Acknowledgments This work was supported by a grant from the Next-Generation BioGreen 21 Program (PJ009520 to Sun-Hyung Lim and PJ00994 to Sun-Hwa Ha) and the 2013 Postdoctoral Fellowship Program (to Min Kyoung You) of the National Academy of Agricultural Science, Rural Development Administration, Korea. This work was also supported by a Young Scientist Supporting Grant (20130686 to Sun-Hwa Ha) of Kyung Hee University, Korea and by Basic Science Research Program through the National Research Foundation of Korea (NRF) funded by the Ministry of Education, Science and Technology (NRF-2013R1A1A2062998 to Sun-Hwa Ha).