Prevalence of Antibodies to Four Human Coronaviruses Is Lower in Nasal Secretions than in Serum ᰔ Downloaded from Abstract Little is known about the prevalence of mucosal antibodies induced by infection with human coronaviruses (HCoV), including HCoV-229E and -OC43 and recently described strains (HCoV-NL63 and -HKU1). By