Acknowledgments The assistance of Tarja Hovivuori in qPCR and Anneli Virta in Sanger sequencing is greatly appreciated. This study was funded by the Academy of Finland (http://www.aka.fi/). The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. The authors declare that they have no competing interests. D.F. participated in the bioinformatic analysis of the data, developed the in-house R-package HoardeR, analyzed the data using HoardeR, and helped draft the manuscript. A.L. performed the Solid 4 sequencing run and participated in the mapping and DE analysis of the data. A.G. participated in the design and execution of the sequencing protocol. A.S. designed and coordinated the study, performed the qPCR and Sanger sequencing and part of the bioinformatic analysis, and drafted the manuscript. All authors read and approved the final manuscript.