A subsequent study by Mohanty et al. (2008) also investigated motivation effects on attention, this time manipulating motivational state, namely hunger. Specifically, in the context of a covert-orienting task with a central cue, participants detected motivationally relevant (food) or irrelevant (tools) targets under conditions of hunger and satiety. As in the study by Small et al., responses in sites in parietal cortex (e.g., intraparietal sulcus, IPS) exhibited correlations with the speed of attentional shifts that were sensitive to not just motivational state, but also to the motivational value of the target. Similar patterns were also observed in the PCC and the orbitofrontal cortex (OFC). Furthermore, amygdala, PCC, locus coeruleus and SN/midbrain showed sensitivity to food-related cues when hungry, but not when satiated, an effect that did not generalize to tools. These findings demonstrate that motivational state (hunger) modulates spatial attention via response modulations across several brain regions.