We thank Dr. Andy Dudley for his help in designing mouse genotyping strategies. We also thank Jose Rivera-Feliciano and Dr. Douglas Kim for helpful discussions. We are immensely grateful to Drs. Brigid Hogan and Holger Kulessa for their kind gift of the Bmp4 conditional allele and Drs. Liz Robertson and Andy Dudley for the Bmp7 knockout allele. We would like to acknowledge the help of Dr. Ernestina Schipani and Jan Saxton for help with histology.