FGFR3,ancestor
R3328	T5088	T5087	punct	/,FGFR3
R3329	T5089	T5087	nummod	4,FGFR3
R3330	T5090	T5072	aux	did,include
R3331	T5091	T5072	neg	not,include
R3332	T5092	T5093	det	a,gene
R3333	T5093	T5072	dobj	gene,include
R3334	T5094	T5093	compound	TACC,gene
R3335	T5095	T5072	punct	", ",include
R3336	T5096	T5072	cc	or,include
R3337	T5097	T5098	mark	that,lost
R3338	T5098	T5072	conj	lost,include
R3339	T5099	T5100	predet	such,gene
R3340	T5100	T5098	nsubjpass	gene,lost
R3341	T5101	T5100	det	a,gene
R3342	T5102	T5098	auxpass	was,lost
R3343	T5103	T5104	advmod	very,early
R3344	T5104	T5098	advmod	early,lost
R3345	T5105	T5104	prep	in,early
R3346	T5106	T5107	compound	vertebrate,evolution
R3347	T5107	T5105	pobj	evolution,in
R3348	T5108	T5098	punct	", ",lost
R3349	T5109	T5098	advmod	prior,lost
R3350	T5110	T5109	prep	to,prior
R3351	T5111	T5112	det	the,divergence
R3352	T5112	T5110	pobj	divergence,to
R3353	T5113	T5112	prep	of,divergence
R3354	T5114	T5115	det	the,lineages
R3355	T5115	T5113	pobj	lineages,of
R3356	T5116	T5115	compound	Gnanthostome,lineages
R3357	T5117	T5056	punct	.,provides
R3358	T5119	T5120	advmod	However,await
R3359	T5121	T5120	punct	", ",await
R3360	T5122	T5123	det	the,resolution
R3361	T5123	T5120	nsubj	resolution,await
R3362	T5124	T5123	amod	final,resolution
R3398	T5160	T5159	amod	other,genes
R3399	T5161	T5159	amod	corresponding,genes
R3400	T5162	T5152	prep	in,contain
R3401	T5163	T5164	det	this,segment
R3402	T5164	T5162	pobj	segment,in
R3403	T5165	T5164	amod	conserved,segment
R3404	T5166	T5120	punct	.,await
R2693	T4414	T4412	punct	),Fig.
R2694	T4415	T4398	punct	", ",located
R2695	T4416	T4417	advmod	further,suggesting
R2696	T4417	T4398	advcl	suggesting,located
R2697	T4418	T4419	mark	that,transposed
R2698	T4419	T4417	ccomp	transposed,suggesting
R2699	T4420	T4421	det	this,cluster
R2673	T4392	T4391	acomp	paralogous,are
R2674	T4393	T4392	prep	with,paralogous
R2675	T4394	T4393	pobj	4p16,with
R2676	T4395	T4381	punct	.,separate
R2677	T4397	T4398	advmod	Interestingly,located
R2678	T4399	T4398	punct	", ",located
R2679	T4400	T4401	det	these,clusters
R2680	T4401	T4398	nsubjpass	clusters,located
R2681	T4402	T4401	nummod	three,clusters
R2682	T4403	T4398	auxpass	are,located
R2683	T4404	T4398	prep	on,located
R2684	T4405	T4406	amod	different,chromosomes
R2685	T4406	T4404	pobj	chromosomes,on
R2686	T4407	T4398	prep	in,located
R2687	T4408	T4407	pobj	mouse,in
R2688	T4409	T4408	cc	and,mouse
R2689	T4410	T4408	conj	rat,mouse
R2690	T4411	T4412	punct	(,Fig.
R2691	T4412	T4398	parataxis	Fig.,located
R2692	T4413	T4412	nummod	2,Fig.
R2700	T4421	T4419	nsubjpass	cluster,transposed
R2701	T4422	T4421	prep	of,cluster
R2702	T4423	T4422	pobj	genes,of
R2703	T4424	T4419	auxpass	was,transposed
R2704	T4425	T4419	prep	into,transposed
R2705	T4426	T4427	det	this,region
R2706	T4427	T4425	pobj	region,into
R2707	T4428	T4419	prep	after,transposed
R2708	T4429	T4430	det	the,divergence
R2709	T4430	T4428	pobj	divergence,after
R2710	T4431	T4432	compound	primate,rodent
R2711	T4432	T4430	compound	rodent,divergence
R2712	T4433	T4432	punct	/,rodent
R2714	T4437	T4438	mark	Because,provide
R3363	T5125	T5123	prep	of,resolution
R3364	T5126	T5127	det	the,evolution
R3365	T5127	T5125	pobj	evolution,of
R3366	T5128	T5127	amod	initial,evolution
R3367	T5129	T5127	prep	of,evolution
R3368	T5130	T5131	det	these,segment
R3369	T5131	T5129	pobj	segment,of
R3370	T5132	T5131	amod	paralogous,segment
R3371	T5133	T5120	aux	will,await
R3372	T5134	T5135	det	the,sequencing
R3373	T5135	T5120	dobj	sequencing,await
R3374	T5136	T5135	prep	of,sequencing
R3375	T5137	T5138	det	the,genomes
R3376	T5138	T5136	pobj	genomes,of
R3377	T5139	T5138	nmod	amphioxus,genomes
R3378	T5140	T5139	cc	and,amphioxus
R3379	T5141	T5139	conj	lamprey,amphioxus
R3380	T5142	T5138	punct	", ",genomes
R3381	T5143	T5144	dep	which,have
R3382	T5144	T5138	relcl	have,genomes
R3383	T5145	T5144	advmod	only,have
R3384	T5146	T5147	nummod	one,gene
R3385	T5147	T5144	dobj	gene,have
R3386	T5148	T5147	compound	FGFR,gene
R3387	T5149	T5144	punct	", ",have
R3388	T5150	T5144	cc	and,have
R3389	T5151	T5152	advmod	therefore,contain
R3390	T5152	T5144	conj	contain,have
R3391	T5153	T5152	aux	should,contain
R3392	T5154	T5152	advmod	only,contain
R3393	T5155	T5156	nummod	one,copy
R3394	T5156	T5152	dobj	copy,contain
R3395	T5157	T5156	prep	of,copy
R3396	T5158	T5159	det	the,genes
R3397	T5159	T5157	pobj	genes,of

craft-ca-core-ex-dev

Below, discontinuous spans are shown in the chain model. You can change it to the bag model.

Id	Subject	Object	Predicate	Lexical cue
T3973	611-621	GO_SO_EXT:chromosome	denotes	chromosome
T3974	640-645	NCBITaxon:9606	denotes	human
T3975	646-656	GO_SO_EXT:chromosome	denotes	chromosome
T3976	720-732	NCBITaxon_EXT:invertebrate	denotes	invertebrate
T3977	778-782	GO_PR_EXT:fibroblast_growth_factor_receptor	denotes	FGFR
T3978	788-792	SO_EXT:0000704	denotes	gene
T3979	839-850	SO_EXT:sequence_duplication_entity_or_process	denotes	duplication
T3980	886-896	_FRAGMENT	denotes	paralogous
T3981	909-917	SO_EXT:0000854	denotes	segments
T3982	897-917	GO_SO_EXT:chromosomal_part_or_position_or_region_or_site	denotes	chromosomal segments
T3983	1013-1017	SO_EXT:0000704	denotes	gene
T3984	1021-1032	NCBITaxon:7742	denotes	vertebrates
T3985	1058-1076	GO_SO_EXT:chromosomal_part_or_position_or_region_or_site	denotes	chromosomal region
T3986	1080-1085	PR_EXT:000001451	denotes	FGFR4
T3987	1143-1148	PR_EXT:000001856	denotes	RHAMM
T3988	1149-1153	SO_EXT:0000704	denotes	gene
T3989	1191-1196	NCBITaxon:9606	denotes	Human
T3990	1197-1202	PR_EXT:000001856	denotes	RHAMM
T3991	1211-1221	GO_SO_EXT:chromosome	denotes	chromosome
T3992	1250-1254	PR_EXT:000008212	denotes	GPX3
T3993	1259-1264	PR_EXT:000011245	denotes	NKX2E
T3994	1302-1307	SO_EXT:0000704	denotes	genes
T3995	1311-1316	NCBITaxon:9606	denotes	human
T3996	1317-1327	GO_SO_EXT:chromosome	denotes	chromosome
T3997	1339-1349	SO:0000859	denotes	paralogous
T3998	1422-1433	GO_SO_EXT:chromosome	denotes	chromosomes
T3999	1437-1442	NCBITaxon:10088	denotes	mouse
T4000	1447-1450	NCBITaxon:10114	denotes	rat
T4001	1501-1506	SO_EXT:0000704	denotes	genes
T4002	1511-1521	GO_SO_EXT:sequence_transposition_process	denotes	transposed
T4003	1549-1556	NCBITaxon:9443	denotes	primate
T4004	1557-1563	NCBITaxon:9989	denotes	rodent
T4006	2274-2278	SO_EXT:0000704	denotes	gene
T4007	2328-2332	SO_EXT:0000704	denotes	gene
T4008	2333-2343	GO:0065007	denotes	regulation
T4009	2395-2414	SO_EXT:0000854	denotes	paralogous segments
T4010	2432-2438	SO_EXT:0001026	denotes	genome
T4011	2446-2454	NCBITaxon:7712	denotes	tunicate
T4012	2455-2470	NCBITaxon:7719	denotes	C. intestinalis
T4013	2509-2519	NCBITaxon:7742	denotes	vertebrate
T4014	2520-2526	SO_EXT:0001026	denotes	genome
T4015	2546-2557	NCBITaxon:31033	denotes	T. rubripes
T4016	2613-2619	SO_EXT:0001026	denotes	genome
T4017	2623-2634	NCBITaxon:31033	denotes	T. rubripes
T4018	2661-2681	GO_SO_EXT:chromosomal_part_or_position_or_region_or_site	denotes	chromosomal segments
T4019	2682-2692	SO:0000859	denotes	paralogous
T4020	2718-2729	NCBITaxon:7742	denotes	vertebrates
T4021	2758-2769	SO_EXT:0000855	denotes	orthologues
T4022	2773-2779	PR_EXT:000008258	denotes	GPRK2L
T4023	2784-2789	PR_EXT:000013947	denotes	RGS12
T4024	2803-2814	NCBITaxon:31033	denotes	T. rubripes
T4025	2815-2823	SO_EXT:0001248	denotes	scaffold
T4026	2866-2867	CHEBI_SO_EXT:base	denotes	b
T4027	2885-2890	NCBITaxon:9606	denotes	human
T4028	2901-2912	NCBITaxon:31033	denotes	T. rubripes
T4029	2913-2924	SO_EXT:0000855	denotes	orthologues
T4030	2928-2933	PR_EXT:000001450	denotes	FGFR3
T4031	2980-2981	CHEBI_SO_EXT:base	denotes	b
T4032	2982-2989	SO_EXT:0001026	denotes	genomic
T4033	2990-2998	SO_EXT:0001248	denotes	scaffold
T4034	3050-3055	NCBITaxon:9606	denotes	human
T4035	3056-3067	SO_EXT:0000855	denotes	orthologues
T4036	3077-3082	SO_EXT:0000704	denotes	genes
T4037	3112-3113	CHEBI_SO_EXT:base	denotes	b
T4038	3150-3155	PR_EXT:000016008	denotes	TACC3
T4039	3160-3165	PR_EXT:000007506	denotes	FGFRL
T4040	3189-3198	SO_EXT:0001248	denotes	scaffolds
T4041	3258-3262	SO_EXT:0000704	denotes	gene
T4042	3297-3302	PR_EXT:000008260	denotes	GPRK6
T4043	3306-3311	NCBITaxon:9606	denotes	human
T4044	3312-3322	GO_SO_EXT:chromosome	denotes	chromosome
T4045	3354-3364	NCBITaxon:31031	denotes	pufferfish
T4046	3375-3386	NCBITaxon:31033	denotes	T. rubripes
T4047	3387-3398	SO_EXT:0000855	denotes	orthologues
T4048	3402-3406	PR_EXT:000011441	denotes	NSD1
T4049	3408-3413	PR_EXT:000001451	denotes	FGFR4
T4050	3429-3433	SO_EXT:0000704	denotes	gene
T4051	3445-3453	SO_EXT:0001248	denotes	scaffold
T4052	3514-3518	SO_EXT:0000704	denotes	gene
T4053	3544-3548	SO_EXT:0000704	denotes	gene
T4054	3625-3634	SO_EXT:biological_sequence	denotes	sequences
T4055	3663-3668	PR_EXT:000016006	denotes	TACC1
T4056	3677-3682	SO_EXT:0000704	denotes	genes
T4057	3690-3700	NCBITaxon:31031	denotes	pufferfish
T4058	3734-3735	CHEBI_SO_EXT:base	denotes	b
T4059	3736-3744	SO_EXT:0001248	denotes	scaffold
T4060	3795-3806	SO_EXT:0000855	denotes	orthologues
T4061	3818-3823	SO_EXT:0000704	denotes	genes
T4062	3835-3840	NCBITaxon:9606	denotes	human
T4063	3841-3851	GO_SO_EXT:chromosome	denotes	chromosome
T4064	3872-3880	SO_EXT:0001248	denotes	scaffold
T4065	3913-3918	PR_EXT:000016006	denotes	TACC1
T4066	3919-3938	GO_SO_EXT:chromosomal_part_or_position_or_region_or_site	denotes	chromosomal segment
T4067	3965-3976	NCBITaxon:7742	denotes	vertebrates
T4068	4010-4014	SO_EXT:0000704	denotes	gene
T4069	4038-4039	CHEBI_SO_EXT:base	denotes	b
T4070	4040-4048	SO_EXT:0001248	denotes	scaffold
T4071	4077-4085	SO_EXT:0001248	denotes	scaffold
T4072	4104-4115	NCBITaxon:31033	denotes	T. rubripes
T4073	4116-4127	SO_EXT:0000855	denotes	orthologues
T4074	4131-4135	PR_EXT:000010686	denotes	MSX1
T4075	4137-4142	PR_EXT:000015790	denotes	STX18
T4076	4160-4174	SO_EXT:0000996	denotes	predicted gene
T4077	4201-4210	SO_EXT:biological_sequence	denotes	sequences
T4078	4216-4224	SO:0000857	denotes	homology
T4079	4228-4232	PR_EXT:000009884	denotes	LOXL
T4080	4234-4237	PR_EXT:000007232	denotes	EVC
T4081	4276-4284	SO_EXT:0001248	denotes	scaffold
T4082	4297-4302	SO_EXT:0000704	denotes	genes
T4083	4316-4326	_FRAGMENT	denotes	regions of
T4084	4333-4343	GO_SO_EXT:chromosomal_part_or_position_or_region_or_site	denotes	chromosome
T4085	4327-4332	NCBITaxon:9606	denotes	human
T4086	4384-4389	PR_EXT:000016008	denotes	TACC3
T4087	4394-4399	PR_EXT:000016007	denotes	TACC2
T4088	4459-4466	SO_EXT:0001026	denotes	genomic
T4089	4507-4518	SO_EXT:sequence_duplication_entity_or_process	denotes	duplication
T4119	5298-5303	PR_EXT:000001856	denotes	RHAMM
T4120	5304-5309	SO_EXT:0000704	denotes	genes
T4121	5313-5328	NCBITaxon:7719	denotes	C. intestinalis
T4122	5379-5398	SO_EXT:0000854	denotes	paralogous segments
T4123	5416-5420	GO_PR_EXT:fibroblast_growth_factor_receptor	denotes	FGFR
T4124	5426-5431	SO_EXT:0000704	denotes	genes
T4125	5512-5525	GO_SO_EXT:sequence_rearrangement_entity_or_process	denotes	rearrangement
T4126	5529-5538	NCBITaxon:7712	denotes	tunicates
T4127	5583-5594	NCBITaxon:31033	denotes	T. rubripes
T4128	5595-5601	SO_EXT:0001026	denotes	genome
T4129	5674-5685	SO_EXT:sequence_duplication_entity_or_process	denotes	duplication
T4130	5693-5712	GO_SO_EXT:chromosomal_part_or_position_or_region_or_site	denotes	chromosomal segment
T4131	5732-5737	PR_EXT:000001450	denotes	FGFR3
T4132	5772-5776	SO_EXT:0000704	denotes	gene
T4133	5793-5797	SO_EXT:0000704	denotes	gene
T4134	5821-5831	NCBITaxon:7742	denotes	vertebrate
T4135	5874-5886	NCBITaxon:61463	denotes	Gnanthostome
T4136	5961-5979	SO_EXT:0000854	denotes	paralogous segment
T4137	6013-6022	NCBITaxon:7737	denotes	amphioxus
T4138	6027-6034	NCBITaxon:7745	denotes	lamprey
T4139	6035-6042	SO_EXT:0001026	denotes	genomes
T4140	6064-6068	GO_PR_EXT:fibroblast_growth_factor_receptor	denotes	FGFR
T4141	6069-6073	SO_EXT:0000704	denotes	gene
T4142	6113-6117	SO_EXT:sequence_copy_entity	denotes	copy
T4143	6145-6150	SO_EXT:0000704	denotes	genes
T4144	6159-6168	SO_EXT:biological_conservation_process_or_quality	denotes	conserved
T12592	1783-1789	_FRAGMENT	denotes	region
T12593	1813-1826	GO_SO_EXT:chromosomal_part_or_position_or_region_or_site	denotes	on chromosome
T12594	1798-1802	PR_EXT:000008212	denotes	GPX3
T12595	1807-1812	PR_EXT:000011245	denotes	NKX2E
T12596	1816-1826	GO_SO_EXT:chromosome	denotes	chromosome
T12597	1881-1886	SO_EXT:0000704	denotes	genes
T12598	1887-1895	SO_EXT:sequence_insertion_process	denotes	inserted
T12599	1902-1913	SO_EXT:sequence_duplication_entity_or_process	denotes	duplication
T12600	1948-1953	SO_EXT:0000704	denotes	genes
T12601	1989-1997	_FRAGMENT	denotes	syntenic
T12602	2016-2023	SO_EXT:0005858	denotes	regions
T12603	1998-2003	NCBITaxon:10088	denotes	mouse
T12604	2004-2023	GO_SO_EXT:chromosomal_part_or_position_or_region_or_site	denotes	chromosomal regions
T12605	2045-2062	NCBITaxon:31033	denotes	Takifugu rubripes
T12606	2063-2072	SO_EXT:0001248	denotes	scaffolds
T12607	2116-2126	SO:0000857	denotes	homologous
T12608	2127-2131	SO_EXT:0000704	denotes	gene
T12609	2188-2198	SO:0000859	denotes	paralogous
T12610	2199-2204	SO_EXT:0000704	denotes	genes
T12784	6218-6235	NCBITaxon:31033	denotes	Takifugu rubripes
T12785	6240-6258	NCBITaxon:7719	denotes	Ciona intestinalis
T12786	6282-6289	SO_EXT:0001026	denotes	genomic
T12787	6290-6299	SO_EXT:0001248	denotes	scaffolds
T12788	6315-6332	NCBITaxon:31033	denotes	Takifugu rubripes
T12789	6355-6360	SO_EXT:0000704	denotes	genes
T12790	6362-6370	SO_EXT:0001248	denotes	Scaffold
T12791	6401-6405	SO_EXT:0000704	denotes	gene
T12792	6415-6420	SO_EXT:0000704	denotes	genes
T12793	6439-6449	SO_EXT:0000853	denotes	homologues
T12794	6453-6464	SO_EXT:0000855	denotes	orthologues
T12795	6484-6490	_FRAGMENT	denotes	arm of
T12796	6497-6507	SO_EXT:0000105	denotes	chromosome
T12797	6491-6496	NCBITaxon:9606	denotes	human
T12798	6497-6507	GO_SO_EXT:chromosome	denotes	chromosome
T12799	6526-6534	SO_EXT:0001248	denotes	scaffold
T12800	6622-6627	PR_EXT:000016006	denotes	TACC1
T12801	6628-6633	PR_EXT:000016007	denotes	TACC2
T12802	6634-6653	GO_SO_EXT:chromosomal_part_or_position_or_region_or_site	denotes	chromosomal segment
T12803	6676-6684	SO_EXT:0001248	denotes	scaffold
T12804	6705-6716	SO_EXT:0000855	denotes	orthologues
T12805	6720-6725	SO_EXT:0000704	denotes	genes
T12806	6754-6760	_FRAGMENT	denotes	arm of
T12807	6767-6777	SO_EXT:0000105	denotes	chromosome
T12808	6761-6766	NCBITaxon:9606	denotes	human
T12809	6767-6777	GO_SO_EXT:chromosome	denotes	chromosome
T12810	6786-6804	NCBITaxon:7719	denotes	Ciona intestinalis
T12811	6825-6830	SO_EXT:0000704	denotes	genes
T12812	6840-6859	SO_EXT:0000854	denotes	paralogous segments
T12813	6863-6868	NCBITaxon:9606	denotes	human
T12970	6900-6907	SO_EXT:0001026	denotes	Genomic
T12971	6930-6935	SO_EXT:0000704	denotes	genes
T12972	6937-6946	SO_EXT:biological_conservation_process_or_quality	denotes	Conserved
T12973	6964-6968	CHEMINF_GO_EXT:chemical_binding_or_bond_formation	denotes	bind
T12974	6969-6976	CHEBI_PR_EXT:protein	denotes	protein
T12975	6992-6997	NCBITaxon:9606	denotes	human
T12976	7003-7011	CHEBI_PR_EXT:protein	denotes	proteins
T12977	7031-7037	SO_EXT:sequence_repeat_unit_or_region	denotes	repeat
T12978	7041-7046	NCBITaxon:9606	denotes	human
T12979	7047-7052	PR_EXT:000016006	denotes	TACC1
T12980	7067-7071	CHEMINF_GO_EXT:chemical_binding_or_bond_formation	denotes	bind
T12981	7072-7077	PR_EXT:000017527	denotes	GAS41
T12982	7116-7122	SO_EXT:0000417	denotes	domain
T12983	7123-7128	CHEMINF_GO_EXT:chemical_binding_or_bond_formation	denotes	binds
T12984	7129-7135	PR_EXT:000005512	denotes	ch-TOG
T12985	7155-7169	PR_EXT:000035365	denotes	Aurora kinases
T12986	7162-7169	GO_EXT:0016301	denotes	kinases
T12987	7177-7184	NCBITaxon:species	denotes	species
T12988	7208-7213	SO_EXT:sequence_or_structure_motif	denotes	motif
T12989	7236-7243	SO_EXT:sequence_coding_function	denotes	encoded
T12990	7254-7259	SO_EXT:0000147	denotes	exons
T12991	7272-7277	SO_EXT:0000704	denotes	genes
T12992	7303-7310	SO_EXT:0001060	denotes	isoform
T12993	7320-7324	SO_EXT:0000704	denotes	gene
T12584	1586-1592	SO:0000987	denotes	Linear
T12585	1609-1613	SO_EXT:0000704	denotes	gene
T12586	1642-1658	GO_SO_EXT:chromosomal_part_or_position_or_region_or_site	denotes	chromosomal loci
T12587	1666-1670	GO_PR_EXT:fibroblast_growth_factor_receptor	denotes	FGFR
T12588	1671-1676	SO_EXT:0000704	denotes	genes
T12589	1680-1686	NCBITaxon:9606	denotes	humans
T12590	1688-1698	SO:0000859	denotes	Paralogous
T12591	1699-1704	SO_EXT:0000704	denotes	genes
T3949	17-37	GO_SO_EXT:chromosomal_part_or_position_or_region_or_site	denotes	chromosomal segments
T3950	58-63	SO_EXT:0000704	denotes	genes
T3951	93-97	GO_PR_EXT:fibroblast_growth_factor_receptor	denotes	FGFR
T3952	98-103	SO_EXT:0000704	denotes	genes
T3953	134-144	NCBITaxon:7742	denotes	vertebrate
T3954	145-150	PR_EXT:000016006	denotes	TACC1
T3955	153-158	SO_EXT:0000704	denotes	genes
T3956	195-214	GO_SO_EXT:chromosomal_part_or_position_or_region_or_site	denotes	chromosomal regions
T3957	230-234	GO_PR_EXT:fibroblast_growth_factor_receptor	denotes	FGFR
T3958	235-239	SO_EXT:0000704	denotes	gene
T3959	250-256	NCBITaxon:9606	denotes	humans
T3960	258-263	NCBITaxon:10088	denotes	mouse
T3961	272-280	NCBITaxon:6656	denotes	arthopod
T3962	281-296	NCBITaxon:7227	denotes	D. melanogaster
T3963	315-327	SO_EXT:biological_conservation_process_or_quality	denotes	conservation
T3964	331-341	_FRAGMENT	denotes	paralogous
T3965	354-362	SO_EXT:0000854	denotes	segments
T3966	342-362	GO_SO_EXT:chromosomal_part_or_position_or_region_or_site	denotes	chromosomal segments
T3967	377-386	NCBITaxon:1	denotes	organisms
T3968	486-505	GO_SO_EXT:chromosomal_part_or_position_or_region_or_site	denotes	chromosomal segment
T3969	510-520	SO_EXT:sequence_duplication_process	denotes	duplicated
T3970	534-544	NCBITaxon:7742	denotes	vertebrate
T3971	571-582	SO_EXT:sequence_duplication_entity_or_process	denotes	duplication
T3972	605-610	NCBITaxon:9606	denotes	human
T4005	2258-2270	SO_EXT:biological_conservation_process_or_quality	denotes	conservation
T4090	4536-4546	_FRAGMENT	denotes	paralogous
T4091	4559-4566	SO_EXT:0000854	denotes	segment
T4092	4547-4566	GO_SO_EXT:chromosomal_part_or_position_or_region_or_site	denotes	chromosomal segment
T4093	4568-4577	SO_EXT:biological_conservation_process_or_quality	denotes	Conserved
T4094	4578-4588	SO:0000859	denotes	paralogous
T4095	4644-4649	SO_EXT:0000704	denotes	genes
T4096	4682-4689	SO_EXT:0001026	denotes	genomic
T4097	4690-4696	SO_EXT:0000149	denotes	contig
T4098	4778-4788	_FRAGMENT	denotes	paralogous
T4099	4801-4809	SO_EXT:0000854	denotes	segments
T4100	4789-4809	GO_SO_EXT:chromosomal_part_or_position_or_region_or_site	denotes	chromosomal segments
T4101	4826-4837	NCBITaxon:7742	denotes	vertebrates
T4102	4856-4862	SO_EXT:0001026	denotes	genome
T4103	4870-4881	NCBITaxon:7712	denotes	urochordate
T4104	4882-4897	NCBITaxon:7719	denotes	C. intestinalis
T4105	4922-4933	SO_EXT:0000855	denotes	orthologues
T4106	4937-4941	GO_PR_EXT:fibroblast_growth_factor_receptor	denotes	FGFR
T4107	4947-4952	PR_EXT:000030426	denotes	WHSC1
T4108	4954-4970	GO_EXT:0004180	denotes	carboxypeptidase
T4109	4954-4972	PR_EXT:000005841	denotes	carboxypeptidase Z
T4110	5008-5009	CHEBI_SO_EXT:base	denotes	b
T4111	5024-5039	NCBITaxon:7719	denotes	C. intestinalis
T4112	5040-5046	SO_EXT:0001026	denotes	genome
T4113	5055-5060	NCBITaxon:9606	denotes	human
T4114	5061-5072	SO_EXT:0000855	denotes	orthologues
T4115	5097-5116	SO_EXT:0000854	denotes	paralogous segments
T4116	5220-5225	SO_EXT:0000704	denotes	genes
T4117	5235-5245	NCBITaxon:7742	denotes	vertebrate
T4118	5246-5265	SO_EXT:0000854	denotes	paralogous segments
R8157	T12593	T12592	_lexicallyChainedTo	on chromosome,region
R8158	T12602	T12601	_lexicallyChainedTo	regions,syntenic
R2438	T3965	T3964	_lexicallyChainedTo	segments,paralogous
R2439	T3981	T3980	_lexicallyChainedTo	segments,paralogous
R2440	T4084	T4083	_lexicallyChainedTo	chromosome,regions of
R2441	T4091	T4090	_lexicallyChainedTo	segment,paralogous
R2442	T4099	T4098	_lexicallyChainedTo	segments,paralogous
R8280	T12796	T12795	_lexicallyChainedTo	chromosome,arm of
R8281	T12807	T12806	_lexicallyChainedTo	chromosome,arm of

craft-ca-core-dev

Below, discontinuous spans are shown in the chain model. You can change it to the bag model.

Id	Subject	Object	Predicate	Lexical cue
T3806	235-239	SO:0000704	denotes	gene
T3807	250-256	NCBITaxon:9606	denotes	humans
T3808	258-263	NCBITaxon:10088	denotes	mouse
T3809	272-280	NCBITaxon:6656	denotes	arthopod
T3810	281-296	NCBITaxon:7227	denotes	D. melanogaster
T3811	331-341	_FRAGMENT	denotes	paralogous
T3812	354-362	SO:0000854	denotes	segments
T3813	377-386	NCBITaxon:1	denotes	organisms
T3814	534-544	NCBITaxon:7742	denotes	vertebrate
T3815	605-610	NCBITaxon:9606	denotes	human
T3816	640-645	NCBITaxon:9606	denotes	human
T3817	788-792	SO:0000704	denotes	gene
T3818	886-896	_FRAGMENT	denotes	paralogous
T3819	909-917	SO:0000854	denotes	segments
T3820	1013-1017	SO:0000704	denotes	gene
T3821	1021-1032	NCBITaxon:7742	denotes	vertebrates
T3822	1080-1085	PR:000001451	denotes	FGFR4
T3823	1143-1148	PR:000001856	denotes	RHAMM
T3824	1149-1153	SO:0000704	denotes	gene
T3825	1191-1196	NCBITaxon:9606	denotes	Human
T3826	1197-1202	PR:000001856	denotes	RHAMM
T3827	1250-1254	PR:000008212	denotes	GPX3
T3828	1259-1264	PR:000011245	denotes	NKX2E
T3829	1302-1307	SO:0000704	denotes	genes
T3830	1311-1316	NCBITaxon:9606	denotes	human
T3831	1339-1349	SO:0000859	denotes	paralogous
T3832	1437-1442	NCBITaxon:10088	denotes	mouse
T3833	1447-1450	NCBITaxon:10114	denotes	rat
T3834	1501-1506	SO:0000704	denotes	genes
T3835	1549-1556	NCBITaxon:9443	denotes	primate
T3836	1557-1563	NCBITaxon:9989	denotes	rodent
T3837	2274-2278	SO:0000704	denotes	gene
T3838	2328-2332	SO:0000704	denotes	gene
T3839	2333-2343	GO:0065007	denotes	regulation
T3840	2395-2414	SO:0000854	denotes	paralogous segments
T3841	2432-2438	SO:0001026	denotes	genome
T3842	2446-2454	NCBITaxon:7712	denotes	tunicate
T3843	2455-2470	NCBITaxon:7719	denotes	C. intestinalis
T3844	2509-2519	NCBITaxon:7742	denotes	vertebrate
T3845	2520-2526	SO:0001026	denotes	genome
T3846	2546-2557	NCBITaxon:31033	denotes	T. rubripes
T3847	2613-2619	SO:0001026	denotes	genome
T3848	2623-2634	NCBITaxon:31033	denotes	T. rubripes
T3849	2682-2692	SO:0000859	denotes	paralogous
T3850	2718-2729	NCBITaxon:7742	denotes	vertebrates
T3851	2758-2769	SO:0000855	denotes	orthologues
T3852	2773-2779	PR:000008258	denotes	GPRK2L
T3853	2784-2789	PR:000013947	denotes	RGS12
T3854	2803-2814	NCBITaxon:31033	denotes	T. rubripes
T3855	2815-2823	SO:0001248	denotes	scaffold
T3856	2885-2890	NCBITaxon:9606	denotes	human
T3857	2901-2912	NCBITaxon:31033	denotes	T. rubripes
T3858	2913-2924	SO:0000855	denotes	orthologues
T3859	2928-2933	PR:000001450	denotes	FGFR3
T3864	3077-3082	SO:0000704	denotes	genes
T3865	3150-3155	PR:000016008	denotes	TACC3
T3866	3160-3165	PR:000007506	denotes	FGFRL
T3867	3189-3198	SO:0001248	denotes	scaffolds
T3868	3258-3262	SO:0000704	denotes	gene
T3869	3297-3302	PR:000008260	denotes	GPRK6
T3870	3306-3311	NCBITaxon:9606	denotes	human
T3871	3354-3364	NCBITaxon:31031	denotes	pufferfish
T3872	3375-3386	NCBITaxon:31033	denotes	T. rubripes
T3873	3387-3398	SO:0000855	denotes	orthologues
T3874	3402-3406	PR:000011441	denotes	NSD1
T3875	3408-3413	PR:000001451	denotes	FGFR4
T3876	3429-3433	SO:0000704	denotes	gene
T3877	3445-3453	SO:0001248	denotes	scaffold
T3878	3514-3518	SO:0000704	denotes	gene
T3879	3544-3548	SO:0000704	denotes	gene
T3880	3663-3668	PR:000016006	denotes	TACC1
T3881	3677-3682	SO:0000704	denotes	genes
T3882	3690-3700	NCBITaxon:31031	denotes	pufferfish
T3883	3736-3744	SO:0001248	denotes	scaffold
T3884	3795-3806	SO:0000855	denotes	orthologues
T3885	3818-3823	SO:0000704	denotes	genes
T3886	3835-3840	NCBITaxon:9606	denotes	human
T3887	3872-3880	SO:0001248	denotes	scaffold
T3888	3913-3918	PR:000016006	denotes	TACC1
T3889	3965-3976	NCBITaxon:7742	denotes	vertebrates
T3890	4010-4014	SO:0000704	denotes	gene
T3891	4040-4048	SO:0001248	denotes	scaffold
T3892	4077-4085	SO:0001248	denotes	scaffold
T3893	4104-4115	NCBITaxon:31033	denotes	T. rubripes
T3894	4116-4127	SO:0000855	denotes	orthologues
T3895	4131-4135	PR:000010686	denotes	MSX1
T3896	4137-4142	PR:000015790	denotes	STX18
T3897	4160-4174	SO:0000996	denotes	predicted gene
T3898	4216-4224	SO:0000857	denotes	homology
T3899	4228-4232	PR:000009884	denotes	LOXL
T3900	4234-4237	PR:000007232	denotes	EVC
T3901	4276-4284	SO:0001248	denotes	scaffold
T3902	4297-4302	SO:0000704	denotes	genes
T3903	4327-4332	NCBITaxon:9606	denotes	human
T3904	4384-4389	PR:000016008	denotes	TACC3
T3905	4394-4399	PR:000016007	denotes	TACC2
T3906	4459-4466	SO:0001026	denotes	genomic
T3907	4536-4546	_FRAGMENT	denotes	paralogous
T3908	4559-4566	SO:0000854	denotes	segment
T3909	4578-4588	SO:0000859	denotes	paralogous
T3910	4644-4649	SO:0000704	denotes	genes
T3911	4682-4689	SO:0001026	denotes	genomic
T3912	4690-4696	SO:0000149	denotes	contig
T3913	4778-4788	_FRAGMENT	denotes	paralogous
T3914	4801-4809	SO:0000854	denotes	segments
T3915	4826-4837	NCBITaxon:7742	denotes	vertebrates
T3916	4856-4862	SO:0001026	denotes	genome
T3917	4870-4881	NCBITaxon:7712	denotes	urochordate
T3918	4882-4897	NCBITaxon:7719	denotes	C. intestinalis
T3919	4922-4933	SO:0000855	denotes	orthologues
T3920	4947-4952	PR:000030426	denotes	WHSC1
T3921	4954-4972	PR:000005841	denotes	carboxypeptidase Z
T3922	5024-5039	NCBITaxon:7719	denotes	C. intestinalis
T3923	5040-5046	SO:0001026	denotes	genome
T3924	5055-5060	NCBITaxon:9606	denotes	human
T3925	5061-5072	SO:0000855	denotes	orthologues
T3926	5097-5116	SO:0000854	denotes	paralogous segments
T3927	5220-5225	SO:0000704	denotes	genes
T3928	5235-5245	NCBITaxon:7742	denotes	vertebrate
T3929	5246-5265	SO:0000854	denotes	paralogous segments
T3930	5298-5303	PR:000001856	denotes	RHAMM
T3931	5304-5309	SO:0000704	denotes	genes
T3932	5313-5328	NCBITaxon:7719	denotes	C. intestinalis
T3933	5379-5398	SO:0000854	denotes	paralogous segments
T3934	5426-5431	SO:0000704	denotes	genes
T3935	5529-5538	NCBITaxon:7712	denotes	tunicates
T3936	5583-5594	NCBITaxon:31033	denotes	T. rubripes
T3937	5595-5601	SO:0001026	denotes	genome
T3938	5732-5737	PR:000001450	denotes	FGFR3
T3939	5772-5776	SO:0000704	denotes	gene
T3940	5793-5797	SO:0000704	denotes	gene
T3941	5821-5831	NCBITaxon:7742	denotes	vertebrate
T3942	5874-5886	NCBITaxon:61463	denotes	Gnanthostome
T3943	5961-5979	SO:0000854	denotes	paralogous segment
T3944	6013-6022	NCBITaxon:7737	denotes	amphioxus
T3945	6027-6034	NCBITaxon:7745	denotes	lamprey
T3946	6035-6042	SO:0001026	denotes	genomes
T3947	6069-6073	SO:0000704	denotes	gene
T3948	6145-6150	SO:0000704	denotes	genes
T12565	1586-1592	SO:0000987	denotes	Linear
T12566	1609-1613	SO:0000704	denotes	gene
T12567	1671-1676	SO:0000704	denotes	genes
T12568	1680-1686	NCBITaxon:9606	denotes	humans
T12569	1688-1698	SO:0000859	denotes	Paralogous
T12570	1699-1704	SO:0000704	denotes	genes
T12571	1798-1802	PR:000008212	denotes	GPX3
T12572	1807-1812	PR:000011245	denotes	NKX2E
T12573	1881-1886	SO:0000704	denotes	genes
T12574	1948-1953	SO:0000704	denotes	genes
T12575	1989-1997	_FRAGMENT	denotes	syntenic
T12576	2016-2023	SO:0005858	denotes	regions
T12577	1998-2003	NCBITaxon:10088	denotes	mouse
T12578	2045-2062	NCBITaxon:31033	denotes	Takifugu rubripes
T12579	2063-2072	SO:0001248	denotes	scaffolds
T12580	2116-2126	SO:0000857	denotes	homologous
T12581	2127-2131	SO:0000704	denotes	gene
T12582	2188-2198	SO:0000859	denotes	paralogous
T12583	2199-2204	SO:0000704	denotes	genes
T12757	6218-6235	NCBITaxon:31033	denotes	Takifugu rubripes
T12758	6240-6258	NCBITaxon:7719	denotes	Ciona intestinalis
T12759	6282-6289	SO:0001026	denotes	genomic
T12760	6290-6299	SO:0001248	denotes	scaffolds
T12761	6315-6332	NCBITaxon:31033	denotes	Takifugu rubripes
T12762	6355-6360	SO:0000704	denotes	genes
T12763	6362-6370	SO:0001248	denotes	Scaffold
T12764	6401-6405	SO:0000704	denotes	gene
T12765	6415-6420	SO:0000704	denotes	genes
T12766	6439-6449	SO:0000853	denotes	homologues
T12767	6453-6464	SO:0000855	denotes	orthologues
T12768	6484-6490	_FRAGMENT	denotes	arm of
T12769	6497-6507	SO:0000105	denotes	chromosome
T12770	6491-6496	NCBITaxon:9606	denotes	human
T12771	6526-6534	SO:0001248	denotes	scaffold
T12772	6622-6627	PR:000016006	denotes	TACC1
T12773	6628-6633	PR:000016007	denotes	TACC2
T12774	6676-6684	SO:0001248	denotes	scaffold
T12775	6705-6716	SO:0000855	denotes	orthologues
T12776	6720-6725	SO:0000704	denotes	genes
T12777	6754-6760	_FRAGMENT	denotes	arm of
T12778	6767-6777	SO:0000105	denotes	chromosome
T12779	6761-6766	NCBITaxon:9606	denotes	human
T12780	6786-6804	NCBITaxon:7719	denotes	Ciona intestinalis
T12781	6825-6830	SO:0000704	denotes	genes
T12782	6840-6859	SO:0000854	denotes	paralogous segments
T12783	6863-6868	NCBITaxon:9606	denotes	human
T12956	6900-6907	SO:0001026	denotes	Genomic
T12957	6930-6935	SO:0000704	denotes	genes
T12958	6992-6997	NCBITaxon:9606	denotes	human
T12959	7041-7046	NCBITaxon:9606	denotes	human
T12960	7047-7052	PR:000016006	denotes	TACC1
T12961	7072-7077	PR:000017527	denotes	GAS41
T12962	7116-7122	SO:0000417	denotes	domain
T12963	7129-7135	PR:000005512	denotes	ch-TOG
T12964	7155-7169	PR:000035365	denotes	Aurora kinases
T12965	7177-7184	NCBITaxon:species	denotes	species
T12966	7254-7259	SO:0000147	denotes	exons
T12967	7272-7277	SO:0000704	denotes	genes
T12968	7303-7310	SO:0001060	denotes	isoform
T12969	7320-7324	SO:0000704	denotes	gene
T3801	58-63	SO:0000704	denotes	genes
T3802	98-103	SO:0000704	denotes	genes
T3803	134-144	NCBITaxon:7742	denotes	vertebrate
T3804	145-150	PR:000016006	denotes	TACC1
T3805	153-158	SO:0000704	denotes	genes
T3860	2982-2989	SO:0001026	denotes	genomic
T3861	2990-2998	SO:0001248	denotes	scaffold
T3862	3050-3055	NCBITaxon:9606	denotes	human
T3863	3056-3067	SO:0000855	denotes	orthologues
R8156	T12576	T12575	_lexicallyChainedTo	regions,syntenic
R8278	T12769	T12768	_lexicallyChainedTo	chromosome,arm of
R2434	T3812	T3811	_lexicallyChainedTo	segments,paralogous
R2435	T3819	T3818	_lexicallyChainedTo	segments,paralogous
R2436	T3908	T3907	_lexicallyChainedTo	segment,paralogous
R2437	T3914	T3913	_lexicallyChainedTo	segments,paralogous
R8279	T12778	T12777	_lexicallyChainedTo	chromosome,arm of

2_test

Id	Subject	Object	Predicate	Lexical cue
15207008-12777526-9666000	397-399	12777526	denotes	13
15207008-12808028-9666001	1186-1188	12808028	denotes	10
15207008-12481130-9666002	2472-2474	12481130	denotes	11
15207008-12142439-9666003	2559-2561	12142439	denotes	14
15207008-12481130-9666004	4899-4901	12481130	denotes	11

PMC:441373 / 14596-21930 JSONTXT

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2_test

PMC:441373 / 14596-21930 JSON TXT