λ G U   = ( n G U  +   n a l l  +    + n G U  −   n a l l  −    ) × 100 ,   MathType@MTEF@5@5@+=feaafiart1ev1aaatCvAUfKttLearuWrP9MDH5MBPbIqV92AaeXatLxBI9gBaebbnrfifHhDYfgasaacH8akY=wiFfYdH8Gipec8Eeeu0xXdbba9frFj0=OqFfea0dXdd9vqai=hGuQ8kuc9pgc9s8qqaq=dirpe0xb9q8qiLsFr0=vr0=vr0dc8meaabaqaciaacaGaaeqabaqabeGadaaakeaaiiGacqWF7oaBdaWgaaWcbaGaem4raCKaemyvaufabeaakiabg2da9iabcIcaOmaalaaabaGaemOBa42aa0baaSqaaiabdEeahjabdwfavbqaaiabgUcaRaaaaOqaaiabd6gaUnaaDaaaleaacqWGHbqycqWGSbaBcqWGSbaBaeaacqGHRaWkaaaaaOGaey4kaSYaaSaaaeaacqWGUbGBdaqhaaWcbaGaem4raCKaemyvaufabaGaeyOeI0caaaGcbaGaemOBa42aa0baaSqaaiabdggaHjabdYgaSjabdYgaSbqaaiabgkHiTaaaaaGccqGGPaqkcqGHxdaTcqaIXaqmcqaIWaamcqaIWaamcqGGSaalaaa@5120@